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BUMEE TINS
AMERICAN
PALEONTOLOGY
VOL. 9
March, 1921 — June, 1922
Harris Co.
Cornell Univ., Ithaca, N. Y.
LOSES oa
CONTENTS OF VOL. IX
Bull. No.
37 —New Eocene Species from Alabama.
Bye eten VAUl cliriela ys) ae ee even Pl’s 1-3
38 —Recent Mollusca of the Gulf of Mexico
and Pleistocene and Pliocene
Species from the Gulf States.
Part 2.—Scaphopoda, Gastropoda,
Amphineura and Cephalopoda.
By Ce iaiMaunyansneeee eee aaeies
39 —The Miocene of Northern Costa Rica
with Notes on its General Strati-
graphic Relations.
hy Js oe (OSS0ysljgadenssessuasansose Pl’s 4-35
Page
1-32
33-173
174-482
13, 96)
BULLETINS
OF
AMERICAN PALEONTOLOGY
Vol. 9
No. 37
NEW EOCENE SPECIES FROM ALABAMA
BY
T. H. ALDRICH
March 6, 1921
Ne
<
* Harris Co.
Coriell Univ., Ithaca, N. Y.
LB SEN
PREEACEH.
The late Herbert H. Smith, curator of the Alabama State
Museum of Natural History University, Alabama, made exten-
sive collections in the Tertiary of Alabama, and at the time of
his sudden and lamented death was engaged in classifying and
arranging his material. The collections were made on joint ac-
count for the Carnegie Museum of Pittsburg and the Alabama
Museum of Natural History. The condition of the material
made it necessary to go over all of it, and classify it for division
as well as to describe what was considered new. The writer has
also added a few new species from his collecting. It is regretted
that Mr. Smith was not spared to complete his labors, because -
his ability in specific determinations was almost unique. The
types of the new species collected by him are all in the Univer-
sity Museum. We are indebted te—Dr. W. H. Dall, Dr. Paul
Bartsch, Prof. G. D. Harris and Dr. F. A. Smith for generous
aid in both specific and generic determinations of doubtful forms.
a ey
leak
NEw HocENE SPECIES FROM ALABAMA
BY
he ae eA RICE
Mitramorpha turriculata, n. sp. lel iG Sebo 10
Shell small, biconic, whorls eight, three embryonic and
smooth, the fourth with strong longitudinal ribs, which continue
over the other whorls, dying down some on the body whorl.
Spiral strize numerous, and impressed, a strong double band bor-
dering the suture, aperture narrow, smooth within. Length
6 mm.
Locality.—Choctaw Corner, Ala. Woods Bluff horizon.
Type.—My cabinet.
Remarks.—This little shell seems to be nearest to Witramor-
pha, it has a glazed appearance ; there is some evidence of a
slight notch on and above the periphery of the body whorl.
Turris (Pleurotoma) nodoideus, n. sp. Pls te Riss ya
Shell small, spire pointed, whorls nine ; first four embryon-
ic, the fifth bearing longitudinal ribs in part, and revolving lines
in part with the ribs also present but without the subsutural col-
lar below. ‘The sixth, seventh and eighth whorls with the bor-
dering part of the ninth carrying first, a subsutural collar which
becomes doubled and thicker towards the body whorl and is
marked with very numerous raised lines belonging to the fasciole;
second, just below is a rather broad concave revolving space
reaching to the peripheral parts of the whorls carrying very
numerous close set semicircular anal markings, these continue
6 BULLETIN 37 6
over the body whorl in curved raised lines : third, the periphery
sometimes carrying a couple of coarse lines followed below by
from four to six raised lines, also raised longitudinal ribs from
the periphery to suture. Body whorl with raised revolving lines
extending to the end of the canal, carrying a few alternating
ones belowthe periphery ; aperture smooth within ; canal slightly
rostrate.
Locality.—Cave Branch, 6 miles E. of Thomasville, Ala.
Woods Bluff beds.
Type.—Ala. Museum. University, Ala.
Remarks.—This species seems to differ in some respects from
any heretofore described, but it may be only a variety of some
existing species like Zurris denticula Bast. et. al. The fact is
thatthe Woods Bluff beds at Cave Branch contain so many numer-
ous varieties that it is almost impossible to locate them property.
The whole subject of species in this family needs careful study.
The young are often obtuse, and differ in shape and ornamenta-
tion from the adult so that they could be called new species, and
pass muster while a series would connect them all together. Am
not prepared to go into details but hope this subject will be
looked into by some of our younger students. Some specimens
may be hybrids, and are altogether perplexing. One ought to
have authentic specimens of the described species to even begin
the examination. All the species of this group from this locality
seem somewhat stunted.
Turris specus, n. sp., JEN Neb eR WAR se
Shell small, whorls probably seven (the embryonic tip is de-
fective) first three embryonic, then four spiral whorls ornamented
with a strong subsutural collar, then a slightly concave revolving
fasciolar space covering about one fourth of the whorl, followed
by a coarse cancellated surface ; the ribs cut by about four im-
pressed revolving lines making a series of nodes, body whorl with
the same ornamentation above followed by coarse revolving lines
growing finer to the base ; outer lip smooth inside, columella
thickened, base somewhat rostrate.
Locality,—Cave Branch, Ala. Woods Bluff beds.
7 ALDRICH, KOCENE MOLLUSCA 7
Type.—Ala. Mus. of Nat. History, University, Ala.
Microdriilia rostratuia Csy.
This species described by Col. Casey in 1903 bears a certain
resemblance to Pleuvotoma insignifica Heilpr. (Fusus nanas Lea)
and has been referred to this species, but specimens before me
show differences as follows: Col. Casey says the ‘‘body whorls
are about four in number.’’ My specimens have five, also in the
embryo there is sometimes but one costulate whorl, while
Pleuro. insignifica Heilpr. has but three embryonic whorls, all
smooth, and no costulate ones. The two species seem to be
quite distinct. The Texas form mentioned by Prof. Harris is
probably the same as Heilprin’s species.
SPECIES TO BE ADDED TO THE Woops BLUFF FAUNA—
FROM CAVE BRANCH, ALA.
Exilia pergracilis Con.
Cylichna deKayt Lea.
Cancellarta alveata Con.
ae costata Lea.
Pasithea guttula Lea (H. H. Smith).
Cancellaria panones Harris.
Ancillaria subglobosa Con.
Levifusus sutert Aldr.
Caricella dolita Con.
Drillia lonsdalit Vea.
Barnea alatoidea Aldr.
Diplodonta nana Vea.
Egerella subtrigonia Lea.
Tellina leana Dall.
Turris bimoniatus, n. sp. Pliitea), Hig:
Shell small, whorls probably ten, spire on the type partially
missing. Another specimen shows four embryonic smooth
whorls. The four spiral whorls and part of the fifth ornamented
with curved sinuosities covering the main part of the whorls, a
8 BULLETIN 37 8
strong collar below the sutures, the balance showing a double
collar with a concave space between ; the whole of the surface
with numerous spiral raised lines. Aperture about a third of
the length of shell, lines on the base have a tendency to alter-
nate. Length 14 mm. !
Locality.—Cave Branch, Ala. Woods Bluff beds.
Type.—Ala. Museum, University, Ala.
Remarks.—'This seems to combine the characteristics of two
or more species, and if there were only one specimen, could be
considered a variety. It seems to be related to 7. monzliata
Heilpr.
Pleurotoma (Peratotoma) Gardneri, n. sp. TEN sey) Teds ich
Shell medium, substance rather thin, broadly conic ; whorls
nine to ten, embryonic ones four in number, the spire very small,
but the whorls rapidly expanding, the permanent whorls profuse-
ly ornamented with both longitudinal and spiral raised lines, the
peripheral part and below carrying nodes while the spirals are
alternate in size and thickness. Body whorl large, shouldered,
profusely ornamented, the spirals nodular ; just below the suture
there are a couple coarser than the others and also at base ; aper-
ture half the length of the shell, smooth within, inner lip witha
thin callus ; canal recurved and slightly spatulate. Length
21 mm.
Locality.—Pugh’s branch of Satilpa Creek, Ala., Gosport
Sand.
Type.—Ala. Museum of Nat. History, University, Ala.
Remarks.—This species is on the borderland between /uszs,
Strepsidura and Pleurotoma. It isvery close to /l. hetlprini,
nobis from Jackson, Miss. The largest specimen is 25 mm. in
length. The slit is not pronounced. Named in honor of Miss
Julia A. Gardner of the National Museum.
Odostomia (Evalea) Bartschi, n. sp. eal ats abe ey,
Shell small, whorls six, spire almost totally immersed, shell
rather rapidly expanding. Basal whorl large, suture deep, aper-
ture oblong. One strong, straight fold on the columella placed
9 ALDRICH, HOCENE MoLLUSCA 9
centrally; umbilicus open, inner lip somewhat recurved at base.
Surface not smooth, carrying microscopic incised spiral lines.
Length 3 mm.
Locality.—Found at both Gregg’s and Bell’s Landings.
Type.—From Bell’s Ldg. In the Ala. Museum.
Turbonilla (Ptycheulimella) clinensis, n. sp. LAL eg) Vetkerh 10),
Shell small, narrow, whorls eight, two embryonic, the next
three longitudinally striated, balance smooth. Spire blunt,
suture distinct, aperture small. Outer lip smooth within, inner
lip with a strong fold curving up into the aperture and bordering
the base. Length 3+ mm.
Locality.—Gregg’s Landing, Ala.
Type.—Alabama Museum, University, Ala.
Remarks.—Seems to be a distinct species. There are three
other species of 7urbonzlla in the State Collection but too imper-
fect to warrant description.
Turbonilla (Cinguiina) tuscahomensis, n. sp. enh 3640), Teme ath
Shell small, whorls nine, the first three embryonic ; apex
small, the fourth and fifth whorls with raised spirals, balance
carrying five strong spirals, and somewhat cancellated ; suture
deeply impressed. The space above the first or lowest spiral
slightly wider than between the other four. Aperture broadly
ovate ; inner lip slightly reflected ; base slightly spatulate.
Length 4mm. Figured specimen from Bell’s Ldg.
Locality.—Both Gregg’s and Bell’s Ldg., Alabama.
Type.—Alabama Museum of Nat. History.
Remarks.—\Looks like a Lzttium.
Turbonilla (Ptycheulimella) tardiusculus, n. sp. Pie Rigen 2.
Shell with nine whorls, the first two embryonic, the balance
smooth and of a dull brown color, substance of shell rather solid,
suture lightly impressed, banded below. Aperture small, inner
lip reflected at base, imperforate ; no fold showing on the col-
umella. Two specimens found. Length4% mm.
Locality.—Cave Branch, Ala. Woods bluff beds.
Type.—My collection. ©
fe) BULLETIN 37 10
Pyramidetta (Iphiana) anita Ald.
The Nautilus, Vol. XXI, No. I, p. 9, Pl. 1, Fig. 12, May, 1907.
This species was described as a 7urbonilla, but is now put
into the Pyramidellide by Dr. Bartsch.
Pyramideila (Syrnola) mitchelliana Aldr.
Bull. of American Paleontology, No. 22, P. 8, Pl. 3, Fig. 3, IgII.
Is also removed from 7urbonzlla.
The Ala. Museum has a number of new forms of small spe-
cies, collected by Mr. Herbert A. Smith which are described
herein. They have been submitted to Dr. Bartsch who has kind-
ly indicated their generic position.
Pyramidella (Iphiana) obtusoides, n. sp. IP yes Webel Tey
Shell small, with four whorls capped by two embryonic ones
turned abruptly to the back side of shell and partially immersed.
The specimens, three in number, have a dull surface, apparently
smooth, but under a glass numerous spiral lines are seen.
Suture not deeply impressed ; aperture oblong-ovate ; outer lip
sharp, smooth within ; inner lip reflected, somewhat twisted,
carrying a single smalltooth. Length 3% mm.
Locality.—Found at both Gregg’s and Bell’s Landings, Ala.
River, Ala.
Type.—Ala. Museum of Natural History.
Remarks.—A short, dull-looking species.
Epitonium munistriatum, n. sp. Pion. pehigse tA a5
Shell small, aciculate whorls, probably toor more. Embry-
onic ones three in number, apex pointed ; the next two or three
with longitudinals only, balance cancellated. Whorls rounded,
suture deep. Spirals 5 to 7, rather coarse, ribs raised with con-
cave spaces between, base with smaller alternate spirals between
the primaries ; intersections pointed. ‘The base of this species
(15) is in the Museum from Gregg’s Landing, Ala. The other
specimen is from Woods Bluff, Ala. The aperture shows a
a thickened inner lip where it joins the base, also a thin callus.
Locality.—As above.
Type—Ala. Museum of Nat. History.
II ALDRICH, KOCENE MOLLUSCA Il
Epitonium multiliniferum, n. sp. Veda GD ahkeg atop
All the specimens are fragmentary, but well marked. Em-
bryonic whorls missing ; suture distinct. Ribs on body whorl
34 in number, extending over the base, much weaker there ;
very numerous close set spiral lines between the ribs. | Aperture
round, inner lip thickened, no umbilicus.
Locality.—Cave Branch, Ala. Woods Bluff beds.
Type.—My cabinet.
Remarks.—This is a distinct species and it deserves differen-
tiation from the other forms. ‘The figure given by Prof. Harris
in Bull. Am. Paleontology, No. 11, Pl. 12, Fig. 8, is probably a
young example of this species. The sides are more nearly par-
allel than in most Apztonzums.
Epitonium subacutum, n. sp. Pir Bigs s 175 0o:
Shell thin, narrowly acute, whorls 12. The first five are em-
bryonic and smooth ; the balance spirally striated ; cancellated.
The vertical ribs on the body whorl about twenty-six in number,
curving into the umbilicus. They are thin or acute ; the pri-
mary spirals about ten in number with finer alternate lines be-
tween ; inner lip with a slight callus. Length 10 mm.
Locality.—Cave Branch, Ala. Woods Bluff beds.
Type.—In Ala. Museum. One specimen shows the base and
the other the embryonic shell.
Teinostoma subangulata Mr. var. Smithii, n. var. VEAL id ty |i dietary, ako}, VIO
Shell small, flattened above, rounded below ; umbilicus not
large ; a callus on the base which enters the umbilicus and con-
tinues within. Bordering the suture is an appressed area raised
above the whorls ; surface above with numerous close set revolv-
ing lines ; periphery smooth with a few revolving lines below.
Aperture circular ; outer lip somewhat thickened ; lines of
growth perceptible ; interior somewhat pearly. Diameter 8 mm.
Localtty,—Bell’s Landing, Ala.
Type.—Ala. Museum of Natural History.
Remarks.—Very much larger than the type of 7. subangu-
lata Mr. Looks very much like a land shell. It differs some-
what from Meyer’s species in the shape of the aperture.
12 BULLETIN 37 12
Teinostoma reguiaris, n. sp. Bie igs. 2 1-2»
Shell small but larger than the ordinary species of this
genus ; whorls five, depressed above ; periphery rounded ; sur-
face covered with fine lines which are crossed with lines of
growth. Under a glass the intersections give the surface a
beaded appearance ; a depressed space just below the suture.
The base has coarser lines with a smooth space around the um-
bilicus which is open ; aperture oval. Diameter 4 mim.
Locality.—Pugh’s Branch of Satilpa Creek, Clark Co., Ala.
Gosport Sand.
Type.—Ala. Museum at University, Ala.
Remarks.—This little shell may possibly belong to the sub-
genus Omphalius of Chlorostoma but it seems closer to Zeznos-
toma.
Ancillopsis Tuomoyi, n. sp. Teds ttys | JE hes\, Ae oy,
Shell solid, thick, oblong, smooth and shining ; whorls 5-6 ;
spire rather pointed ; the first four whorls small, the others rap-
idly increasing ; a swollen callus reaching above the body whorl
and also covering the front of this whorl and lapping over the
back of this whorl. A deep suture reaches the aperture where
it is rather wide ; a few irregular lines of growth are visible ;
aperture about three-fourths the length of the shell, oblong-ovate;
columella smooth, concave. The basal groove shows strongly at
base, but only showing a short distance up where it is covered by
the enamel. Length 28 mm ; breadth 21 mm.
Locality.—Bell’s Landing, lower bed, also one example from
Gregg’s Landing, Ala. The lower bed at Bell’s Landing is the
same bed as at Gregg’s Ldg. ‘Type from Bell’s Landing, lower
bed.
Type.—Ala. Museum of Nat. History.
Remarks.—This species resembles //acron in some respects
but is evidently an Azeczllopsis.
Nassa pleona, n. sp. Plvmeebigsy 25.26:
Shell small, nuclear whorls broken off. Four remaining
whorls carry longitudinal ribs that are crossed by two spirals,
13 ALDRICH, KOCENE MOLLUSCA 13
nodular at intersections. Two close set spirals border the suture
above and below same; suture channelled ; the last whorl car-
ries about Io widely spaced spirals, raised and nodular with con-
cave spaces between. Outer lip thickened with the spirals ex-
tending over same, denticulated ; inner lip enameled with the
spirals showing through the same. Canal very short ending in
a semicircular opening. Length 12+ mm. ‘Two specimens ob-
tained.
Localtty—Lisbon, Ala.
Type.—My cabinet.
Glivella semilignitica, n. sp. Te ebhers: By, ates
Shell small, whorls six, four smooth ; the fourth and subse-
quent ones bounded by a raised band just below the suture ; apex
blunt, body whorl showing impressed lines of growth ; aperture
asin the genus. Length 5% mm.
Locality.—Clark County, Ala. Woods Bluff horizon.
Type. —Ala. Museum of Nat. History.
Remarks.—The sutural callosity pronounced on the last
whorl only. Differs from O. mediavia Harris in being smaller,
more blunt, with fewer folds on the columella, and by a raised
sutural band.
Gilbertia esteliensis, n. sp. AL Ds | Jaber eas
Shell globular, whorls three ; the first embryonic ; the sec-
ond spirally striated but very narrow ; the third very globose
and spirally striated ; outer lip thickened and broadly reflected ;
a single tooth projecting into the aperture from it ; inner lip
thickened with a broad spatulate tooth above, and two prominent
teeth near the base ; the outer lip partially covering the second
whorl. Under a glass the strize show that they are pitted as in
Actgon. Height 2% mm. ; breadth 2 mm.
Locality.—Sucarnoochee beds, 3 miles south of Estella, Ala.
Ty pe.—Ala. State Museum, University, Ala.
Canceilaria Gilberti, n. sp. Ble ie REO 20%
Shell with seven whorls remaining ; the spire is damaged ;
cancellated ; suture deeply impressed and constricted, especially
14 BULLETIN 37 14
on the lower whorls. Below the suture is a raised cord marked
with a spiral line upon the lower whorls; above the suture is a
wider space bounded by strong raised spirals with other raised
spirals above. Body whorl expanded with a strong spiral at the
periphery ; those above this are close together, those below al-
ternated with a finer spiral; lines of growth show on body whorl
reversing near the base ; aperture oblong-ovate, the spiral lines
rounding into same ; columella with two folds ; shell has a small
umbilicus. Labrum lirate within. Length 9 mm.; breadth 4mm.
Locality.—Grege’s Ldg., Ala. River, Ala.
Type.—Ala. Museum of Nat. History.
Remarks.—Only one specimen found. This species has
many of the characters of Admete.
Qvula reguiaroidea, nu. sp. LEI Se |) Deabezy evo
Shell small, elongate ; both anterior and posterior with fine
revolving lines ; main body of shell smooth ; outer lip thickened
and crenulated from beak to base ; inner lip also crenulated its
whole length ; a heavy callus on the outer surface of the col-
umella at the posterior end with the canal extending beyond the
callus. On the smaller of the two specimens before me there are
fine revolving lines half way down from the posterior ; aperture
narrow, expanding back of the anterior canal ; ends somewhat
flattened. Length of largest specimen 14 mm.
Locality.—Bell’s Landing mari, Ala. River, Ala.
Type.—Ala. Museum of Nat. History.
Cyprea estellensis, n. sp. ISR) erica
Shell small ; rotund ; smooth ; aperture narrow ; outer lip
curving into the apex which is open. Lengthi2mm. Breadth
ro mm.
Locality.—Sucarnoochee beds, Pursley Creek, Wilcox Co.,
Ala.
Type.—Ala. State Museum.
Remarks.—This specimen has only preserved the shell in
part but the interior is hard rock and gives the form quite accur-
ately. The aperture is all filled with matrix, hiding the denti-
15 ALDRICH, HKOCENE MOLLUSCA 15
tion. Prof. Harris mentions finding one species at Fort Gaines
on the Chattahoochee and another (which is probably the one
subsequently described % mile north of Ripley, Miss.) They
are figured and named as they seem to be rare and peculiar to
this horizon.
Cyprea sp.? Pia i Bigsuauae
Shell small ; surface smooth ; basal callus extending par-
tially over the sides ; outer lip expanded ; extremities somewhat
pointed. One fragment shows the dentition, consisting of over
20 small teeth on the inner lip. Length of the larger fragment
15 mm.
Locality.—Sucarnoocchee beds, one and three miles south of
Estella, Ala. on’ Pursley Creek.
Types.—Ala. Museum of Nat. History.
Seems to be rather close to Cyprea Smutht nobis from
Grege’s Ldg., Ala.
Siphonalia quadrilineata, n. sp. Panos Ak ie
Shell resembling the Ausus subscalarinus Heilpr. Whorls
eight, two embryonic, the balance more or iess cancellated ;
longitudinal costee somewhat rounded; spiral striee very numer-
ous, four fine ones between a coarser line ; the volutions are ap-
pressed at the suture ; the spiral striz cross over the costee and
the coarse lines make nodes of intersections ; canal reflected and
twisted : the outer lip on the type is broken away but appears to
have been smooth internally ; inner lip somewhat thickened.
Length 17mm.
Locality.— Pugh’s Branch of Satilpa Creek, Clark Co., Ala.
Gosport Sand.
Type.—In Ala. Museum, University Ala.
Remark.—One is rather rash to attempt a new form herewith
but the surface ornamentation is peculiar and seems to be new.
Cerithiopsis esteliensis, n. sp. VEIL LN Deikex, 1K)s
Shell minute ; whorls eleven or more, the first two smooth,
the next two with raised ribs ; the balance showing two strong,
heavily beaded spirals on the peripheral part of each whorl with
16 BULLETIN 37 16
a third spiral much finer just below the suture ; one or more
whorls are missing from the basal part.
Length 3 mm.
Locatity.—Sucarnoochee clays, near Estella, Ala.
Type.—in Ala. Museum at University, Ala.
Leevibuccinum (Euryochetus ?) harrisi, n. sp. Ply 2h) VEtes. 7-9:
Shell small ; whorls eight, three smooth, the other five be-
low {with close-set raised lines ; outer lip smooth, slightly in-
curved ; inner lip with a callus, thicker and slightly raised on the
canal, which is short, rather wide ; shell somewhat truncated an-
teriorly ; suture distinct. Length of largest specimen 44% mm.
Locality.—Cave Branch, Ala., about 6 miles east of Thomas-
ville, Ala. Woods Bluff beds.
Type.—Ala. Museum of Nat. Hist., University, Ala.
Remarks.—This shell is quite fragile, the body whorl break-
ing away from the spire. Prof. Harris figured the spire of this
species under Aesopus erectus (Proc. Acad. Nat. Sciences, Phila.,
1896, p. 476, pl. 21, fig. 3), which it resembles, but later on
states it isnotthesame. ‘The Ala. Museum has three specimens,
and there are two broken ones in my cabinet.
Type.—Ala. Museum of Nat. History, University, Ala.
TENUIACTAGN, n. g.
Shell resembling Actegon but much more slender ; whorls
nine ; aperture small, spire turned to the left and partially im-
mersed ; suture deep; columella with one fold ; surface with
numerous incised revolving lines.
Tenuiactzon pertenuis, n. sp. Ply 237) Eig: wo:
Shell small, slender ; whorls 9 ; surface with numerous re-
volving incised lines, stronger at base but almost disappearing at
the shoulder of body whorl ; then just below the suture showing
two or three stronger ones ; apex rounded and turned to the left,
partially immersed ; outer lip somewhat incurved ; base round-
ed ; columella with one strong fold ; the inner lip reflected over
the body whorl ; suture deep, giving each whorl the appearance
of being inserted into the next succeeding one. Length 13 mm. ;
breadth of body whorl 4 mm. ; aperture 5 mm.
17 ALDRICH, EKOCENE MoLLUSCA 17
Locality.— Gregg’s Ldg. marl at Bell’s Ldg. and Gregg’s
Ldg., Ala. River, Ala.
Type.—Ala. State Museum at University, Ala.
Teredo ringens, n. sp. JEM as) i saab ao
Shell substance thin, closed at the larger end like Auphus ;
body of shell with raised, rather acute rings parallel to each
other, and virtually at right angles to the longer diameter.
Other fragments are not so stongly marked. Length of type
‘specimen 37 mm. Average breadth 5 mm.
Locality.—Sucarnochee clay bed, 3 miles south of Estelle,
Ala.
Type.—Alabama Museum, University, Ala.
Tellina estellensis, n. sp. PEI aon | Deb vedsyy ney Yl
Shell small, exceedigly thin, exterior smooth, except fine
growth lines ; these are stronger near the ventral margin ; in-
equilateral ; right valve with small laterals, larger in the oppo-
site valve ; pallial sinus large, rounded ; anterior somewhat
pointed ; posterior rounded. Length 13-15 mm. ; height 9-11 mm.
Localityx—Sucarnochee beds, near Estelle, Ala.
Type.—Ala. Museum of Nat. History, University, Ala.
Remarks.—This species is the same figured by Prof. Harris
(Bulletins Am. Pal., Vol. 1, p. 182) asacast. It is rather com-
mon but so very fragile it was almost impossible to procure a per-
fect example.
Tellina cynoglossa Dall.
This is the common species in the Woods Bluff beds. It
seems to be only a variety of 7ellina subtriangularis Aldr., which
was described from half grown specimens, which is more regu-
larly rounded posteriorly and also lacks the fold seen in older
specimens. The full grown examples from Woods Bluff on the
Tombigbee River measure as much as 25 mm. in length
Tellina bellsiana, n. sp. Pay Rigs.) 15) LO:
Shell large, flat, thin ; nearly equilateral ; surface faintly
concentrically striate ; lines:growing coarser on the anterior end ;
18 BULLETIN 37 18
beaks very small ; the dorsal area strongly bent, pallial area gib-
bous ; teeth normal. Height about 24 mm.
Locality.—Both Gregg’s and Bell’s Landings, Ala. River,
Ala.
Type.—Ala. Museum of Nat. Hist.
Remarks.—One example in the Carnegie Museum. The
type is broken some, shell thin for its size.
Tellina semirotunda, n. sp. peas Maleate 1077,
Shell medium ; nearly equilateral ; anterior somewhat
pointed ; surface smooth, except lines of growth irregularly
spaced ; a slightly depressed area extending from beak to base
behind the anterior fold. Cicatrices rather large, are well
marked ; pallial area large ; cardinals bifid. Length 16 mm. ;
height 9 mm.
Locality.—Both Gregg’s and Bell’s Ldg., Ala. River, Ala.
Type.—Ala. Museum of Natural History.
Tellina semipapyria, n. sp. Vey ay) Dene, u's),
Shell medium, resembling 7Tel/ina semtrotunda nobis ; sur-
face smooth except growth lines ; pallial sinus large, partially
confluent with the pallial line ; only left valve known.
Locality.—Jackson’s Rockhouse branch. Woods Bluff beds.
Ala. Length 23 mm. ; height 9mm.
Type.—Ala. Museum of Nat. History.
Remarks.—Bears a strong resemblance to the Claibornian
species Tellina papyria Con. but is somewhat more rotund.
Semele langdoniana, n. sp. Jes mle, 16), AO:
Shell oblong, oval ; surface practically smooth, showing
very fine lines that are coarser on the umbonal slope ; some spec-
imens show also fine radial lines reaching to the ventral margin ;
posterior shortest ; ventral margin entire ; pallial sinus profound.
Shell looks like a 7el/ina externally. Length of largest spect-
men is about 27 mm.
Locality.—Bell’s Landing, Monroe Co., Ala. River, Ala.
Type.—(Left valve) Ala. Museum, University Ala.
Remarks.—There are several fragmentary specimens in the
19 ALDRICH, KOCENE MOLLUSCA 19
lot. ‘This species is named after the late Dr. W. Langdon, an
Assistant State Geologist of Alabama and the discoverer of the
Floridian Miocene.
Semele monroensis, n. sp. 2a REN SIS 2 22.
Shell small, rather flat ; one right valve only in the collec-
tion ; umbonal slope strongly bent ; surface smooth above with
several widely spaced lines towards the ventral margin ; hinge
stout ; ligament long for the genus ; the umbonal slope marked
interiorly by a few radial lines. Length 5 mm.
Locality.—Bell’s Landing, Ala. River, Ala.
Type.—Ala. Museum of Nat. History.
Remarks.—The specimen described is quite young but dis-
tinct. Looks like a Weretrix in shape and ornamentation.
Lucina primoidea, n. sp. RIM 2aNOnigSN 22h 2A.
Shell with many raised lines ; beaks recurved, higher than
long. ‘This species is figured because it is from an horizon close
to the Cretaceous and is doubtless an ancestral form of Luczna
cornuta Conr. of the Claibornian. The interior is not accessible.
Locality.—Black Bluff, Tombigbee River, Ala., about 20 feet
above the base.
Type.—My collection.
Martesia recurva, n. sp. RANI eS 5.20.
Shell small, short, rotund; with two accessory plates just
forward of the umbones ; groove running nearly vertical from
beaks to base and strongly marked ; concentric strize strongly
marked on the posterior end, bending abruptly at the groove and
thence running up behind the umbones and under the accessory
plates ; the anterior end rounded and smooth ; cardinal margin
bordered with a thickened and raised process. Length 8 mm. ;
height 4% mm.
Locality.—Fleming’s Mill, Ala.. on Pea River. Nanafalia
beds.
Type.—My collection.
Remarks.—This species is more obtuse than the usual forms.
Three examples found. ‘The interior is not accessible.
20 BULLETIN 37 | 20
Rochefortia minuta, n. sp. Pl 2tyibiigss 27.928
Shell, small, ovate ; nearly equilateral ; periostracum still
adherent, surface showing a few incremental lines ; beaks point-
ed, with a resiliary pit underneath with short lamelle on each
side, the posterior one the longest ; muscular scar slightly im-
pressed ; the right valve has a space for the left lateral ; pallial
line simple. Resembles somewhat Rochefortia Stimpsont Dall.
The space in our specimen under the beak looks as though it had
been excavated out of the dorsal side, but such is not really the
case. Prof. W. H. Dall has examined this specimen, and veri-
fied its generic position. Am greatly indebted to him in a num-
ber of cases. Length 3 mm. ; height 2 mm.
Locality.—Bell’s Ldg., Alabama River, Ala.
Type.—Ala. Museum.
Montacuta bicuspidata, n. sp. * Pl. 2, Figs. 29, 30.
Shell minute, oblong-ovate ; surface with incremental lines ;
beaks carrying the prodissoconch, nearly central ; viewing the
shell from the outside and above, it shows two tooth-like projec-
tions, one on each side of the beak, rising from the dorsal mar-
gin and another shorter one which does not show like the two
first mentioned ; ends rounded ; base arcuate ; hinge with a small
subtrigonal tooth and also a space for a lateral in the other valve.
Pallial line simple ; adductors distinct but not deep. Length 234
mim ; height 2 mm.
Locality.—Bell’s Ldg., Ala.
Lype.—Ala. Mus. of Nat. History, University, Ala.
Basterotia ? prima, un. sp. PLE aaatkigs amt anne
Shell small ; valves rather thick ; subquadrangular ; beaks
subterminal ; shell profoundly gaping at the anterior ventral
part ; a depressed space running from umbo to ventral margin ;
surface nearly smooth, showing growth lines which also show in
the interior. Anterior adductor scar deep, with a raised
border buttressed to hinge plate ; posterior scar slightly im-
pressed, pear-shaped and both rather large for the size of shell ;
one cardinal tooth inright valve, corresponding to a deep socket
20 ALDRICH, FOCENE MOLLUSCA 43
in the other ; a longer tooth below. Three single valves in the
collection. Length 8mm. ; height about 7 mm.
Locality.—Gregg’s and Bell’s Landings, Ala. River, Ala.
Type.—In Ala. Museum.
Remarks.—This species is doubtfully placed in Basterota ;
the large gap suggests an attachment to some other form, but is
not irregular. Prof. Dall has kindly examined it, and suggests
that it might be commensal.
Saxicavella alabamensis, n. sp. Rl ae igs a Aays.
Shell inequivalve ; thin ; posterior much broader than the
anterior ; outer surface showing irregular lines of growth,
stronger over the umbonal slope ; an upright cardinal tooth
directly under the umbo, with a short lateral on posterior side in
the right valve ; anterior muscular impression distinct ; pallial
line faint. Only one valve in the collection. Length 6 mm. ;
height 4% mm.
Locality.—Bell’s Landing, Monroe Co., Ala.
Type.—In Ala. Museum.
Montacuta Herberti, n. sp. Pee eS MOL 7.
Shell small; inequilateral ; dorsal and ventral margins
rounded ; each end rather blunt; the posterior somewhat
angulated ; surface smooth with very fine concentric striations,
and a few lines of growth near the ventral margin ; shell inflat-
ed ; cardinals feeble, showing in left valve only ; beaks low,
rather pointed, a small buttress under them. Length 9 mm ;
height 6% mm.
Locality—Cave Branch, Ala.
Type.—Ala. Museum of Nat. History.
Remarks.—One valve in my collection. Named in honor of
the late Herbert H. Smith.
Ervilia lignitica, n. sp. TMi gin! | TEAS toys
Shell small, inflated ; nearly inequilateral ; rather solid ;
the posterior slightly longer than the anterior ; both ends round-
ed ; surface carrying a few irregularly spaced grooves of growth;
interior polished, shining ; pallial sinus small, not reaching to
22 BULLETIN 37 22
center ; umbones small, dentition as in the genus with short and
strong teeth. Length 6 mm. ; height 3% mm.
Locality.—Both Gregg’s and Bell’s Landing beds, Ala.
River, Ala.
Type.—Ala. Museum of Nat. History, University, Ala.
| Remarks.—This species has a strong resemblance to A7vvzlia
polita Dall from the Floridian Pliocene. The young seem to be
higher in proportion to their length than the full grown speci-
mens. :
Panopea bellsensis, n. sp. AL Ss Seite, GO) 18) 6
Shell rather large, thin, with a brown epidermis ; lines of
growth irregularly spaced ; shell compressed, nearly equilateral ;
beaks small, appressed to and partly covered by the reflected dor-
sal margin; one large cardinal tooth ; ligamental attachment
short ; interior porcellaneous with growth lines showing through;
pedal scars small. Length 30 mm. ; height about 13 mm.
Locality.—Bell’s Ldg., Ala. River, Ala.
Type.—Ala. Museum of Natural History.
Remarks.—One valve and some fragments in the collection.
This species has its epidermis preserved. ‘The shell is extremely
thin and delicate.
Donax acutangula, n. sp. NG gy 2 IWikex, 102),
Shell nearly equilateral ; pointed at both ends, the umbonal
slope bent abruptly to almost a right angle in old specimens ;
surface nearly smooth ; fine growth lines showing ; ventral
margin entire ; beaks small, pointed, smooth ; hinge well devel-
oped ; laterals very long, striated on the inner edges. Length
about 31 mm. ; height 15 mm.
Locality.—Bell’s Landing, Ala. River, Ala.
Type.—Ala. Museum of Nat. History, University Ala.
Remarks.—One whole valve and several fragments of both
valves received.
Arca (Cucullaria) ozarkensis, un. sp. Bee enbigs nian eran
Shell small, rounded at both ends, giving the shell a quad-
23 ALDRICH, HOCENE MOLLUSCA 23
rangular outline ; beaks small, compressed and pointed ; surface
with rather close set radial lines on the posterior slope ; balance
of the surface smooth ; teeth as in the subgenus, three or four
anterior and two posterior ; muscular scars shallow but plainly
marked. Length 5 mm. ; height 3 mm.
Locality.—Woods Bluff beds near Ozark, Ala.
Type.—My cabinet.
Ostrea intermedoides, n. sp. Ne Debby ans sip
Shell medium, valves rather thin ; surface in older speci-
men with numerous raised concentric ribs ; the lower valve bent
downwards near the beak ; the umbo shows radial strize ; mar-
gins crenulated internally ; a fragment of the upper valve ap-
pears to be flat. Length of lower valve 32mm. ; breadth about
20 mm.
Locality.— Bell’s Landing marl, Bell’s Landing, Ala. River,
Ala,
Type.—Ala. Museum of Nat. History.
Remarks.—Have hesitated to add another oyster to our Eo-
cene, but I have been unable to find a place for this species.
Four lower valves and a fragment of the upper valve are in the
collection. The ornamentation recalls O. falco Dall from the
Jacksonian.
Psammobia Smiihi, n. sp. Ply sah iesa. l7, plo:
Shell large ; substance of shell thin ; oblong-ovate ; rather
compressed ; surface shining, with fine concentric sculpture
medially ; both ends coarser and elevated ; on the posterior
slope the laminze are rough and Somewhat mammillated ; beaks
very sinall ; muscular scars distinct but shallow. Length 60
mm. ; height 32 mm. ; Breadth about 12 mm.
Locality.—Gregg’s Ldg. marl at Bell’s Landing, Ala. Riv-
er, Ala.
Type.—Alabama Museum of Nat. History, University Ala.
Remarks.—Both valves of this splendid species are in the
collection. It bears a general resemblance to P. /i/osa Conrad,
but the dimensions are very different. Collected by the late H.
H. Smith and named in his honor.
24 BULLETIN 37 24
Psammobia harrisi, n. sp. Pl. 12.) Bigs..19,, 20!
Shell nearly equilateral ; truncate behind, with fine concen-
tric sculpture, coarser at each end ; ligamental area short ; cica-
trices distinct ; lateral tooth long ; posterior one short. Length
about 37 mm.
Locality.—Gregg’s Landing, Ala. River, Ala.
Remarks.—Vhis species resembles P. ozarkana Harris but is
more obtuse. ‘The type is not perfect.
€
DIVERSA
(Echinocyamus ?) meyeri, n. sp. PIS a Mies Ha 1 228
Test small, ovate ; flattened below ; deeply pitted with ovate
scars above which become nearly square on the sides and below ;
substance of the test of moderate thickness ; apicial system near-
ly central ; ambulacral petals relatively short, straight, with
numerous round pores. Peristome is medium circular, and
slightly depressed ; periproct not small, closer to the peristome
than the side. Length 5 mm. ; width 3% mm. ; height 3 mm.
Locality.—Gosport Sand, Claiborne, Ala.
Type.—My cabinet.
Remarks.—This species is peculiar in the deep pits covering
the surface. It is larger than &. huxleyanus Meyer and more
obese though egg-shaped. Received from Dr. Otto Meyer in
whose honor I have named it.
NOTES
BLACK BLUFF OR SUCARNOCHEE HORIZON.
Prof. H. H. Smith and wife collected a number of species
from these clays not far from Camden, Ala., in the vicinity of
Estelle, Ala. The deposit consists of an aluminous clay, dark
chocolate in color, and contains the greater part of the species
mentioned by Prof. G. D. Harris from the bluff at Fort Gaines
on the Chattahoochee River where they seem to be represented
principally by casts. Those near Estelle are rather numerous,
but very fragile. A few new species are described in this paper.
25 ALDRICH, HOCENE MoLLUusCcA 25
A surprising fact is that a minute Corbula, Corbula (Aloid:s)
milium Dall, originally described from the Wilcoxian is extreme-
ly common. ‘The supposed differences in the specific fauna is
explained. The two localities are over 100 miles apart. The
list of species from near Hstelle is as follows :
Leda saffordana Harris.
‘“ parva Rogers.
guercollis Harris.
mitlameensts ‘*
Corbula (Aloidis) milium Dall.
Glycymeris cf aviculoides Con.
Pinna sp. ?
Teredo ringens Aldr.
Yoldia eborea Con.
Venericardia wilcoxensis Dall.
Tellina estellensts Aldr.
Protocardia Harrist Dall.
Pecten alabamensis Aldr.
Ostrea ? young.
Nucula ovula Lea.
6c
6
Ce.
medtavia Harris.
Modtola saffordi Gabb.
Meretrix ripleyana Gabb.
Cucullaa saffordt Gabb.
i macrodonta Witt.
Crassatellites sepuicollis Warris.
a gabbi Safford.
Arca sp.?
Corbuls subcompressa Gabb.
Strepsidura fleilprint Aldr.
Volutilithes rugatus Con.
a limopsts Con.
Turritella tennesseensts Gabb.
i mortont Con.
Aumerosa Con.
Turritella alabamiensts Whitt.
be
BULLETIN 37
Triton showaltevz Con.
Solarium periscelidum Dall.
Solariclla alabamensis Ald.
Rissoina alabamensis Ald.
Pseudoliva unicarinata Ald.
ng scalina Heilpr.
Pleurotoma quercollis Harris.
i persa Whitt.
meedtavia Harris.
os adeona Whitt.
longipersa Harris (var.)
Calyptraphorus velatus, var. compressus Aldr.
Olivella mediavia Harris.
Natica saffordia? **
‘¢ reversa Whité.
onusta a
eminula Con.
Limulany, Wes
Amaura tombigbeensis Harris.
Mesalia watsonensis Harris.
‘(pumila Gabb (var. )
ae alabamiensis Whitt.
Levifusus pagoda Heilpr.
: dalei Harris.
Fusus quercollis Farris.
PN NOLLOMES EN Gite
Coll coneeles NN ad he
‘< —mohrt Aldr.
Dentalium mediaviense Harris.
Cadulus turgidus Mr.
Atys robustordes Aldr.
Tornatella quercollis Harris.
Cylichna meyert Aldr.
Gilbertia estellensis Aldr.
Cypre@a 2 sp.
a
29
27
ALDRICH, HOCENE MOLLUSCA
Exilia pergracilis Con.
Nautilus sp. Fragments.
Flabellum conoideum Vaughan.
Balanophylla haleana M. Ed. & H.
Stenocyathus n. sp.
Foraminifera, 12 species.
Figure
W7h
19,
Zip
23,
25;
27,
~
Il.
BULLETIN 37
EXPLANATION OF PLATE 1
28
Page
Mitramorpha turriculata, n. sp., Choctaw Cor, Length6mm... 5
Turris MODOLACUSMMENSD AN Gave ranle bate iecen in eneenette ae gimm... 5
TUIGUSESPECH STIS DM CAN CM OBTA TCH ewer sere eeaititatsasesee 8mm... 6
Turris bimoniatus, n. sp., Cave Branch..........2.. sese..0 14mm... 7
Pleurotoma ( Peratotoma) Gardneri, n. sp., Pugh’s Br. 21 mm... 8
Odostomia (Evalea) Barischi, n. sp., Bell’s Ldg............ Bi youbodlane |) tS)
Turbonilla (Ptycheulimella) clinensis, n. sp., Gregg’s
A Bfelr aX 6 Vester aunt aA Hae aa aaa iat Apa mtn 3mm... 9
Turbonilla (Cingulina) tuscahomensis, n. sp., Gregg’s
I Brahals boa bea Waa Re aS MOUNTS Wn A aa eUO Ue Plame Pa HSIN a ar 4mm... 9
Turbonilia ( Ptycheulimella) tardiusculus, nu. sp., Cave
Ba ee a Shaan Wa LE alate aid sabia a Meet ona 4.5mm... 9
Pyramidella ([phiana) obtusoides, n. sp., Gregg’s Ldg. 3.5mm... 10
Epitontuim muntstriatum, n. sp., Jackson’s Rock House
UB} of 010) a Maaienieai aM MU ant Ore RE cS a ERIE 4mm... 10
Epitonium munistriatum, n. sp., Gregg’s Landing ..... 3mm... 10
Epitonium multiliniferum, n, sp., Cave Branch......... iomm... II
Epitonium subacutum, n. sp., Cave Branch...............- Io mm... II
Teinostoma subangulata Mr., var. Smithi, n. var.,
Bell Sy arra dani patie auetun wp iauabamacuinsta at blu ca uionuiaualyan 8 mm... II
Teinostoma regularis, n. sp., Pugh’s Branch..............+. 4mm... 12
Anctllopsis Tuomeyi, n. sp., Bell’s Landing............... 28 mm... 12
IN ASSO PICOM ENTS D Aly ISON Meni ceeee een ieee eae 12+ mm... 12
Ohwvella semilienitica ims spy Clarkin Con ieee ison 5.5mm... 13
Cancellaria Gilberti, n. sp., Gregg’s Landing...............9 mm... 13
Ovula regularoides, n. sp., Bell’s Landing........... 2... 14mm... 14
Sy;
a
a
So
>
é @ %
17
Za
Meets
megs
29
ALDRICH, KOCENE MOLLUSCA
PLATE 2
(2)
ey.
T5,
27;
29,
28.
30.
BULLETIN 37
EXPLANATION OF PLATE 2
30
Page
Gypr aa estellen sts. Ne Sp) \eursleya Chascsscceseecensetesres ce 12mm... 14
@ypraaispybursley | Cree iece you eee enestececee ee sseeeeeee I5 mm... 15
Siphonalia quadrilineata, nu. sp., Pugh’s Branch......... Gp scobadlinn 105)
Gilbertia estellensis, n. sp., Estelle.................eeeeeeee 2.5x2 mm... 13
Levibuccinum (Euryochetus ?) harrisi, n.sp., Cave
Branch); largest specimens wscerse.c ses -ssasses io 4.5 mm... 16
Tenutacteon pretenuis, n. gen. and sp., Gregg’s Ldg....13 mm... 16
Gerithtopsts esicllensis. 1.) iSps. ee Stelle enn accneee seeseceeee 3mm... 15
IRA AHO) (HISTALS oo VYSUAVS ooecaodoabbacnasooseoya, oaticased 37 mm... 17
Tellina estellensis, nu. sp., Estelle......... largest sp....I5XII mm... 17
Tellina bellsiana, n. sp., Bell’s Ldg.; height.............. 24mm... 17
Tellina semirotunda, n. sp., Gregg’s Ldg. ; length......16 mm... 18
Tellina semipapyria, n.sp., Jackson’s Rockhouse
Branchy wen oth cove sssucesaveseecscer sone eeee ees 23 mm... 18
Semele langdoniana, n. sp., Bell’s Ldg.. ..............060++ 27 mm... 18
Semele monroensts, nu. sp., Bell’s Ldg...........ceeeeeeeeeee ee 5 mm... 19
Lucina primoidea, n. sp., Black Bluff................006 9x9x6 mm... 19
Martesia vecurva, n. sp., Fleming’s Mill, Nanafalia
| OY e10 USAMA AL a eGR Sc acy 8x4.5 mm... 19
RROCHE/OV UG IHINULAA NeSp (Bells ie daaauee ses ee eee ere eee 3mm... 20
Montacuta bicuspidata, n. sp., Bell’s Ldg............... 2x234 mm... 20
Vol. 9, Pl. 2
Paleont.
Amer.
Bull.
Pl. 2, No. 37
7;
MY
Zi <
Ky
_
<7
<
7 Tis. : )
See OZ
Coe
7
ui ae
ria
wy)
cae
wr
ho Sy
leat
31
ALDRICH, EOCENE MOLLUSCA
PLATE 3
(3)
aI
20.
22.
BULLETIN 37 32
EXPLANATION OF PLATE 3
Page
Basterotia ? prima, n. sp., Gregg’s Ldg.................--- 8x7 mim... 20
Sartcavella alabamensis, n. sp., Bell’s Ldg.......... 6x4.5 mim.... 21
WIONLACULA ELCLOCH LIM SPs CAVE MD ae tae etre eens ee sass ee g mm... 21
Eyvilraligniieca.) Te Spi Belly Siy i, digmees | ean eseicusin: senate 6mm... 21
Fanopem bellsensts, 1. Spi Bells ide ys eee esa 30X13 mm... 22
Donax acutangula, n. sp., Bells’? Ldg............ ee. 31XI5 mm... 22
Arca (Cucullaria) ozarkeusis, nu. sp., near Ozark...... 5X3 mim... 22
Ostrea intermedoides, n. sp., Bell’s Ldg. No. 15, 24x19 mm.
No. 16, 32x22 mim... 23
Psammobia Simitht, n: sp., Bells Wde. tes 60x32 mm... 23
Psamunooia harrvist,)m.\sp., Gregers dere ce sale cesens 37 mm... 24
(Echinocyamus ?) meyer, n. sp., Claiborne...... 5X342x3 mm... 24
Vol. 9, PI. 3
Paleont.
Amer.
Bull.
Pl. 3, No. 37
Ce
<—
Bo
te
a
hoe ;
ce
nes
BULLETINS
OF
AMERICAN PALEONTOLOGY
soy TE
Vol 9
sk
No 38
RECENT MOLLUSCA OF THE GULF OF MEXICO AND
PLEISTOCENE AND PLIOCENE SPECIES FROM
THE GULF STATES
Part 2
Scaphopoda, Gastropoda, Amphineura, Cephalopoda
BG:
CARLOTTA JOAQUINA MAuRY
January 31, 1922
[Part I, PELECYPODA is Bulletin No. 34, 1920. ]
Harris Co.
Cornell University, Ithaca, N. Y.
Wi SeAw
i
My)
a
A MOLLUSCA OF THE GULF OF MEXICO 24
GWASS V/SGNAai@ @ibDA
ORDER SOLENOCONCHA
Genus DENTALIUM Linnzeus
laqueatum Verrill, Trans, Conn., Acad, Arts Sci., vol. 6. pt. 2,
Pas pl 44. fi 18) 18857 Walli Bull} Musi Comp. Zool:
EMO D AZO DI ts Tresor) Bill ara WLS A Wine.
GIS ypu TOM ply 27.) fn pl AG. li Tey) T8SOr.bilsahyiand
Shanp, yinvyon s Man. Conch.,.17) ps ro, (pli 70 fa\ta2-) pluis)
Peso?) tlenderson,, Bulli ci0.2U. Ss. Nat. Maas. p. 23;
PAT te Oni'7,), 1O2O.
Distribution—Hatteras to Barbados, Gulf of Mexico: West
Florida and off the Mississippi delta, 60-68 fms.
texasianum Philippi, Zeitschr. Malak, p. 144, 1848; Pilsbry &
Sharp, Tryon’s Man. Conch., 17, p. 22, 1897; Henderson,
cr OR SaNe VE 1.) 28 nile Enea 5 Oe, LO2O.
sexangulare Hilgard, House of Rep. Ex. Doc. 1, pt. 2, pp.
37,1889, pl. 3, f. 7, 1878. Not of Lamarck:
gouldiz Singley, 4th Ann. Rept. Texas Surv., p. 331, 1892;
Dall, List Cameron Sh. Not the true gould7? of Dall.
Note.—This may be identical with Chenu’s 2. americanum
1843. Closely related to gouldzz Dall, with which it has
been confused.
Distribution.— N.Carolina to Yucatan, 3-9 fms. Gulf coast:
Point au Fer and Cameron, La.; Galveston, Corpus Christi
and Pass Cabello, Tex. Pleistocene, Lake Borgne bor-
ings, La.
texasianum cestum Henderson, Bull, 111, U.S. N.M., p. 29,
DIN Ark MAA. 8) (O20:
Distribution.—Pass Cabello, Texas. Probably in very
shallow water.
rebeccaensis Henderson, Bull. r11, U. S. N. M. pp. 16, 31,pl. 3,
LN TO2@)
5 BULLETIN 38 35
Distribution.—West Florida, 7-16 fms. and the Tortugas.
(Anialis) antilarum d’Orbigny, In de la Sagra’s, Hist Pol. y
Natyisla deliCubal 2iipszo2t plist Ota reroll site
& Sharp, Diyouis) Man. Conch) 17). 57, ple) 14) f 22-25)
seven Jealssavaleqsropal,| JEU Wessse VOL, Sy ONY VES! oy Au jolly. 96, t=
4, 6-8, 1920.
Note.—Confined to shallow water. Deep water citations are
reterable to different species.
Distribution.—Florida to Barbados. Gulf Coast: Charlotte
Harbor and other stations off West Florida, 2-14 fms.
(Antalis) pseudohexagonum Ihering. See Henderson Bull. 111,
Ue SNe Mee py oy ple Gu tet- 2 192e)
Distribution.—Florida to Brazil. Gulf coast: Marco, Tampa
and Charlotte Harbor, West Florida, 2-5 fms.
(Antalis) disparile d’ Orbigny, Hist. Pol. y Nat. Isla de Cuba,
2. Puzo plicit lati lo4o: (ci Vianattay) Proc mncade
Ne SiPhilaissi) pe7isi741 19033 taenderson, Bulle now WUEioe
ING Meo ae pln On hac on i O20:
Note.— Frequently confused with D. aztillarum by authors
althougn really closer to dexasianum.
Distribution.—Florida to Rio de Janeiro, Brazil. Gulf
coast; Tampa, Fla. Littoral zone.
(Antalis) ceratum Dall, Bull. Mus. Comp. Zool., 9, p. 38, 1881;
TOGO US. 0, MAM OL AG). Wel I ONL, \ Ag) sts yi atshexoy Dowoudly a), U).
SNE ME pe On pl 20 Ss TOSOn me enderson is allay mune
Sa) NE) Meipper49-514) ploy. hf. 227, 1920:
Distribution—Florida to Barbados. Gulf of Mexico: Off
Mississippi delta, 321 fms.; Yucatan Banks, 84 fms.
(Antalis) taphrium Dall, Bull. M. C. Z., 18, p. 422, 1889; U.S.
N.M.,p. 76, 1889; Pilsbry & Sharp, Tryon’s Man. 17, p. 58,
L8O7 lenderson Bulltaian Ue SaiNEwver Nps 260 pluie ies
1920.
36 MOLLUSCA OF THE GULF OF MEXICO 6
Distribution—Hatteras to Cuba, 22-182 fms. Type dredged
between Mississippi delta and Cedar Keys in 30 fms.
(Antalis) bartietti Henderson, Bull. 111, U.S. N. M., p. 55, pl.
CMH hy LOZO:
rmonliagine Dall Bulle Me ICA eZ Ni'o.)paig7s) USSt + Bullaia 7.
eS SINE MO Dp, 76) (pars), “1889.) Not) anilarum)) a Or-
bigny, 1846.
Note.—True antidllarum is semi-littoral while déarflett7 is in
deeper, colder waters.
Distribution.—Florida to Martinique. Gulf of Mexico: Be-
tween Mississippi delta and Cedar Keys, 227 fms.; Yucatan
Banks, 399 fms.
(Heteroschisma) cailithrix Dall, Bull. M. C. Z., 18, p. 427, pl. 27,
LOMO CO MP oUluia 7. Sa Na Me pio, plea 7Mit:) TOmisse:
ilsbryce Suarp, Uryons Many Conchin7)) p62, ply 7, f.13;
Noone Elenderson, Bully 111) Uo S4) Ne Mi p57, plewsa fs 3,
dks On) MOO’
Distribution—North Carolina to off Rio de la Plata, Ar-
gentina. Gulf of Mexico: Between Mississippi delta and
Cedar Keys, 1181 fms.; Yucatan Strait, 640 fms. (type lo-
cality).
(Fissidentalium) meridionale Pilsbry and Sharp, Tryon’s Man.
Conehy17) 99.73, ple15, £. 32-34) 1897, \(as variety of ca7z=
didum): Henderson, Bull. 111, U. S. N. M., p. 61, pl. 9, f.
2TO2O:
Lona sep alle alle NE KOCH) Z-6,) 9: 37.) kool iLO Ps) 4:25.00) LOOOs
Proc. U. S. N. M. 12, p. 294, 1889. Not ceras Watson.
Distribution—Type dredged by the Albatross, 240 miles south-
east of Rio de Janeiro, 671 fms., 37° temperature bottom.
Also found Yucatan Banks, 1568 fms. and off Barbados.
The largest Dentalium of the western Atlantic and charac-
terized by its unique, ashen color.
7 BULLETIN 38 37
(Grapiacme) eboreum Conrad, Proc. Acad. N.S. Phila., 3, p._
27, 1846; Pilsbry. and Sharp, Tryon’s Man. Conch., 17, p.
89) pl: 16,\f.)'47-405)) 55, 56) 1897,;Ekenderson))) Ball) irae
NGM ph OO. pl MO NEM sH5 4) SiO mip lamin canon ZO
leptin) Bush ihrans, Conny Acad aw ArtSsGandsS cla wisere nite
CD NAT OmpIUA SMe Sh Tea iT SS Seii Daly tele aa an Ole
NMP pO LS!
matara Mal OB ME COZ 3h pa AZO TS SO. 1 Ellas
WS ceN MOMS So)
Distribution.— Hatteras to Barbados. Gulf coast: Sarasota
Pass (type locality), Charlotte Harbor, Fla. and between
Mississippi delta and Cedar Keys. Beach to 111 fms.
Note.—Mr. Henderson unites matara and Jleptum with
eborcum, noting that szafara is the perfectly smooth adult
form of the species.
(Grapiacme) semistriolatum Guilding, Trans. Linn. Soc. Lon-
doniin7 pty ips s4vplagi tet os read Chemin Comehemldile
De Ty pia i iain Set itendersons, Hull mre Opn Sener
Me piOG yp lenin stank 54S MO n O2@:
Distribution.— An abundant littoral species. Florida to
Barbados. Gulf coast; West Florida in shallow water,
7-12 fms.
(Graptacme) sericatum Dall, Bull. M. C. Z., 9, p. 37, 1881; 18,
DV AZZ DL ZO MTs son a dll ary MONS. ING IMs: i mii oruple
2On eT SSO. VElendersony ME mile MNT Oe IN an Veep Deals
PMI TA OM O2Or
Distribution.—Type only found. Dredged by the Alake, 640
fms., Yucatan Banks.
(Laevidentalium) callipeplum Dall, Bull. M. C. Z., 18. p. p. 410,
OLN AAS Saaceon = CMa ifereayanies) IBY EVOL) ak AUS AN SI INI dy bas Ob 6) Ol! 2347
fyi 2'O) Lacon lrans: | \Viaoner wimSt Cia 2) \p. (442 1sQoR
Pilsbry and Sharp, Tryon’s Man. Conch., 17, p. 100, pl. 19,
38 MOLLUSCA OF THE GULF OF MEXICO 8
MPOMnso7)) Henderson) Bullen) (Ws) S\N Mpa 4. pls) 12),
“its (Gy) OPee
Distribution.—Florida to Porto Rico. Gulf of Mexico,
dredged to 169 fms. between Mississippi delta and Cedar
Keys, (type locality).
(Laevidentalium) perlongum Dall, Bull., M. C. Z., 5, p. 61, 1878,
Women nUudure) Gy 9) 26.1883) 18). Anos pler27 fs, 6,
MoCo Bulls7 US. Ney Mes ps6) pla 27.) On Taso blen=
derson Bull ir1r US) N2 Meio 75.0 pk. (o,f. 1) 1920:
Distribution.—Hatteras to Rio de la Plata, 11—1330 fms.
Gulf of Mexico between Mississippi delta and Cedar Keys
at 940 and 1181 fms. also elsewhere in the Gulf at 1330
fms.
(Episiphon) sowerbyi Guilding, Trans. Linn. Soc. London, 17
pass fy 7a T8345) Pilsbry and) Sharp,)nyonrs, Man,)
Counc ptr 7.) pl. 20, f..2om1so7> blenderson)) Bull. 111
We SHON VE pps 77-80, Diya ity 2a LOwTO2@:
Za Dally VB ww 37 Ue Si Ne ME pal 76 T88O.) | Pro:
bably not f/um of Sowerby, 1860, which was from the east
Atlantic.
Distribution.—Hatteras to Barbados. Gulf of Mexico: Off
Mobile, Ala., 27 fms.; between Mississippi delta and Cedar
Keys, 60 fms.
sowerbyi pelliceri Henderson, Bull. 111, U. S. N. M., p. 80,
IE NEL 7S) Out O2 0:
Distribution.— Florida to Cuba, Cape Catouche, Yucatan,
25 fms.
(Bathoxiphus) ensiculus Jeffreys, Ann. and Mag. Nat. Hist. ser.
ERO MOU SAN LS 7h: Me hOC. LOOM SOC PE OOOW plan 4Qute0 4);
1882. Watson, Challenger Rept., (Scaphopoda), p. 12, pl,
Zeb 2 Sos. LOall: (Bully 370 Ul Sa Ne Me py 7On18 36.
lenderson, Bullieriny Ws oS: Ney Vi pa Stanplanie iat 14.5,
TOW LQ2O:
9 BULLETIN 38 39
Distribution.—East and West Atlantic. A deep.water
species of wide range. Gulf of Mexico: Yucatan Strait,
640 fms.
(Bathoxiphus) didymum Watson, Jour. Linn. Soc. London, 14, p.
517, 1879; Henderson, Bull) 11.0. S. NM. p. 83) spline
10, MOKA,
Distribution.--Culebra Isl. and Barbabos. Doubtful specimen,
Yucatan Banks, 25 fms.
(Compressidens) pressum Pilsbry and Sharp, Tryon’s Man. Conch.
TiO w 2A Ole mut. MIMD 22a NGOs 2h ed SO). Wlendersone
Bull nee NG Mi ons Zep lealaaien ss soo, sLO2O.
compressum Watson, Jour. Linn. Soc. London, 14, p. 516, 1879;
DallsyBullwMe C7 Zo passa rons Watson, Challe kept
(ScaphQeip: 0, pla at] Guinges malls ull 27. US. Niner
p. 76, 1889. Not compressum of d’Orbigny, 1850.
Distribution.—Florida to the Antilles. Gulf of Mexico:
Off Cedar Keys, 196 fms.; between Mississippi delta and
Cedar Keys, Fla.
(Compressidens) ophiodon Dall, Bull. M. C. Z., 9, p. 38, 1881; 18,
D427. pl 26) OE S8o; MBLIllNe TORS. UN Melt oe 7G ple
ZO tO 1889" Itenderson,, Bulle minim Us So. NY) Met) Saki:
WA ATOZ @.
Distribution.—Florida to Barbados. Gulf of Mexico: West
Florida, 169 fms.
quadrangulare Harris, Bull. Amer. Paleontology, vol. 1, p. 95,
1895. Upper Miocene, Galveston well at 2465-2871 feet
(Harris).
tetragonum Sowerby, Thesaurus Conch., 3, p. 103, pl. 224, f.
21, 22, 1860. Upper Miocene, Galveston well, at 2158-
2920 feet (Harris).
Remark.—A fragment of an undetermined, Pleistocene, Den-
talium was obtained from Knapp’s No. 2 well, Terrebonne
Parish, La. at 1190-1430 feet.
A0 MoLLUSCA OF THE GULF OF MEXICO 10
Genus CADULUS Philippi
(Polyschides) quadridentatus Dall, Bull. M. C. Z. 9, p. 36, 1881;
SOM AZo Plit2 7 i 5 rssory BULLEN a 7U) 1) SIN Mp.) 76;
Plea Tat. Su 889) lenderson, Bull) iin Wh) SNe) MES pp 97-
TOOM MNT 7a the 221) LOZ!
toms bush. Vln: Conn. Acad.,\6) p. 471.) pl 45, f, 20) 18855
Dalal sree Sy NW IME pkigGh plo An. st) 201 8 5o
Distribution.—Hatteras to Rio de la Plata, 3-52, fms. Gulf
of Mexico: West Florida, 10-25 fms.; Yucatan Banks 25
fms.
(Poiyschides) carolinensis Bush. In Verrill, Rept. Fish Com. (for
LISA PSST LSo54 Li Conny Acad’) 6.0 p. 47) Pla 5) be
Momissse Dall Bull: 27 Ua SuiNe Mp. (48 (pars). ply au,
f. 19, 1889; Pilsbry and Sharp, Tryon’s Man. Conch., 17, p.
m2 pl. 25, 4. 64) pli 30, f. 241-127), 1898; Henderson, Bull!
ORO EL SMN EUV) Os 102) Ply mhz Mt eNO) LOZO}
Distribution.—Hatteras to Key West, 3-87 fms. Gulf of
Mexico, off Mobile, in 27 fms.
(Platyschides) poculum Dall, Bull., M. C. Z,, 18, p. 429, 1889;
Ble 7 Oo. Ne MES pp) 765 1889 5) Pilsbry and) sharp;
direyonys Vian. Conch 17. p41 72, Dleisgeten 50, 57) L398) 5
itenderson, Bulli, U.S: Ni Mey py 10s pli nz, 1.18, 1920.
Distribution.— Type found off St. Vincent, W. I., 424 fms.
Aso dredged, 640 fms., Yucatan Banks.
(Platyschides) parvus Henderson, Bull. 111, U.S.N. M., p. 113,
OTS hor 52.04) LO20.
amtantus Dall, Bull. 37, U, S. N. M., p. 78, 1889. Net Dall’s
typical amzantus.
Distribution.— Off Barbados, too fms. (type); Yucatan
Banks, 1002 fms.
(Platyschides) elongatus Henderson, Bull. 111, U.S. N. M., p.
LAB. ‘solo WO, sey sy sO VAOY
II BULLETIN 38 4r
Distribution.—Gulf of Mexico, off the mouth of the Missis-
sippi River, 68 fms.
(Platyschides) arctus Henderson, Bull. 111, U. S. N. M., p. 124,
DILLON LO MmmOZOs
Distribution.—Gulf of Mexico off Cape San Blas, Florida,
60 fms.
(Gadila) mayori Henderson, Bull. 111, U. S. N. M., p. 133, pl.
LOW Lao LOZ)
Distribution.—Florida, Gulf of Mexico, off Cape San Blas,
Fla., 60 fms.
(Cadulus) transitorius Henderson, Bull. m1, p. 143, pl. 19, f. 6,
1920.
Distribution.—Florida to Old Providence Isl.,north of Colon.
Gulf of Mexico, off Cape San Blas, Fla., 196 and 169 fms.
42 MOLLUSCA OF THE GULF OF MEXICO [2
GOMASS GINS @ 7 @ aX
ORDER PTEROPODA
Genus CAVOLINA Abiidgaard
inflexa Lesueur Nouv. Bull. Soc. Philom., 3, p. 285, pl. 5, f. 3,
Iok(aseeyalca). Dalit Bulle We S Nati Mus.) (37, p. 82,
LOCO ETOC. Ueno. Nate MUS i 137.\\0.) 1.7) LOIO:
Distribution.—West Atlantic, Lat. 42°N.—42° S., Hast
Pacific, Lat. 13° N.—42° S. Gulf of Mexico, West Florida.
Pelagic. Pleistocene to Recent.
uncinata Rang, in d’Orbigny’s, Voy. Ame’r Me'’r., p. 93, pl. 5,
MEME TS 26.8 (as Eyalea) Dalley Balle 27 WU AS: Ne Mars,
MSZ Mlle G6, hy) 1lO} 18803) brocwU) SN Mius: 375 pa t98,
1910; Johnson, Occ. Papers, Boston Soc. Nat. Hist., 7, p.
156, 1915.
Distribution.—West Atlantic, Lat. 40°N.-40°S. East Pacific,
Lower California to Ecuador and Galapagos Isls. Gulf of
Mexico; West Florida. Pelagic. Pleistocene to Recent.
(Diacria) trispinosa Lesueur, in Blainville; Dict. des Sciences Nat.
227 82, 1821, (as Ayalea); (Gould) inv. Mass.,) p: 504,
ms7ou(as Yzacra); Wall, Bull. 375% pi'82, pl. 66) f)115,, 1889,
Sumner, Bull. Bur. Fish., 31, pt. 2, p. 703, 1913. Johnson,
Oces Papalbosk. SOC47,) DP: 15 7, LORS:
Distribution.—West Atlantic, Lat. 60°N.—4o° S. Gulf of
Mexico, West Florida. Pelagic. Pleistocene to Recent.
Genus CLIO Linnceus
pyramidata Linnzeus, Systema Nature, 12th ed., p. 1094, 1767;
Dalle Bulbe7. Wie SevNG MN peahsonmTssoy. (as Clcodora):
Broce Se Ne Mea 7p. 198) TOLOwmjonnson: (Ocei rap:
BOSH SOCK 7a Dulisou LOLS.
Cleodora lanceolata Souleyet, Zool. Bonite, 2, p. 179, pl. 6, f.
17-25, 1852.
Distrib ution.—Atlan tic, Spitzbergen to Lat. 40° S. South-
3 BULLETIN 38 43
eastern Pacific, Lat. 27° S. Gulf of Mexico, West Florida
and Texas. Pelagic. Pleistocene to Recent.
recunva) Children,’ Jour//Roye\lust.;) 15). 220, pli, fa 107.) 1820)
(as Balantium); Dall, Bull. 37, p. 82, 1889; Johnson, Occ-
Papi BOsts! SOC 7 i ps lon LONss
Distribution.—Lat. 40° N.—33° S. Gulf of Mexico, West
Florida. Pleistocene to Recent.
Genus CRESEIS (Rang) Sowerby
subula Quoy and Gaimard, Ann. des Sci. Nat., ser. I, 10, p. 233,
pls Dy hea ins274 \(asi@/eocoxa) Walla Balle 27. paoos
1889; Dall and Simpson, Bull. U. S. Fish Com. for 1900, 1,
Dp: 360, (plNS 7) fo LOOP DallivenocwWES. INEM 37s.
LQLO) Johnson. Occ) Lap.) Bost Soc 7.) p- 158s LOLS) (as
S7yleola) Dall Ball.) 12) Wey SeeNataiMiidsy.0p4) 5O.y1o2ie
Distribution.—Atlantic, Lat. 41° N.—40° S. Type locality,
Teneriffe. Pacific, Peruvian. Gulf of Mexico, West
Florida and Texas. Pelagic. Pleistocene to Recent.
Genus STYLIOLA (Lesueur) Gray
virgula Rang, Ann. Sci. Nat., ser. 1, 13, p. 316, f. 2, 1828, (as
Greseis): Verrill, Trans. (Conn: “Acad.,5, -p. 557, 1882, (as
styizola) Dall) Bull iag Ue SssuNe oe MiuS.. )\ 1). 80.) 7 Sonntas
Gresezs) (Proc. US. INE MES 27ers 18, LOLO)(as”.Sty/zola))e
Johnson, Occ. Pap. Bost. Soc., 7, p. 157, 1915, (as Cvesezs).
Distribution.— Atlantic, Lat. 41°N. - 35° S. Pacific, off Juan
Fernandez Isl. Gulf of Mexico, West Florida, Pelagic.
Pleistocene to Recent.
Genus VAGINELLA Daudin
chipolana Dall, Trans. Wagner Inst. Sci.,3, pt. 2, p. 431, pl. 23, f.
4, 5, 1892.
Lower Miocene, Chipola beds, Bailey’s Ferry, Calhoun Co.,
Florida. An ancient, Cretaceous to Recent, genus.
44 MOLLUSCA OF THE GULF OF MEXICO 14
Genus CUVIERINA Boas
columnelia Rang, Ann. des Sci. Nat., ser. 1, v. 12, p. 323, 1827,
Atlas 4°, pl. 45 B, f. 1-8. (as Cuvierta); Pelseneer, Challen-
gen) Eteropoda, Zool.,.22 art.) 1, p. 67, Teese; Dall Buall.27, Ul:
Soe Marsh) 182) i ply 66s) faint Son na Waser inst:
Sew synpte 2, ps 4ar i T892s Peck. Proc) U's) Nii Mus.) 16;
PAAOe Less ty ito. re9ge! Dall Proc.) War Su Na Mie 7) 1:
KOS) TOLO; Jolson) Occ Pap. Bost..S0C!)) 7p. L595 LOLS:
Triptera columnella Verrill, Trans. Conn. Acad., 5, p. 557,
L382) 62 py 2iA, 1884:
Distribution.—West Atlantic, Lat. 43° N.—4o° 5. East
Pacific, Lat. 23° N.—42° S. Gulf of Mexico, West Florida
and Texas, recent. Lower Miocene Chipola bed at Alum
Bluff, Calhoun Co. Florida.
/
Note.—The specific name is often, but erroneously, written
columella. "The correct form is colummnella since Rang prints
it thus twice in the original description. The generic name
Cuvieria is preoccupied.
Genus EMBOLUS Jeffreys
inflatus d’Orbigny, Voy. Ame’r. Me’r., p. 174, pl. 12, f. 16-19,
HSZ6) (asdvanta)- Wall, Bull) 27 UK SaiN) Musi p. 80,
MSOOMETOC Wis. Ni MEWS: 27) pa lOom Oro; ounson,)|Oce:
Rap BOSt. SOC.) 7.) p) 155, 1915. (as “2zveacia)).
Spirialts rostralis Verrill, Trans, Conn. Acad., 6, p. 431, 1885.
Distribution.—West Atlantic, Lat. 42° N.—40°S. East
Pacific. Lat. 42° N.—40° S. Gulf of Mexico, West Florida.
Pelagic, Pleistocene to Recent.
Genus PERACLE Forbes
reticulata d’Orbigny, Voy. Ame’r, Me’r., p. 178, pl. 12, f. 32-
C0 Jie
BSsO, oso. (asia Hana) Dally Bulle 27a UL Sa Ne Muss
PMCs sO roe Weis.) Ne MUS! Nam Ds nO LOLO:
15 BULLETIN 38 45
Distribution.—West Atlantic, Lat. 37° N.—o° S. East
Pacific, Lat. 20° S. Mediterranean. Gulf of Mexico, West
Florida. Pelagic. Pleistocene to Recent.
ORDER OPISTHOBRANCHIATA
Genus ACTAESN Montfort
punctostriatus C. B. Adams, Jour. Bost. Soc. Nat. Hist., 3, p.
3235 pli 3; £70; 11840, \as) Lozvatela)\ i Dally uli) 274 WER.
ING Mus: ((p. 7840) ipl. (41 is A ols 2a 22 Sori dlbiey NVialon er
Inst. Sci., 3. p. 14, 90; Singley, 4th Ann Rept. Geol. Surv.
Texas, p.)332, 92; Pilsbry,) ainyon’s) Man.) Conch! sem im
TS) DN 157, Dl Tet OS, OR plano 22" 22 0 co2.\)Dalilvannad
Simpson, Bull: U.S. Fish Come an). 262)! 1901. Johnsons
OcePapers) Bost) Soc (Ne Lie omar) NOS:
punctatus d’Orbigny, In La Sagra, Hist. Pol. y Nat. Isla de
Cuba, 1, p. 230, Atlas, pl. 17, f. 10-12, 1842; (as Zornatella):
Dall Proc. US) Ne Me Ole pa e247 r88iae Not 1a pacar
Ferussac, 1823. Not of Lea, 1833. Not of Piette, 1855.
cubensis Gabb, Tr. Amer. Phil, Soc., 15, p. 245, 1873.
Note.—The recent Cuban Acteon punctatus d’Orbigny
(preoccupied, renamed cubenszs by Gabb) is now considered
identical with the northern punctostriatus. Act@on punc-
tatus Wea) (Contr (Geoltio. jiammiaply 4st. (96) 1s), Claiborue
Eocene, Ala.; and, fide Dall, Pliocene; Caloosahatchie beds,
Fla. Not Recent.
Distribution.—Cape Cod to the Antilles, 2-63 fms. Miocene
(of Shiloh, New Jersey) to Recent. Gulf coast: Cedar Keys,
Fla,, Corpus Christi Bay, Tex. Pliocene, Caloosahatchie
beds, Fla.
exilis Jeffreys, Ann. & Mag. Nat. Hist., 4th, ser., 6, p. 85, 1870;
Watson, Chall. Rept. Gastr., p. 624, ’86; Dall, Bull. M. C.
Lp wharve Coll i 8i ipa gon .SOsmotillen a7 Onna No Mie anon
46 MOLLUSCA OF THE GULF OF MEXICO 16
B3o Meilspigy, Mane Conch 1 Ser, 154 pe 156) ply to, 1! 4-6,’ 93.
Punic a cescuipra | Verrmill:| Proc.) W..5) N, M5 32," p. 381,
1880.
wavdus Nernst hr. Connl Acad), 5, p..540) pl. 58) f..21, 1882.
Distribution.— West Atlantic, Martha’s Vineyard, Mass. to
Florida. Also Mediterranean and Bay of Biscay. 92-1465
fms!) (Gulf of Mexico; -Campeche Bank, at ' 200° fms.
Pliocene to Recent.
pusillus Forbes, Rept. Brit. Assoc. Adv. Sci., p. 191, 1843, (as
Tornatella); Jeffreys, Ann. & Mag. N.H.,4th ser., 6, p. 84,
5th ser. 10, p, 34; Watson, Chall. Rept. Gastr. p. 627; Dall,
Bulls a7. Ws S.No M., p. 84, 1889: Pilsbry, Man? Conch: ;
TSC NSHP 5ONI OOS:
An unfigured species.
Distribution.— Mediterranean, Madeira; West Indies. 20-
450 fms. Gulf of Mexico, off Sand Key, West Florida, at
eT istaNS
perforatus Dall, Bull. M. C. Z., 9, p. 96, 1881; 18, p. 42, pl. 18,
Peso bull 27 Ua oo) NaN DN nS 4a ula TSM iit 35.089)
Bilobiyw Man Conch. 1 Ser. 5) pel 5Om ple 204 ta 2On ESO"
Distribution.—Gulf of Mexico, 805 fms.
McIsSUS Dall NEC 7G .i Ds 05.) SSI kon puraze lei toute TO:
POOP BUN a TA UR e SHON). Men sp. San Olen 7a fil TOs 218.0):
il Sbinyag Mani Conehis 1 Sets) 15. pa Loluinog”
Distribution.—Yucatan Strait, off Cape San Antonio, 640
fms. Also off Fernandina, N. EF. Fla., 294 fms.
delicatus Dall. Bull. M. C. Z., 18, p. 41, pl. 17, f. 5, 1889: Proc.
We Se Ne METZ spy 296, /8o 7. Bullet anne SUNG Ma hoc 7Sae
plate peins 62S.) balsbtaya Mans Conch aisers 1.154) p41 102.-pl:
DO, ty Bin, OB
Distribution.—Gulf of Mexico, West Florida, region. Also
Barbabos 73-400 fms.
17 BULLETIN 35 47
galvestonensis Harris, Bull. Amer. Pal., vol. 1, p. 96, pl. 3, f. 1,
1895.
Distribution—Upper Miocene, Galveston well at 2552-287:
feet.
Genus RNIGICULA Deshayes
RitidaVeralh, Am, our Seu auisen i) 5 pu Lows 730 br. Cons
INCAd Ms DAS wlan Tuan tact Dalla olleeNten Cp Zi nS aioe
AB WSO Bite aan Da Ott MDl Marte norns @ qi Ens otivgy (UMM taltay
Conchiy ni sen sy is5..p- 290.) ple downs 36 1O3 +) OLSON. Oces
Pay eBOSt SOC INIa UL a0) Tio) i sliy | OMe
leplockevla Brugnone,) Misc! Male ipuiriiuipls)) Py it. 073) L87se
Morelet wound de (Conch) \ppuimerm28 5. pla ssa tem7 aor
Agassiz, Three Cruises of the Blake, 2, p. 70, f. 291, 1888.
Distribution.—Massachussetts to Pernanibuco, Brazil. Also
Pliocene of Italy (Brugnone) and recent in the Mediter-
ranean. 19-1075 fms. Gulf of Mexico, Yucatan Strait,
640 fms.
Genus ACTEOCINA Gray
bullata Kiener, Sp. et Icon. Cog. Viv., p. 5, pl. 1, f. 4, (as
Toratela) Wall Bulli 7 Onis Ne WME SA o oor
Palsbirya) Mans (Concha Ser nuis 4) pus 30) ple 50, bt.) 301 pluj22.
fa i7- tol (Os) iblarnis Bull yAcne mals ol it; ps OOumOsE
Dall sr Simpsoni Bully Sakisky (Com) 1. p. 362, plisyn
LOM uOO!
Bulla canaliculata d Orbigny, Hist. Pol. y. Nat. Isla de Cuba,
HpsLsee pla) Diswei2ic24u Noto Say.
Tornatina olivula A. Adams, Thes. Conch., 2, p. 569, pl. 121,
an gat
Distribution.—Florida Strait to Trinidad Island. Upper
Miocene, Galveston well at 2410-2425 ft. (Harris).
recta d’Orbigny, In Sagra’s Hist. Pol. y. Nat. Isla de Cuba, 1,
DIU tpl 4) bis) t4 Mazo m(aswauiia) WlOall yy Ball a7) UNS.
48
MoLLUSCA OF THE GULF OF MEXICO 13
NGNME p84) 18897 (Pilsbry, Man) Conchvtiser,15,/p. 184,
pl. 22, f. 13-15, ’93, (as Zornatina); Maury, Bull. Amer.
PANO 20) pital plalai ty 2hTol7:
Distribution.— Florida Keys, Antilles and St. Helena. Gulf
coast, Tampa. Also Lower Miocene Santo Domingo and
Jamaica. A closely related, Older Pliocene, species is
Acteocina wetherilli Lea, from Deal, N. J. and the Orbitolite
bed, Tampa, Fla.
canaliculata Say. Jour.. Acad. Nat. Sci. Phila., 5, p. 211, 1822,
(as Volvaria); Holmes, Post. Pl. Foss. S. Car., p. 78, pl. 12,
Mme iian Gouldw Binney) vedu nips 259 401115 kw S704 CaS
Werculus) + Wally Proc: Us SiiiNaiMe) Os p. 324.) S35 Bulle 37,
WM Sa Ni Ives woS4) pli SohteM 27 Sonia (asl onmarz7a)):
Singley, 4th Ann. Rept. Geol. Tex., p. 332, 92; Pilsbry,
Mane Conch iiisene 15) ipl o4epI M22 ib 2 2M) ple 5 Ousn2'5,
Pow og.) Johnson, Occ.) Pap: Bost oo0ci) Ni En (70. 247,
Honorary Vall) Am, PaliineNow 2oqn pila.) Plush.) 2),
IQI7.
obstricta Gould, Silliman’s Jour. Sci. 38, p. 196, 1840, (as
Bulla wAdams.))\ Hes), 2.) pae15OOut 2h.) 200), (as
Tornatina).
Distribution.—Portland, Maine to Haiti, 0-63 fms. Lower
Miocene (Santo Domingo) to Recent. Gulf coast.—Recent:
Cameron and Point au Fer, La., (very abundant); Galves-
ton and Corpus Christi, Tex. Pleistocene: North Creek,
Manatee and Labelle, Fla.; Grand Chenier, La , Knapp’s
wells, Terrebonne Parish, No. 2 at 1050-1790 ft., No. 3 at
258-1525 ft. Pliocene; Caloosahatchie River, Fla. Miocene;
Gilbert well No. 10, Bateson, Tex., 323 ft.
candei d’Orbigny, Moll. Cuba, 1, p. 128, pl. 4, f. 1-4, (as Bulla);
Daal a 700) SING Me pa Saem Olas fs (13.339),
Vanatta, Pr. A. N.S. Phila., 55, p- 757, 1903.
Distribution. Hatteras to Martinique, o-48 fms. Gulf
1g BULLETIN 38 49
coast, St. Joseph’s Bay and Crooked Isl., West Fla.
Genus RETUSA Brown
frielei Dall; Bull. M. C. Z., 9, p. tor, 188%, (as Uirzculus): 18,
Diaz pl 175) te 4s (SOs MOM STA MOE SisilNis av leis Omastoleunlicge
tia.) 89°) Pilsbry., Man. Conchi (a isera Wins saps 2uO nla inte
8, 1893.
Note.—I follow Cossmann in placing Ufricudus Brown,
1827 (not of Schumacher, 1817) in the synonymy of
Retusa.
Distribution.—Yucatan Strait, 640 fms.
Genus SCAPHANDER Montfort
punctostriatus Mighels, Pr. Bost. Soc. N. H., 1, p. 49, 1841, (as
Bulla. Wall, Ball 127.) Ula S Nee Mees Op ba 2 sete Aan Ors
Pilsbry Man: Conch.) 1isereng ep 246 (aol hah ommOes
Johnson, Occ! Pa. Bost: S: Nee aie pai4on Doms.
librarius Love’n, Index Moll. Scand., p. 10, 1846.
Distribution.—Iceland & Norway to Bay of Biscay. Maine
to Barbados, 46-1467 fms. Gulf of Mexico, Lat. 28° N.,
Lon. 88° W., 533 fms. Inhabits shallow water in its north-
ern habitats.
watsoni Dall, Bull) MC. Zo); pgonimSSir aime p52 ,iyole elie
for 78o.0 Bull 271) Wi. Su Nee Mes py SONI aie mons,
Distribution.—Hatteras to Barbados, 54-324 fms. Gulf of
Mexico, West Fla. region, dredged U.S. Fish Com., bot-
tom temperature 46° F.
nobilis Merrill, Tr. Conn.) Acad: /Seni6, sp izogy plazas:
Dall Ball. Moi Z.. 718, p53) 1869s leilsbmy.wNtaney © onelny
TS CIAM EllS NDA 24 OmOoe
Distribution.—Martha’s Vineyard 906-1309 fms., Delaware
Bay log1-1209 fms., Tobago 880 fms., Gulf of Mexico, U.
S. Fish Com. Station 2127, at 1639 fms.
50 MOLLUSCA OF THE GULF OF MEXICO 20
Genus SABATIA Bellardi
bathymophila Dall, Bull. M. C. Z., 9, p. 98, 1881, (as Azys?);18,
PSs to NOD. Son Bulls) 27. Wel SiNoy Meni ps 36, pl.
Mt Os OD. SOx
Distribution.—Fernandina to Guadeloupe, 294-1568 fms.
Gulf of Mexico, Yucatan Strait, 740 fms.
Genus CYLICHNELLA Gabb
bidentata d’Orbigny, Moll. Cuba 1. p. 125, pl. 4, f. 13-16, 1841,
Gsve7da) Gabb.) (Pr. AYN Se Phila 240 ou 27h) pla iO i
ae alle (Bulle 37. Ul Se Neri py. 86) plu anaite) wa 780);
Wenattay Pr AU NS. Phila. 5 5.yp9075 75) L903:
Bulla biplicata Lea. Pr. Bost, Soc. N. H., 1, p. 204, 1844.
Distribution.—Hatteras to Barbados, 7-168 fms. Pliocene
to Recent. Gulf coast, Recent: West Florida and Texas.
Pleistocene, Louisiana, Knapp’s No. 2 "well, Terrebonne
Parish, 1519-1542 ft., Crowley No. 4 well, Jennings, 1663-
1670 feet.
bidentata var. gaivestonensis Harris, Bull. Am. Pal., vol. 1, p.
GoOmpla it: 2) 1895;
Distribution.—Upper Miocene, Galveston well at 2600-2733
HECE:
Genus BULLARIA Rafinesque
striata Bruguiere, Ency. Meth., 1, p. 572, (as Bulla); Dall,
Bullen 37, Wl SNe Me) 1p. 88,) 1889; Pilsbry,)) Man. Conch.) 1;
Sell SD See i Dle a7 uty 42-40... Oze
Distribution.—Recent, Mediterranean ahd Atlantic coasts of
Portugal and Morocco. Also living at Clearwater Harbor,
West Florida; New Harbor Isls. and Chandeleurs, Louisi-
ana; Texas. Pliocene, Caloosahatchie marls, Fla.
occidentalis A. Adams, In Sowerby’s Thes., 2, p. 577, pl. 123, f.
VERO S On (as bulla) Wall ei Ui os Ni MEMO np 1324)
Se uRully a7 UES) Ne Mi ip.) 88.0780; )oimeley; 4th Amn:
21 BULLETIN 38 st
Rept (Geol Surv We pag320 02 Naiatca, wet weno
Phila 5 5. pai s7 se koos
Distribution.—Florida to St. Vincent, W. I. Gulf coast:
Cedar Keys, St. Marks, Tampa, Crooked Isl (Calhoun Co.),
Fla.; Corpus Christi and Matagorda Bays, Texas.
solida Gmelin, Syst. Nat., 13, p. 3434, (as Bulla); Dall, Bull.
Bh nO SNe MESS py) SSeS ocubilsbiey ny Miany Conch imisets
U4: Oy NS Olly AUG Teh Ia, VECO).
Distribution.—West Indies, Vera Cruz and Maracaibo.
Cited from Texas region, Gulf of Mexico (Dall, ’8q).
abyssicola Dall, Bull. M. C. Z., 9, p. 97, 1881, (as Bulla); 18,
SO) pla i7e tai So; Bulle es 7 aaOr oN in DSS) sp ka aly
Tl SOselisbry, Man. (Comchessiasemen 5.) 15 23o. plan sor
21, 2S93"
pinguiscula Jeffreys, Ann. & Mag. N.H., p. 318, 1880, (women
nudum); Watson, Chall. Exp. Gastr., p. 638, 1886.
Distribution.—Bay of Biscay, Azores, Antilles and Yucatan
Strait, 450-640 fms.
Genus HAMINGEA Turton
elegans Gray. Ann. of Philos. new ser. 9, p. 408; 1825; Index
estac.) ‘Suppl. Apliii2) wigienuas) Pella) Pilsbry, Vian
Conchre ar iseraerss ps5 5 ale wate 37-30), Pl 4ONmer Sos
1893; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 364,
IQOI.
guilding?t Swainson, Malacol, pp. 251, 360, f. 46, 1840; Adams,
hes!) P4580.) pl m2aaikas 7 Somalia ull 277, Unis siN eaves
p. 88, 1889.
Note.—/ide Pilsbry, the true elegans of Gray is this species
and not a European shell.
Distribution.—Antilles to Rio de Janeiro. Gulf coast, West
Florida and Texas.
52 MOoLLUSCA OF THE GULF OF MEXICO 22
Succinea Conrad, Pr. A. N. S. Phila., 3, p. 26, pl. 1, f. 5, 1846,
Geyaiaye Dally eri ioaM Ney MeO ps 6324,1 08837 Bull,
PUR Oe Nig Me pa 88, 1803) Mitchellt Mist) Mex) Sh;
ilcbiye Many | Conch, 1Ser) 15.) py 35 jy Dla 484i 119,393;
Dall and Simpson, Bull: U. S. Fish Com., 1, p. 364, Igot.
Distribution.—Indian River, E. Fla., to Texas. West
Florida at Tampa (Conrad’s type locality), and Cedar Keys;
Chandeleurs, La.
aniillarum d’Orbigny, Moll. Cuba 1, p. 124, pl. 4, f. 9-12;
Mosrehy Mal Bl. 22) p. 1755) Walle, Bulls (37. U2 5. No M.;
MSs Tsso-) Pilsbry, Mant iConchy. 1Sen) 15. ps1358,) ple 41,
f. 35, 36, 1893; Mitchell, List Texas Shells.
cerina Menke, Zeitschr. fur Mal., p. 142, 1853.
Distribution.—W. Florida to Porto Rico and St. Thomas.
Gulf coast at Tampa, Fla., and Espiritu Santo Bay, Texas.
antillarum var. guadalupensis Sowerby, Conch. Icon., f. 14, 1868;
eilsbraya Mian (Conch, 1 (ser. 5.) pa a5on) Dis) 405/11 30-34,,
1893.
Distribution.—Guadeloupe, Cuba &c., Gulf coast at White
Water Bay, West Florida.
petiti d’Orbigny, Moll. Cuba 1, p. 130, pl. 4, bis. f. 13-16, (as
Bava Dally Bulli i272 Ui. SusNe a Mops Soup TSso a bllsbiny,
Mane Conch... 1) Ser, 15, p. 359, plied) 234) 245, 1893.
Distribution.—St. Thomas, W. I. and Tampa, West Fla.
virescens Sowerby, Genera, No. 39, f. 2; Dall, Tr. W.1.5., 3,
Pas 1890, kilsbry, Man. Conch:))1 ser, 15" p: 360, pl. 4o,
temo Pl 43, 1-119; 1893.
Distribution.—Recent on the Pacific coast, California to
Mexico. Pliocene, Caloosahatchie beds, Fla. Not in the
recent Atlantic fauna.
Genus CYLINDROBULLA Fischer
beaui Fischer, Jour. de Conch., p. 275, pl. 8, f. 8, 9, 1856; Dall,
23 BULLETIN 38 ' 53
Bull. 37, U. S. N M., p. 88, 1889; Pilsbry, Man. Conch., 1
SehS ps 280 pla A 2uit IO 2 OMS aOeE
Distribution.—Guadeloupe (type locality), St. Thomas, W.
I.; Gulf coast at Cedar Keys, 2-95 fms. ‘This species is the
genotype.
Genus HYDATINA Schumacher
physis Linnzeus, Syst. Nat., ed. x., p. 727, 1758, (as Bulla);
Dall Bulli ie7 00s 1S INS Ores ais oOr ealsbinyan Wate
Conch, 'ser1015, 9. 387.) plague 8037) Dallvand
Simpson) /Bull7U.S. Fish Comayarip. 364) noon.
Distribution.—Atlantic, Pacific and Indian Oceans. Gulf
coast: Sarasota, West. Fla.
Genus PHILINE Ascanius
flexuosa M. Sars, Nyt. Mag. f. naturvidens., 17, p. 181, pl. 11, f.
23-26; Christianiafjordens Fauna, pp. 69, 70, pl. 11, f. 23-
26-4 Wall Bull Me iC Zins Som sou reso bull 27. Un Senin
Mi p88, 1889; Pilsbry,)ManynConch!) im ser.) 16) ;pyi214iple
4, f, 86-89, 1896.
Distribution.—Christiania, Norway; Yucatan Strait, 640 fms.
Genus TETHYS Linnzeus
protea Rang, Hist. Nat. des Aplysiens, p. 56, pl. 10, f. 1-3,1828,
(as Aplysia): d’ Orbigny, Moll. Cuba, 1, p. 117; Beau, Cat.
Coq. Guadeloupe, p. 20; Dall, Pr. U.S. N. M., p. 324,
193; Pilsbry,. Man) iConcheiiraseruno.np. 78. pl. 3 7an2o-22.
1895; Dall & Simpson, Bull. U. Shi Fish Com: 1) pies:
TQOL.
Distribution.—Florida to Carthagena, Columbia. Very
abundant in the Antilles. Gulf of Mexico, West Florida
(Dall).
willcoxi Heilprin, Pr. A. N. S. Phila., p. 364, 1886, (as Aplysza) ;
Da Bialls337, Wi SaNeeNieMpa GO SO) loll SbivaiMiane
54 MOLLUSCA OF THE GULF OF MEXICO 24
Conch., I ser. 16, p. 80, pl. 35, f. 30-32, 1895; Johnson, Occ.
aoe BOSte SOC. ING Tine 5 An) ROWS.
Distribution.—Little Gasparilla Bay and Marco West
Florida. Also Massachussetts coast.
ORDER PULMONATA
Genus AURICULA Lamarck
(Auriculastrum) pellucens, Menke, Syn. Meth. Moll., p. 131, 1830;
Daler WSN): MESA pha Tsui WTS. fay Sos (ioctl le
BOS) Ne) Menip! oo. plac sete SenlS so:
Distribution.—Cedar Keys, Florida, to Demerara, British
Guiana.
Genus PEDIPES Scopoli
mirabilis Muhlfeld, Mag. Ges. Nat. Freude, Berlin, 8, p. 8, pl.
MPN nae ay TSS 4) Dall Bullies SN Given pa O2)\ipll
47, f. 17, 1889: Dall & Simpson, Bull. U. S. Fish Com., I,
DP BOOW ples 53,11.3;) LOOl.
guadridens Pfeiffer, 1839: ovalis Adams, 1849, ¢ridens Pfeiffer,
1854.
naticoides Stearns, Pr. Bost. S. N. H. 13, p. 108, 1869; Dall,
Pr W)o2 INS )MiA8) 5.270) Pi=Ee me a7 Lo O5)
Distribution.—Tampa, Florida,to Guadeloupe Island.
elongatus Dall, Pr. U. S. N. M.,8, p. 279, pl. 18, f. 4, 1885; Bull.
BUL SOHING ME o2s) pl. 47, ita ATs so.
Distribution.—Marco, Southwest Florida.
Genus MELAMPUS Montfort
coffeus Linnzeus,Syst. Nat. ed. x, p. 729, 1758,(as Bulla); Dall,
mB alle s7a Wy. 5. UNG Me ps 92, 9pl4azaih.1/35 R889, Dall &
Simpson, Bull. U. S Fish Com. 1, p. 368, pl. 53, f. 13, Ig0T.
coniformis Bruguiere, Ency. Me’th. 1, p. 339, (as Bulimus).
25 BULLETIN 38 55
Auricula ovula d’Orbigny, Moll. Cuba 1, p. 187, Atlas, pl. 13,
Lea TS 534
Distribution.—Florida to French Guiana. Pleistocene to
Recent. Gulf coast, Cedar Keys, Fla. and Texas.
coffeus var. gundlachi Pfeiffer, Vanatta, Pr. A. N.S. Phila., 55,
P- 757, 1903.
Distribution.—St. Andrews Bay, Washington Co., Fla.
floridanus Shuttleworth, MSS., Pfeiffer, Mon. Aur. Viv., p. 36,
7O56: (Mall, Bulle s70Ue Sine Mies ozanmln anit 2) 153.0.
Distribution.—Tampa to Florida Keys.
flavus Gmelin, Syst. Nat., p. 3436, 1792, (as Voluta); Binney,
Jour.Bost.Soc.) Nat) Hist eipen 60. 1d03--land Galkin
Water Shi-pt..2. p. 12,°65; 2DallkperanWens, IN: Mis) 8) ip:zom.
plMmsit. 2) 85 ullie 27 OM Same enn p 4 O2n mol. Ar aleeoTee
°89; Dall & Simpson, Bull. U.S. Fish Com., 1, p. 368, pl.
54, f. 9, 1901; Mitchell, List Texas Shells.
Voluta flava Gmelin, Syst. Nat. p. 3436, 1792.
Bulimus monile Bruguiere, Ency. Me’th., 1, p. 338, 1780.
torosa Moerch, Cat. Yoldi, 38, 1852.
coronatus Adams, 1849; coronulus Adams, 1854.
Distribution.—Florida to Guadelonpe. Gulf coast: Tampa,
Cedar Keys, Fla.; Point at Fer, Wa.; Port Lavaca and
Espiritu Santo, Tex.
Note.—According to Mr. Mitchell, this species lives near
shore, hiding in the day time under drift wood and sea
weed and goes in search of food nocturnally.
lineatus Say. Jour. A. N.S. Phila., 1 ser. 2, p. 246, 1822; Dall,
Bullve7, Uns. NR IMe poe tnp lag pet) (L240 SQ~.Simedleye
4th Ann. Geol. Rept. Texas, p. 352, ’92: Johnson, Occ. Pa.
BOSE SOC. ENG Hal 74 (pn aiSeomse
Wiaeniatus Says yout.) A. INGi Sikandar Sele 2) up) 2415 O22
Dekay. INV.) Moll pansye Dias ateO2.) mln Zar cau enwar
56 MoLLUSCA OF THE GULF OF MEXICO 26
Binney, Land & Fr. W. Sh., 2, p. 10, ’65; Not dzdentatus
Montagu, 1803, which is European, but introduced on New
England coast.
biplicatus Pfeiffer, Mon. Auric. Viv., p. 21.
Auricula cornea Weshayes, Ency. Me’th., 2, p. 90, 1830;
jaumet Mittre’, 1841.
Distribution.—Maine to Tortola Isl., Gulf coast: Tampa,
Cedar Keys, St. Mark’s, Fla.; Point au Fer. Chaudeleurs,
Belle Isle, La ; Galveston, Tex.
Note.—Northern specimens very distinct; southern grade
towards flavus with which they may hybridize. Inhabits
salt marshes.
(Detracia) bulloides Montagu, Testacea Brit., p. 339, pl. 30, f.
WISes. (as. Volutia). Dall) Pra) Ue S. Na) ME) 8) p.)285, pl
Ieee SSSe) Bull..37), UW. Sa Nee so g2, plia7, f7 7, 89.
Distribution—Cedar Keys, Fla. and the Antilles. Type of
Gray’s subgenus Detracia.
(Sayella) hemphilli Dall, Pr. U.S. N. M., 6, p. 323, pl. 10, f. 6,
1883, (as Leuconza); Pr. U. S. N. M., 8, p. 286, pl. 18, f.
Tso Bulle WS.) ING Mei 7) paoou pls Az rey ith 8O);
Singley, 4th Ann. Rept. Tex. Surv. p. 343, ’92.
Distribution.—Cedar Keys, Florida, and Corpus Christi,
Texas. ‘Type of section Sayella Dall which is intermediate
between Blauneria and Detracia.
(Sayella) crosseana Dall, Pr. U. S.N. M., 8, p.,286, pl. 18, f. 10,
sey Bulllis7. U.S. Na M., p. 92) pliy47,) {2 10,789") oingley,
AthieAnn Rept. Lex. Surv., p-) 343,92).
Distribution.—West Florida at Egmont Key (Tampa Bay);
Corpus Christi, Texas and the Bahamas.
Genus PLANORB1IS Guettard
Ophis Dall Proc. U.S: N. M:, 46, p. 236;;pls 21, f:/3,'\45 1914.
27 BULLETIN 38 57
Distribution.—Upper Miocene or Pliocene. Well near Al-
exandria, La., at 49 feet. Brackish water formation.
Note.—The living species of this fresh water genus are not
included in this Synopsis.
Genus TRALIA Gray
pusilla Gmelin, Syst. Nat. p. 3436, 1792, (as Voluta); Dall, Pr.
Wa Sn NGM Secon a7 6a Wins limite sain eis) ECL a7 Oise Ne
NE o2), (pl Ag in. sii Son mall Sai Simpson allen mss
Fish) Com.) ps) 260; plisaiite 13hinoom.
triplicata Donovan, 1802; ovula Bruguiere, 1789; z¢tens
Wamarck ns22%
Distribution.—Cedar Keys to Guadeloupe.
minuscula Dall, Bull. 37, U.S N. M., p. 92, 1889.
Distribution.—Tampa to Bahamas.
(Alexia) myosotis Draparnaud, Tabl., p. 53, ’o1, (as Auvzcula);
Dall PrUS MN OMS i Dwi 7 Ame se motile 2 Zep Ss) N tem ules
DHOZ Mpls 25 fo 8o!
denticulata Montagu, Test. Brit., p. 234, pl. 20, f. 5, ’02, (as
Voluta)
Distribution.—Europe; Jamaica (introduced); West Florida,
San Francisco (introduced).
Genus BLAUNERIA Shuttleworth
heteroclita Montagu, Test. Brit., Suppl., p. 469, 08, (as Voluta);
Arango) Moll; Cubanayp icoma7s Wall Pro) OL) So uN vies
Bp. 287) pl. 17. fh GN SeiBuliene Wes.) NGM), ip-kozmiple
Az, t.)t4, 80, Dall &iSimpsoum bully) S.)Rish Con: in woe
369, I9O01.
pellucida Pfeiffer, ’40, cubenszs Pir.
Distribution.—Tampa and Marco, West Florida. Also
Antillean.
58 MOoLLUSCA OF THE GULF OF MEXICO 28
Genus SIPHONARIA Sowerby
alternata Say, Jour. A. N, S. Phila., 5, p. 215, ’26, (as Patella);
Bimmer Nand Ge Fol Sis) 25) p. 0535) to. 2540005; Dall Px.
eon NAN Ss pn 28797 S57 Bull i270 WS. Menai pe 192)
1889.
Distribution.—The variety brunnea Hanley (Pr. Z. S., pp.
21,151, 58)is found at Sarasota Bay. West Fla.,and Bermuda.
lineolata d’Orbigny, Moll. Cuba, 1. pl. 17, f. 13-15; Dall, Bull.
BAUS SUN. IVD Sp o2) 80:
Distribution.—Fernandina, Fla., to Braziland Texas region
of the Gulf of Mexico.
Genns WILLIAMIA Monterosato
krebsi Moerch, Dall, Bull. 37, U.S. N. M., p. 92. ’89.
Distribution.—West Florida to Barbados.
Fenus YVERONICELLA Biainville
floridana Binney, Terr. Moll. U.S., 2, pl. 67, f. 17, °51; Dall,
Bulla 7. U5) N. M.; p- 90,1889.
Distribution.—Charlotte Harbor, West Florida to Florida
Keys.
Note.—Since Blainville erroneously attributed a shell to this
slug-like genus, Ferussac’s later name Vagznula is often
used; but Vevonicella has priority.
ORDER CTENOBRANCHIATA
Genus TEREBRA Bruguiere
cinerea Born, Test. Mus. Vind., p. 267, pl. 10, f. 11, 12, 1780;
avon Manki 7p. 21 pl. Oo) tv Ova Toss Dalle spill vie C:
ZL e POs so. Sineley,, 4th) Annee Rept) Dex. \py 332)
qo2; Dall) & Simpson, Bulls) Us Ss ikishy Com: 1 ip. 382)
IQOT.
salleana Deshayes, jamatcensis C. B. Adams.
29 BULLETIN 38 ; 59
Distribution.— West Indies to Colombia. Gulf coast:
Chandeleurs, La., Corpus Christi, Tex. Pleistocene, New
Orleans Pumping Station, No. 7.
dislocata Say. Jour. A. N. S. Phila., 2, p. 236, 1825, (as
Cerithtum); Hilgard, House of Rep., Ex. Doc. 1, pt. 2, p.
S87 785) bryou,) Manwi7, joy TSy (plein Og tino sm steledleiye
4th Amn. Rept), Mexia.) a2. OZ Vanattann ai ya Neno:
Philay (550 P7575, 2903.
rudis Gray, petttiz Kiener. Not petit? Maury, 1917.
Distribution.— Maryland to West Indies and Venezuela.
Miocene to Recent. Gulf coast Recent; Cedar Keys, Fla.;
Chandeleurs, La.; Galveston & Corpus Christi, Texas.
Pleistocene of New Orleans pumping station No. 7; New
Orleans artesian well of 1856; Lake Borgne borings; Knapp’s
wells, Terrebonne Parish, No. I at 1600-1700, 2000-2150 ft.,
No. 2 at 1150-1290, 1731-1739, No. 3 at 1150- 1525 feet.
Also of North Creek, Manatee and Labelle, West Fla. Up-
per Miocene, Galveston well, 2158-2920 ft.
concava Say. Jour. A. N.S. Phila., 5, p. 207, 1827; Dall, Tr. W.
Inst. Sci., 3, p, 24, 1890; Singley, 4th Ann. Rept. Tex.; p.
2325.92 Haris, Bulk vA Mralee wv oleer. ps O71 Os.
Distribution.—N. Carolina, Georgia and Texas, at Galves-
ton . Pleistocene of North Creek, Fla. Pliocene, Caloosahat-
chie, Fla. Upper Miocene, Galveston well at 2552-2920 ft.
Note.—The variety vinosa Dall (Bull. M. C. Z., £8, p. 64,
*89) is found on the Gulf coast at Sarasota Bay and Char-
lotte Harbor, West Fla.
protexta) Conrad, Pr, A.) Nis) Phila’, 2), ps) 26) )184ai (as
Cerithium): Cryon) Mantz 25)pl. (6, 1.168, 11805 salle
Ball37) UliS: NoMa Vpsio4aNe sox air. Wagner lmst.eiah ps
25, 90; Dall & Simpson, Bull, Fish Com. I, p. 382, 1901;
Vaughan, Publ. No. 133, Carn. Iust., p. 171, 1910; Maury,
Boll Am) Pall 20) ps2 aplaquvt tole
60 MOLLUSCA OF THE GULF OF MEXICO 30
Distribution.—Hatteras to Texas 2-50 fms. Lower Miocene
to Recent. Gulf coast: Cedar Keys & St. Marks, Fla.;
Cameron, La.; Galveston, Tex. Pleistocene, North Creek,
Manatee and Labelle, Fla.. The variety J/utescens E. A.
Smith, ranges from N. Carolinato West Florida. It is
more distinctly reticulated than the typical form.
nassula Dall, Bull. M. C. Z., 18, p. 66, pl. 36, f. 8, 1889; Dall
& Simpson, Bull. U. S. Fish Com., 1, p. 383, 1901.
Distribution.— Martinique. Also Gulf of Mexico, Lat. 23°
N., Lon. 88° W., at 95 fms.; Yucatan Strait, 640 fms.
limatula Dall, Bnll. M. C. Z.,18, p. 66, 1889; Bull. 37, U.S. N.
M., p. 94, ’89.
Distribution.—N. Carolina to Barbados. Also Gulf of Mex-
ico between Mississippi delta and Cedar Keys at 111 fms.
texana Dall, Nautilus, 12, p. 45, Aug., 1898; Pr. U. S. N. M.,
PMID aSO2 ply 2On te (Oo) 1902).
Distribution.—Matagorda Island, Texas. Collected by Hon.
J. D. Mitchell. Very rare.
galvestonensis Harris, Bull. Am. Pal., vol. 1, p. 98, pl. 3, f. 5,
1895.
Distribution.—Upper Miocene, Galveston well at 2387-2871
IQIO.
calhounensis Maury, Bull. Am. Pal., No. 21, p. 4, pl. 1, f. 3,
19IO.
Distribution.—Lower Miocene, Chipola marl, Bailey’s
Ferry, Fla., and Bascom No. 2 well Mobile, Alabama, at a
depth of 1241 feet, Chipola horizon.
indenta Conrad, Cited by Mr. Aldrich from the Bascom No. 1
well, Mobile, Ala. at 1500-1556 feet.
langdoni Dall, Pr. U. S. N. M., 18, p. 39, 1895; Tr. W. Inst.,
3, pl. 59, f. 27, 1903.
BULLETIN 38 — 61
Distribution.—Lower Miocene, Chipola marl, Fla.; Upper
Miocene Galveston well at 2158-2920 feet (Harris).
Genus CONUS Linnzus
proteus Hwass, Enc. Meth. vers, 1, pt. 2, p. 682, 1789; Reeve,
Conch Iicontit 210/00 Dalla Bull 27a UE SweN Mes omane
1889 Vaughan Publ: Now 122 Carn. Imstcips 7p volo:
Many.) Bulli) Am Pal No Zov meat pl Gn hanno
leoninus Hwass, 1791, spurius auctores, as of Gmelin.
Distribution.—West coast Florida, 19 fms.
zuela. Pliocene, Caloosahatchie River,
Labelle, Fla.
South to Vene-
and Pleistocene,
centurio Born, Mus. Vind., pl. 7, f. Io, 1780; Tryon, Man., 6, p.
33, f. 9, f£. 68,°1884: Dall, Bull. M. C. Z., 18, p. 69, °89.
Distribution.—Antilles. Also Gulf of Mexico between
Mississippi delta and Cedar Keys at 25 fms.
delessertianus Recluz, Mag. de Zool., pl. 72, 1843; Tryon, Man.,
6))p!133.)) pl. Oo) fy 677) Sam Dall (asi delessextze i iaalleviere
We Sh NEEM Di OAe SOR
Distribution.—Hatteras to West Florida, 22-63 fms. Doubt-
ful specimen, Porto Rico.
flavescens Gray, Sowerby, Conch. IIl., f. 68; Tryon, Man., 6, p.
86, pliimo) he 84,,rssacDallmB allies NOLS. Nien aa ae
1889.
Distribution.— Hatteras to Barbados, and West Florida, 15-
170 fms. Pliocene to Recent.
floridanus Gabb, Am. Jour. Conch., 4, p. 195, pl. 15, f. 4, 1868;
Aryon,)Man:.6,) p)i28splimmeutaAnigh arsed Dalle vp imilia zai
94, 1889; Ir. W. Inst. Sci., 3, p, 27, 1890.
Distribution.—Hatteras to Florida Keys and West Fla.
Pliocene of the Caloosahatchie beds.
64 MOLLUSCA OF THE GULF OF MEXICO 32
peali Green, Tr. Albany Inst.,1, p. 123,pl. 3, f. 3, 1830; Tryon,
Mean 6. ps) Z0.)plai 10. )f2/89-92))) 1884; (Dall) Bulls MM: C. Z.,
18, p. 68,789.
seannseconntad) yAm.4) foun Coneh. is) pi toa pl.) TO} /f.01,
1869.
Distribution.—Hatteras to Isthmus of Panama. Also Cedar
Keys, Florida, o-5 fms. Pliocene, Caloosahatchie beds.
pygmaeus Reeve, Pr. Z. S., p. 179, 1843; Kiener, Coq. Viv.
Conus, p. 174,pl. 111, f. 1 a-b,1848; Dall and Simpson, Bull.
WSS) Bish Com.) 1, p. 383. 190K:
Distribution.—West Florida to TobagoIsl. Pliocene, Ca-
loosahatchie beds. Pleistocene, North Creek, Fla.
amphiurgus Dall, Bull. M, C. Z., 18, p. 70, 1889.
Distribution.—Coast of Yucatan, 27 fms.
chipolanus Dall, Pr. U. S. N.M., 18, p. 42, 1895.
Distribution.—Lower Miocene, Chipola marls, Florida and
of the Bascom No. 2 well, Mobile, Alabama, at 1241 feet.
Genus TURRIS Bolten
albida Perry, Conch. Expl., pl. 32, f. 4, 1811, (as Pleurotoma);
Wall Bull27) We Ss.) IN.) ME p96) 7780; Wall Simpson,
Bulle Wes. Hish! Com). 1) pi 284. 190r- | Maury. (Bulli Am,
Bale INO: )29, (pp. 50; 1917:
Distribution.—Recent, Cedar Keys to Barbados, 26-100 fms.
Oligocene of Yicksburg, Miss., and of Tampa, Fla.; Lower
Miocene, Western Florida and Santo Domingo; Upper Mio-
cene, Galveston well at 2158-2950 feet.
Note.—The variety ¢el/ea Dall (Bull. M. C. Z., 18, p. 72,
1889) was dredged in the Gulf of Mexico between Missis-
sippi delta and Cedar Keys.
Genus LEUCOSYRINX Dall
verrilli Dall, Bull, M. C. Z., 9, p. 57, 1881, (as Pleurotoma); 18,
33 BULLETIN 38 63
PA Siiple LO, 55. 18 So me ulle Brailes pORMensm OO!
Distribution.—N. Carolina to Guadeloupe. Also Gulf of
Mexico, Lat. 25° N., Lon. 84° W., 888 fms., and between
Mississippi delta and Cedar Keys.
sigsbeei Dall, Bull. M. C. Z., 9, p. 57, 1881, (as Pleurotoma) ;
18.270, \pl. li tO. Sos Bulla TOE A SMU Nu NIE Map AOON
Pla Ths OMe So:
Distribution.—Yucatan Strait, 640 fms. Off Bequia, 1591
fms.
tenoceras Dall, Bull. M.C. Z:, 18; p: 76; pli 36, £. 5; 1889;
Bullita7) OS. Nee pi oo" plaizoamanssnioo:
Distribution.—N. Carolina to Guadeloupe. Gulf of Mexico
between Mississippi delta and Cedar Keys at 724 fms.
subgrundifera Dall, In Agassiz’s Three Cruises of the Blake, 2,
p: 66, f: 282, 1388, (as Plezrotoma) Bulle Me Cy Zan sa:
77) Oly Z8e 142) 789. Bills, 27.(U Se eNEy Mes DOO, please
I Oy
Distribution.—Cape Fear, N. C., to St. Kitt’s, W. I. Gulf
of Mexico, Yucatan Strait, 640 fms., and between Missis-
sippi delta and Cedar Keys, 940 fms., U. S. Fish Com. St.
2384.
Genus ANCISTROSYRINX Dall
radiata Dall, Bull. M. C. Z., 18, p. 78, pl. 12, f. 12, 1889; Bull.
87h Uns. Ne Me. p06) plana tegen So. all) Se. Simpsons
Bull U.S. Kish Com, (1, pigsanimoor:
Distribution. —Gulf of Mexico, Yucatan Strait,64o fms. and
between Mississippi delta and Cedar Keys. Also Antillean.
Genus GENOTA Adams
mitrella Dall, Bull. M.C. Z., 9, p. 56, Aug., 1881, (as Pleuro-
LOD) N18 op., 794 Plinnletaisy mySormbtt len 277. ns) SsuN aver
TO) CO) ONY 107) 505 Be 1 Mesoy.
didyma Watson, 1881.
Distribution. —Yucatan Strait, 640 fms. Also off Sombrero.
Weal.
64 MoLLUSCA OF THE GULF OF MEXICO 34
Genus DRILLIA Gray
ostrearum Stearns, Pr. Bost. Soc. N. H., 15, p. 22, 1872; Tryon,
Man Pap mor pla te 7 O a oar Oallem Bullen ise N
M., p. 96, ’89; Dall & Simpson, Bull. U. S. Fish Com., 1,
Pp. 385, 1901.
Distribution.—Hatteras to Cape Catouche, Yucatan and An-
tilles. 15-170 fms. West Florida: Recent, Cedar Keys;
Upper Oligocene, Tampa; Lower Miocene, Chipola beds;
Pliocene, Caloosahatchie beds.
albicoma Dall, Bull. M. C. Z. 18, p. 83, pl. 10, f. 8, ’89; Bull.
We Ss Ne Me; 37,0: 96; pl: 10,/f 8); 1889:
Distribution.—Gulf of Mexico, Lat. 23° N., Lon. 89° W.,
84 fms. Also off Barbados, and St. Thomas.
detecta Dall, Bull. M. C. Z., 9, p. 65, Aug., 1881: 18, p. 84, pl.
Wei Toso.) bull. 37." ps 96) plaio han Sor
circumvoluta Watson, Jour. Linn. Soc., 15, p. 465, Nov. 1881;
ReptyChall, Gastr. p. 357, plai2ik, f.10. 188.5.
Distribution.—Gulf of Mexico, 339 fms. Also Culebra
Sin We
alesideta var. macilenta Dall, Bull. M. C. Z., 18, p. 85, pl. 36, f.
TSS On bulla a7 NU.) S.No VM pooh ply 26nihon 78o)
Distribution.—Cape Fear, N. C. to Barbados. Gulf of
Mexico between Mississippi delta and Cedar Keys, 111 fms.
Typical form of species, off Hatteras.
eucosmia var. canna Dall, Bull, M. C. Z., 18, p, 86, 1889; Bull.
BROS ON Mie. 1: 96; 780.
Distribution.—Cape Lookout, N. C., to Grenada, W. I.
Also Gulf of Mexico, west of Florida, 50 fms.
leucomya Dall, Proc, U.S. N. M., 6, p. 328, pl. 10, f. 8, 1883;
Bel, Vy, Fs COR jello sig sy Sustskeye “Aes WGN DBalsie 7 hy yb eer
1890.
35 BULLETIN 38 65
Distribution.—Shores of Gulf of Mexico from Sarasota,
West Fla., to Yucatan, 3-5 fms. Pliocene, Caloosahatchie
beds, Fla.
albinodata Reeve, Zool. Proc., 6, 1846; Tryon, Man., 6, p. 197,
plyags £5, 1884) DalleBull 27, We SiN MES: oO So:
Distribution.—Charlotte Harbor, West Fla., to Santo
Domingo.
haliostrephis Dall, Bull. M. C. Z., 18, p. 86, pl. 13, f. 3, 1880;
Baller UES. WING MLL pi Oo hp lem TigwEnoNs GO!
Distribution.—Gulf of Mexico, West of Florida, 84 fms.
acestra Dall) 'Bull’'37/°U) S: NVM poor ple ro, tf. 7) So.
Distribution.— West Florida to Grenada 161-400 fms.
pharcida Dall, Bull. M. C. Z., 18, p, 88, pl. 12, f. 2, 1889; Bull.
B27 Mp UOOMple U2 teN2 Os
exasperata Dall. Not of Reeve.
Distribution.—Gulf of Mexico, west of Florida, 229 fms.
Also East Fla. to Barbados 150-1002 fms.
tristicha Dall, Bull. M. C. Z.. 18, p. 88, 1889; Bull. 37, p. 98,
1889.
Distribution.—Three stations Gulf of Mexico between Mis-
sissippi delta and Cedar Keys, 111-210 fms. ‘Temperature
522 - Ore
ebur Reeves, Pr. Zool. Soc., p. 116, 1845, (as Pleurotoma);
Conch!Tcon!) pli 20M ams as dinyony, Man.) 6) pumnser
plas) £56, 84: Dall Bulle Ue Sa uNe EME .\p.O8aisox
Distribution.—Hatteras to Sombrero Isl. Also Gulf of
Mexico, West of Florida, Blake Station 10, at 34 fms.
pagodula Dall, Bull. M. C. Z., 18, p. 90, pl. 13, f. 6, 1889; Bull.
By pO8y ple 138 £6) Soe
Distribution.— Gulf of Mexico, West of Florida, 50fms.
Also Barbados.
66 MOLLUSCA OF THE GULF OF MEXICO 36
ineayDall Pr. U.S. NM.) 6, p. 328) pl) 10) f..5, 2883; Bull, M.
CALS) Oo 7897) Bullvia7. Us Say Nav Muli. O8) pl. i48;
eT SO):
Distribution.— Hatteras to Florida Keys, and West Florida
at Cedar Keys, 3-15 fms.
Note.—The variety carminura Dall (Bull. M. C. Z., 18, p.
gi, 1889) was dredged in Gulf of Mexico, U.S. Fish Com.
Station 2402 at 111 fms.
simpsoni Dall, Pr. Davenport Acad. Sci., 5, p. 54, 1887, (as
ieuvovoma): Bull. M:C.Z., 68) p) or; 1880.
Distribution.— Hatteras to Tampa Bay, West Florida, 15-
18 fms.
lissotropis Dall, Bull. M.C. Z., 9, p. 58, Ang. 1881; 18, p. 91,
VieELINake4 TS SOs Bull a7 MOs,\ Plauen mien sunain SOs
hypsela Watson, Jour. Linn, Soc., 15, p. 433, Oct. 1881; Chall.
Caste s4r ply 21.0. 41885:
Distribution.—Gulf of Mexico, West of Florida at 220 fms.
Also Antillean. Variety ferpolita Dall (Tr. W. Inst., 3, p.
36, ’90), Caloosahatchie Pliocene.
dalli var. cestrota Dall, Bull. M. C. Z., 18, p. 92, 1889; Bull.
375 B- 98, °39.
Distribution.—Between the Mississippi delta and Cedar
Keys, at 196 fms. Typical dallz Verrill, ranges from
Martha’s Vineyard to Delaware Bay.
nucleata Dall, Bull, M.C. Z., 9, p. 62) ’81; 18, p. 92, pl. rr, f.
1 SH ausxexO)p
Distribution.—Gulf of Mexico, 229-339 fms. Also Antillean.
Pleurotoma amblia Watson may be a synonym.
Ven Dalle Balle Me VC.) 59) sp. 68.) 188i TS.) pn Osi ple mony ik.
ZR SO Ulla 2 7Aa Ow OS plese bie tei 2 sO.
Distribution.—Gulf of Mexico, West of Florida, 220-310 fms.
2
~
BULLETIN 38 ; 67
havanensis Dall, Bull. M. C, Z., 9, p. 67, 1881; 18, p. 93, pl. 11,
PASSO!
Distribution.—Yucatan Strait, 640 fms. Also off Havana.
lithocolleta Watson, Jour. Linn. Soc., 15, p. 441, 1881; Chall.
Rept. Gastr., p. 320, pl. 24, f. 6, 1885 (as Pleurotoma);
Dalia whl MC Z ss Vie Os WO. 180: souls yee
OVS Holl aiE aby Kn MAVO\s
a
i)
—
.
4
tI
.
Lanie)
Distribution.—Hatteras to Guadeloupe,W. I. Also Gulf of
Mexico between Mississippi delta and Cedar Keys, at 940
fms.
Genus CYMATOSYRINX Dall
centimata Dall, Bull. M. C. Z., 18, p. 95, pl. 36, f. 9, 1889; Bull.
27 AO Se NL... QS: spl Zowieor oO!
Distribution.—Hatteras and Gulf of Mexico, Lat. 24° N.,
Lon. 84° W. at 1920 fms., and between Mississippi and
Cedar Keys at 1181 fms.
moseni) Wall, Bull. Mi Co Zur 8 hoof plagon 3 TSsor eb ulie
BNW Se IN MES OS) pla gon tna SO!
Distribution.--Cape Hatteras and Sarasota Bay, West Fla.
Genus PLEURQTOMELLA Verrill
packardi Verrill, Am. Jour. Sci., (3) 5, p. 15, 1873; Tr. Conn.,
ENCAG OS D145 25 /PlANAe tite oeiOlemorpmON Shane 2
Distribution.—Gulf of Maineto Rhode Island. ‘The variety
bencdicin Vii &S,,) (Dall Buller Se INGE paozaple
14, f. 4, pl. 60, f. 70 a, ’89) ranges from Gulf of Maine to
Beqttia, 1290-1507 fms.Gulf of Mexico, West Florida region.
leucomata Dall, Bull. M. C Z., 9, p. 63, 1881, (as Drillia); 18,
PMLZoepVictt, £) ne wsoy Bally ia7. ap LO2 ence eNtolnieliaun a.
1889.
68 MOLLUSCA OF THE GULF OF MEXICO 38
Distribution.—Gulf of Mexico between Mississippi delta
and Cedar Keys. Also Florida Strait. 533-940 fms.
agassizi Verrill and Smith, Am. Jour. Sci., 22, p. p. 394, 1880;
pittew Coma cade 54) pl 4 Say pl. 5y70 £308 ay oe
Distribution.—Rhode Island to Cape Fear, N. C. The
Matiety wcxicana Wall (Bull! 37) U) S. NoM., p: 104) pl. 11,
f. 14, 1889) ranges from Western Florida and Yucatan re-
gions of the Gulf to Martinique. 502-640 fms.
filifera Dall, Bull. M. C. Z., 9, p. 56, 1881, (as Bela); 18, p. 123,
MLO Ou Sonn Bill: 27.) pitOsNe Dyn 2 1 tanO toOs
Distribution.—Gulf of Mexico, West of Florida, 331 fms.
HaugaDale Bull) MC. Z.) 18, \p. T25) 18893) Bully) 37,\ pu 1084)
1889.
Distribution.—Cape Fear, N. C., 407 ims. Also Gulf of
Mexico between Mississippi delta and Cedar Keys, at 1181
fms.
extensa Dall Bull: M.C.Z. 9, p. 55, Aug, 1881, (as var. of
icurotoma blakeana), 18, p. 126, pl. 10) t.)2, 789; Bull. 37,
DAMLOA ple TOs) f: 2° So!
FPleurotoma streptophora Watson, Jour. Linn. Soc., 15, p. 464,
Nov. 1881; Chall. Gastr., p. 366, pl. 10, £. 8 a-b) 1885.
Distribution.—Gulf of Mexico, Lat. 23° N., Lon. 88° W.,
804 fms.; Yucatan Strait 640 fms. Also North Atlantic.
blakeana Dall, Bull, M. C. Z., 9, p. 54, 1881; 18, p. 126, pl. 10,
pio Oc ull 27. my TOA... Ply LOm th eaNoGs
breois Neral, br. Conn. Acad!,6, pij4ar7s plyiaa tf: 82785.
Distribution.—Gulf of Maine to Santa Cruz, W. I., 100-1608
fms. Also Yucatan Strait, 640 fms.; Florida Strait, 339
fms.
39 BULLETIN 38 69
Genus GLYPHOSTGMA Gabb
gabbi Dall, Bull Me Co Zui) puto. plan ah et Alas Oat SOOe
B27 SUL SON IMG DA OO. yOl abut HAC ISAM oa oOE
Distribution.—Gulf of Mexico, Blake Station 36, 84 fms.; U.
S.F.C.Sta. 2405, west Florida, 30-50 fms. Also off Barbados.
gratula Dall, Bull. M. C. Z., 9, p. 64, Aug., 1881, (as Pleuro-
LOMAVE TS De LTO pk M2 TOW Soweto.) Os) LOO molars
Le TOMASO!
incilts Watson, Jour. Linn. Soc., 15, p. 425, Oct., 1881; Rept.
ChalliGastrijip. 2o04n) ple 2auhen Sigs
Distribution.—Gulf of Mexico between Mississippi delta and
Cedar Keys, 227 fms. Also Hast Florida and Antilles.
Pliocene, Caloosahatchie beds.
Genus MANGILIA Risso
balteata Reeve, Conch., Icon., 3, pl., 8, f. 57, 1846; Tryon, Man.
Conch 6) pi 2474 plai24 nes D alin stil i277 ies aaING
M., p. 100, ’89; Dall & Simpson, Bull. U.S. Fish Com.,
Dis Zs,) LOOM:
Distribution.—Hatteras to Barbados and West Florida.
Pliocene to Recent.
astricta Reeve, Pr. Z. S. p. 46,'46; Tryon, Man., 6,p.260, pl. 24,
i260 784 Dall Bill a7 Wat SeeN EM VIES ip.) LOO; 30)
Distribution.—West Florida to Florida Keys.
biconica C. B. Adams, Conch. Contr., p. 65; Dall Bull. 37, p.
100, ’89.
Distribution.— West Florida to Jamaica. Unfigured.
plicosa C. B. Adams, Contr. Conch., p. 54, 50, (as Pleurotoma) ;
Dali Buall2 7. Us SNM Me pA MOON DIS @, ft) )l4.. Onsite
ly, 4th Ann: Rept, Tex. Surv.;)p: 333,92; Johnson, Oce:
Palsy Bost soc. IN. URES paulAlsp ymiOiis):
Pleurotoma plicata C. B. Adams, Bost. Jour. N. H., 3, p. 318,
70 MOLLUSCA OF THE GULF OF MEXICO 4O
Pht G, 40%) Not of) Wamarck,
plicatum Kurtz, 1860; drunnea Perkins, ’69.
Clathurvella jewetti Stearns, Pr. A. N.S. Phila., fcr 1873, p.
BAO Dally Pry Uo So NG MEG. pi 220)) T8833"
Distribution. —Massachussetts to Florida Keys and Gulf
coast at Cedar Keys, Fla.; Corpus Christi Bay, Texas, 0-5
fms. Pleistocene, North Creek, Fla.; Pliocene, Caloosa-
hat chie beds, Fla.
rubella Kurtz and Stimpson, Pr. Bost, Soc. N. H., 4, p. 115,751;
Dall Bull 27. USS. NeMi py 100,289: Tr Wagner Inst.
SCZ) Pe 4190.
Distribution.—Cape Lookout, N. C., to Charlotte Harbor,
West Fla. Pliocene, Caloosahatchie beds. Pleistocene,
Knapp’s No. 2 well, Terrebonne Parish, La., at 1800 feet.
stellata Stearns, Pr. Bost. Soc., N. H., 15, p. 22, ’72; Dall, Pr,
eon NE IME, 6.0 3280.87) bnyvont Man 6.) 240) pluiaa.
MOAGon Dally Bal. 2730) pi TOOM SOs Mer aN GON Sin aus
4I, ’90.
Distribution.—Cedar Keys and Tampa to Key West, Fla.,
Pliocene, Caloosahatchie beds.
athoswvlay Dali, Bull!) 37) WU: So Ne Me puino2 plan isa) aay
1889.
Distribution.—Hatteras to Barbados. Also West Florida,
14-333 fms.
l imonitella Dall Pe WS. ING MEO pi B2Om ple TOs) hy TOMS 83 =
PPS oul 7 MP y1O2, Pl. j48, £:) 25°89.
Distribution.—Cedar Keysto Tampa, Fla., o-5 fms. Pleis-
tocene, Knapp’s No. 3 well, Terrebonne Parish, [a., at
1150-1200 feet. Perhaps this species is a dwarfed form of
atrostyla Dall.
cerina Kurtz & Stimpson, Pr.Bost. Soc.,4, p. 115,1851;(as Pleuro-
toma); Hilgard, Rept., Chief Engineers to Sec., War., p.
4I BULLETIN 38 at
Bos 1870; HOUSe OL Rep wah xn DOCAMu pian 2h Dal SS OMMor
Dall Pr Uy So NAM Gh ne28. 82 ilnvonun Maniosnps
Blow plwi22)) faa sae Mall eB illy ea Olas Nien Danilo
plait. 16a, 180.) Vanattan ier. ALONE S: ieinllaiccs hon sie
LOO3;: Johnson), Bost. ,ooc. IN. EA Ocesn Pan einen AG.
IQI5.
Distribution.—Massachusetts to Texas. Gulf coast: Cedar
Keys, St. Joseph’s Bay, Crooked Island, Florida; Galveston
and Corpus Christi, Texas. Pleistocene, New Orleans
well of 1856; Lake Borgne borings; Grand Chenier; Knapp’s
wells, Terrebonne Parish, No. 2 well at 1190-1439, No. 3
at 1500-1525 feet.
cerinella Dall, Bull. M. C. Z., 18, p. 113, 1889; Bull. 37, U. S. N.
MDs KOZ WoO:
Distribution.—Hatteras to West Florida and Galveston, 14-
22 fms. Pleistocene, Knapp’s No. 3 well, Terrebonne
Parish, La., at 1500-1525 feet; North Creek, Fla.
quadrata Reeve, Pr. Zool. Soc., p. 114, 1845, (as Clathurella) ;
Dnyonti Man. 6).p:/278 platen sen SAS malie Baller
a SNE MES OS mO2 8 oO.
Distribution.—Hatteras to Yucatan. Dall regards diminuta
C. B. Adams (Conch. Contr., p. 63) asa variety ranging
from Hatterasto Florida Keys and West Fla.
melanitica Dall, Bull. 37, U. S. N. M., p. 102, 1889.
Distribution.—Hatteras to Haiti and West Florida.
antonia Dall, Bull. M. C. Z., 9, p. 59, 1881; 18, p. 116, pl. 10, f.
Ay planta if 11. So. iBull ey ane oMING VLD. OZ) hp leuimo spt
AvOlMMntte ATL SSO.
Distribution.—Yucatan Strait, 640 fms. Also Guadeloupe.
peripla Dall, Bull. M. C. Z., 9, p. 68, Aug., 1881; (as Drzllza) ;
It) [Oy SES OL MIM Eas MyMac/y ) oxoy>.JeibUUL, Ziaial hoy. Oe OLlANcrg, st) 107),
1889.
73 MOLLUSCA OF THE GULF OF MEXICO 42
chyta Watson, Jour. Linn. Soc., 15, p. 466, Nov., 1881 (as
Defrancia); Rept. Chall. Gastr., p. 358, pl. 18, f. 4, 1885.
Distribution.—Vucatan Strait, 640 fms.
elusiva Dall, Bull. M. C. Z., 9, p. p. 69, Aug., 1881, (as Drillia);
TOUS ply P27 SOU 27 OU Sa NM Ds TO2,
leant 7. SO)
Distribution.—Yucatan Strait, 640 fms. Perhaps identical
with Defrancia perpauxilla Watson.
bandella Dall, Bull. M. C. Z., 9, p. 59, 1881; Verrill, Tr. Conn.
Neada OND 250) 10457 Dalle BullvomMin CH eZsSh ype Glo mepll.
Romig Son Bulla. Us oo Ney MEY pa ro2t) plano sien p ls
Govt 73.80.
Pleurotomella diomede@ Verrill and Smith, Tr. Conn. Acad.,
Opals 2, pli zi ht.) 5, 5a boda.
Distribution.—East coast United States and Gulf of Mexico,
Blake Station 47, at 321 fms, Greatest depth 2100 fms.
comatotropis Dall, Bull. M.C. Z., 9, p. 58, 1881; Verrill, Tr.
Conme Acad. p.452, 17°82; Dall sBallMe C2) 75.18) ps1 bo,
ples 8o7 NB ull) 27) Un Sa Ne ME p. 102, ply nity L2,
Plait. 8. plO1mi i 77) (89: Johnson, Occ! Pal) Bost: Sec:,
NEVE 7. (P- 1405, TOLS.
tiara Watson, Jour. Linn. Soc., p. 440, 1881; Chall. Gastr.,
Pp. 347, pl. 21, f. 7, °85.
Taranis pulchella Verrill, ’82 and 84.
Distribution.—Massachussetts to Barbados, 50-1075 fms.
Also Yucatan Strait, 640 fms.
Helagiaywal Bull MNCZ. 9, p 1Or, Lost, 1S,1p. (uly, Plv nL, 1.
Oso Bally 277-14 So NEN pO 2esple tn falaynt SO):
Distribution.—Gulf of Mexico, Blake Station 44, in 539 fms.,
bottom temperature 39° F. Also Antillean.
43 BULLETIN 38 73
exsculpita Watson, Jour. Linn. Soc., p. 247, 1882, (as Drillia);
Chall 'Gastri py amy Pin aqetbe Nisin (asiG@/orela) Dall
Bulli Me CY Zr Shon plains on Sor Maula ah mos
PLY TS VON oo.
Distribution.—Vucatan Strait, 640 fms. Also Santa Cruz,
Wie
subsida’ Dall, Bull) M:C..Z., 0, p. 62; 18815 18) py 108, pliin2.t
3h (SOs bBulliage) Uli S uN Men pato2 plu i2nit ena inSOE
Distribution.—Gulf of Mexico, West of Florida, 339 fms.
Also Cuba.
para Dall Balle Ni CZ on) oa WSS Sil: eS) Nope i sols si pke una
1889.
Distribution.—VYucatan Strait, 640 fms. Also Martinique,
502 fms,
Genus CYTHARELLA Monterosato
cymeila Dall, Bull. M. C. Z., 18, pl. 12, f. 4, 1889, (as Cythara) ;
Bolla 7 Wii S.. IN ME pu oot plain A So.
Distribution.—Gulf of Mexico, 220 fms. Also Antillean.
galvestonensis Harris, Bull. Am. Pal., vol. 1, p. 100, 1895.
Distridbution.—Upper Miocene, Galveston well, at 2236-
2871 feet.
Genus DAPHNELLA Hinds
jeucophlegma Dall, Bull. M. C. Z., 9, p. 70, 1881; 18, p. 102, pl.
Cito wis sou
Distribution.—Gulf of Mexico, Blake Station 2, in 805 fms.
(Eubela) limacina Dall, Bull. M. C. Z., 9, p. 55, Aug., 1881; Ver-
Tl ire Conn: VA caddis Nip uuias eMs2. NON Dl 205 O4 mall:
Ball Mi CC.) 22.118) p./106;plion inion So.) Bulle) 27.V Oa osuINe
Val LOOMUD LO it ON sop
hormophora Watson, Jour. Linn. Soc., 15, p. 457, Nov. 1881;
Challis Gastra. (pis 5a) polyoma nines:
74 MOoLLUSCA OF THE GULF OF MEXICO 44
Distribution.—Massachussetts to Pernambuco. Gulf of
Mexico, Yucatan Strait, 640 fms. and Blake Station 2, west
of Florida, at 895 fms.
Genus CANCELLARIA Lamarck
reticulata Linnzeus, Syst. Nat., ed. XII, p. 1190, 1767, (as
Voluta); Sowerby, Thesaurus, 2, p. 442, pl. 92, f. 17, 1848;
iieyonn Wan.))7,p. 69, pl.) 2))t,.25, 26,853, Dall) Bull: Me:
C48 sp. 129,\ 89.) Vanatta; hr. AYN |S.) Piila.) 155, p:
757; 1903.
Distribution.—Hatteras to Guadeloupe, 5-30 fms. Gulf
coast: Crooked Isl., Charlotte Harbor and Sarasota Bay,
West Fla.; Chandeleurs, La.; Galveston, Tex. Pleistocene,
New Orleans Pumping Station No, 7.
Note.—The fossil species, C. conradiana Dall, (Tr. Wagner
Inst., 3, p. 42, 1890), is closely allied to C. 7efculata.
galvestonensis Harris, Bull. Am. Pal., vol. 1, p. 100, pl. 3, f. 11,
1895.
Distribution.—Upper Miocene, Galveston well at 2552-2600
feet.
(Norona) clavatula Sowerby, Pr. Zool. Soc,, p. 52, 1832; Conch.
Meee 2h a2-wlinyon) i Mian si 7 pale 75a ipl -hAs uy tenOds Vso:
Inlanris.) Bulls (Am) (Paly vol!) tepeoon os; Wall) Bn U.S.
INN 3375 (200, LO LO.
Distribution.—Recent, Panama to Paita, Peru. Upper Mio-
cene, Galveston well at 2552-2600 feet, (Harris). Notin
the recent Atlantic fauna.
(Trigonostoma) agassizi Dall, Bull. M. C Z., 18, p. 130, pl. 35,
ME MMSSO Mt o7eW. Ss IN MEY pu monmipley ai5iih.) auc 89:
Distribution.—Cape Lookout, N.C., 18 fms. Gulf of Mex-
ico between Mississippi delta and Cedar Keys, 25 fms.
(Trigonostoma) tenera Philippi, Zeitsch. fur. Mal., 5. p. 24, °48;
A5 BULLETIN 38 73
Dall, Bulla 7 Us Sane Ny Mom onsnS ori dlie NN ile OE a Eine
43, 1890.
stimpsont Calkins, Pr. Dav. A. S., 2, p. 250, pl. 8,£. 4, 5, 1878:
Distribution.—Gulf of Mexico, Yucatan region. Pliocene,
Caloosahatchie beds, Fla.
Genus ABDMETE Kroyer
microscopica Dall, Bull. M. C. Z., 18, p. 131, 1889,(as 7rigon-
ostoma sd). Bull a7. U S) IN Mes patos 1 OOnwe i Oo neNe
NEV 24 pA sO4.) plii2o ut {Anat @@2)
Distribution.—Campeche Bank, 200 fms. Also off Fernan-
dina and Cuba.
Genus OLIVA Martyn
circinata Marrat, Sowerby’s Thes. Conch., 4, Oliva, p. 21, pl. 17,
f. 277, 1880; Johnson, Nautilus, 24, p. 123, Mar., 1911.
litterata Lamarck, Ann. du Mus., 16, p. 315, 1810; Not of Bol-
tem 1708. | Hilgard) House) or Repyik x MDoc) 1) pts 2M.
S8ori78)) Dall wBulls 27. Uae SupiNeaev lenin oO planation
$a)) 89; Singley, 4th) Ann) dReptamexa GS.) p. sacung2:
Wanatta,)Pr. A. Ni S) Phila og or sloos.
carvolimensts Conrad,’ Pr. A. Ne Swieiila 4s ip, 563,203:
Distribution. —Hatteras to the West Indies, 0-2 fms. Gulf
coast: Franklin and Calhoun Counties, Fla., Chandeleurs,
Point on Fer, Cameron, La.; Galveston, Corpus Christi,
Matagorda, Tex. Pleistocene, New Orleans artesian well of
1856, Lake Borgne borings, Knapp’s wells, Terrebonne
Parish, No. 1 well at 2000-2450, No. 2 at 1780-1800, No. 3
at 1150-2029 feet. Pliocene, Caloosahatchie beds, Fla.
liodes Dall, Tr. Wagner Inst. Sci., 3, p. 1576, pl. 58, 1903.
Distribution.—Lower Miocene, Chipola marl, West Florida;
Bascom No. 2 well, Mobile, Alabama, at 1241 feet, Chipola
horizon.
76 MOLLUSCA OF THE GULF OF MEXICO 46
Genus OLIVELLA Swainson
mutica Say, Jour. A. N.S. Phila., 2, p. 228, 1822; Hilgard, Rept.
Chief Engineers to Sec. War, p. 358, ’70, House of Rep.,
EomWOCle pt 2) pi Sho, 78; Wally Pry Wiis. NG Me, p:
B24 Se) Bull e327) UL Se Ne ME pp. \1o6)) pl ee ee OS OF
eV AMeN So0.4\5,)) 290%) Vanattay) Pr, A. ING) S. Pinta. 55),
Pavi571 1903; Deussen, U2) S: Gs S:) Water- eine es
INOW ZC5) (D:77; Told.
Distribution.—North Carolina to Trinidad Isl. Gulf coast:
Recent, Cedar Keys, St. Mark’s and St. Joseph’s Bay, Fla.;
. Galveston and Corpus Christi, Tex. Pleistocene, New Or-
leans artesian well of 1856, Lake Borgne borings; Knapp’s
wells, Terrebonne Parish, No. 1 at 1600-1700, No. 2 at 1050-
1842, No. 3 at 1150-1839 feet; North Creek, Manatee and
Labelle, Fla. Pliocene, Caloosahatchie beds, Fla., and New
Orleans Gymnasium well at 1200 feet. Miocene, Gilbert
well No. 10, Bateson, Tex. at 323 feet.
pusilla Marrat, Thes. Conch., t. 21, f. 356-358, 71; Tryon, Man.
Wonchipes.op.1645) ply gent. 45) he Seni Vanattas bi ANG
Seaelailane SS. 171575) 203.
Disiribution.—Cited by Vanatta from St. Joseph’s Bay and
Crooked Isl., West Fla.
nivea Gmelin, Linn. Syst. Nat., ed. XIEI5-p. 3442, 1792; Reeve,
Sonchaylcony, it 640a, 150; Tryon, Manis) pe 67) pl. 15, f
fences Dall Bill a7) UW. 3. Ns (Meaiipe roo, So. Dall Sz
Simpson, Bull. U.S. Kish Com., 1, p.) 393) or.
Distribution.—Sarasota, West Fla. to Haiti, 27-805 fms.
jaspidea Gmelin, Syst. Nat.,ed. 13,p. 3442, 1792; Reeve, Conch.
icone 6, p:)22, 0. 58,1850: Dall & Simpson, Bull, WU), S.) Fish
Comes) ph 202, ‘Toor.
Distribution.—Hatteras to Brazil and West Florida, 27-805
fms.
floralia Duclos, in Chenu, Ill. Conch., p.6; Tryon, Man., 5, p.
47 BULLETIN 38 77
OS plein.) fh: / 2. an ba SS a aallllmin Stilley) aru aS upIN emy os Do
106, ’8o.
Distribution.—Hatteras to Tortola and West Florida.
eulacta) Dall, Tr.) Wagner) Inst.) Seis) 3). 5 76)) pola ison tage
£903.
Distribution.—Lower Miocene, Chipola marl, Fla, and Mo-
bile Oil Co’s No. 2 Bascom well near Mobile, Alabama, at
1241 feet, Chipola horizon.
galvestonensis Harris, Bull. Am. Pal., vol. 1, p. roo, pl. 3, f. 13,
1895.
Distribution.—Upper Miocene, Galveston well at 2410-2871
feet!
subtexana Harris, Bull. Am. Pal., vol. 1, p. 101, pl. 4, f. 1,
1895.
Distribution.—Upper Miocene, Galveston well at 2552-2871
feet
Genus MARGINELLA Lamarck
cared otorer, Jour. Bost))\Soc) .NEiT ps AG5, plionit 2A
1837; Sowerby, Thes., f. 103;) Tryon, Man., 5, p. 33, pl. 10,
P86; 1882.) Dall: Bull srw OE SNe Vien paytOoN SO:
Distribution.—Charlotte Harbor, West Fla., Florida Keys
and Antilles.
storeria Couthouy, Jour. Bost. Soc. N. H., 1, p. 440, pl. 9,f. 1,2,
18373) Dall Ball 37. SUL Sain ven: OOS 89:
Distribution.—Gulf of Mexico, West Fla., to Colon.
cassis Dall, Bull) (M,C. Zin Sexpime7in ply 35, \\\t../ 8 1889. ule
BTM TOO. Dl 2 Shit owas
Distribution.—Gulf of Mexico, Lat.25° N., Lon. 84° W., at
tol fms. Also off Cuba.
apicinca Menke, Syn. Meth. Moll., p. 87, 1828; Tryon, Man., 5,
DA BS,ipl. 10, ef. So) 28a. yDalleiB all se S46 ING Mo aTOos
78 MoLLUSCA OF THE GULF OF MEXICO 48
"89; Singley, ath Ann. Rept. Tex., p. 333, 92; Vanatta,
PrivA. N.S) Phila’, 55, p. 757, 1903; Vaughan, 2nd) Ann.
Rept. Fla..Geol. Surv., p. 148, 1909.
conoitdalis Reeve, Conch. Icon., 15, pl. 18, f. 87.
Distribution.—Hatteras to Jamaica, Gulf coast: West Fla.
and Texas, Pleistocene, Manatee, Orient and Labelle, West
Florida.
pellucida Pfeiffer, Wiegmann’s Archiv; 1, p. 258, 1840; Tryon,
Mansi pl aa pl.) TO fon og mDall bull ian. ss.
ME pa LOOM NSO:
Distribution.—Sarasota, West Fla. and West Indies.
watsoni Dall, Bull. M. C. Z., 9, p, 71, 1881; 18, p. 137, pl. 19, f.
Bplansss by 2486" Bull: 27. Wis wINen Mey pu OG spleToiHt:
Bullen 28h, fein 18 Oe
Distribution.—Yucatan Strait, 640 fms. Also off Havana,
220-805 fms.
virginiana Conrad, Am. Jour. Conch., 4, p. 67, pl. 5, f. 4, 1868,
(as Prunum); Dall, Bull. 37, U. S. N. M., p. 106, ’89; Tr.
NV Salen Seizes 43.) 190}
Distribution.—Recent, North Carolina to Yucatan, 10-294
fms. Pliocene, Caloosahatchie beds, Fla. Miocene, James
River, Va.
denticulata Conrad, Jour. A. N. S. Phila., 6, p. 225, pl. 9, f. 21,
u83o0.) | @Not of Late, /73); Dall) Bullsiay iW! S.)) N° IMs, p:
OSM SOs: WV LS. (35 Pp. Sisuplsnsi ty ommrooo:
Distribution.—Recent, Hatteras to Barbados. Also West
Florida, 5-294 fms. Pliocene, Caloosahatchie beds, Upper
Miocene, Maryland and Virginia.
opalina Stearns, Pr. Bost. Soc. N. H., 15, p. 21, 1872; Dall, Pr.
CN SwNE MS 6ap els 24. 183) Bullea7ipe TOS." 8Q,Cas\ acre
ticulata var. opalina).
Distribution.—Hatteras to Barbados. Recent, Tampa (type
49 BULLETIN 38 7S
locality) and Cedar Keys, 14 fms. Also Barbados, 100 fms.
aureocincta Stearns, Pr. Bost. Soc. N. H., 15, p. 22, 1872; Dall,
Balle 27.1. SON Mie apr rsh iSOe arly Sea ONS 2
1890.
Distribution.—Virginia to West Florida, 3-44 fms. Pliocene,
Caloosahatchie beds.
minuta Pfeiffer, Wiegmann’s Archiv. fur Naturg.1, p. 259, 1840;
Dryon, Mans 5.) p. 43), pl. 12h i oowmmeano Dalla bull 27 mr
SHINGO. O81 SO!
Distribution.—Mediterranean, West Indies, East and West
Florida, 5 -294 fms. Miocene to Recent. Also Pleistocene,
southern Europe and of North Creek, West Fla.
minima Guilding, Sowerby, Thes. Conch., 1, p. 388, pl. 78, f.
220u18a6- Wall Pre U. /Si Naive toMpa 22464) 182i tayore
Mans .p: 44, pl. 12) £./62 8a nDallienBullea en sS: 0 Ni vier
p. 108,’ 80.
Distribution.—Cape Lookout, N. C., to Haiti. Also West
Florida at Cedar Keys, 0-22 fms.
Note.—Tryon placed this species in synonymy of mzz2uta
Pfr. According to Moerch mznzma is asynonym of lavalleana
d’Orbigny, (Moll. Cuba, 2, p. rot, pl. 20, f. 36-38, 1842).
succinea Conrad, Pr. A. N.S) Philavi26.) pl. 1, f- 17, 1840;
(iryon. Man. 5,'p. 34, pli iromnmogmcss, Dall) Bull: 27 -aUs
SAONEYMES ps 108, ple no, fs 64 soy
Distribution.—Fernandina, East Fla., to Sombrero, W. I.
Gulf coast at Tampa, 70-1002 fms. Placed by Tryon in
synonymy with zztzda Hinds.
fusinayDall, Bull. M. C. Zo p72 ir88r: 18. p. 138. pla toy t:
ANS: Mati. 27)\p. (TOON PION n SO,
Distribution.—Yucatan Strait, 640 fms. Also off Fernan-
dina, Fla.
Bo MOLLUSCA OF THE GULF OF MEXICO xe
yucatecana Dall, Bull. M. C. Z., 9, p. 72, 1881; 18, p. 138, pl.
NOM USHE SOV BUlla OK S20 NV py mOoN pI TOR fis,
1889.
Distribution.—VYucatan Strait, 640 fms. Also Sand Key,
Rilaen 25) bas:
seminula Dall, Bull. M. C. ZO puT Za) USSU LSe iP) HO, (Oke LO),
Hen So Bull. 27) U.S. Neve py los. pl Loli, 25.789.
Distribution.—VYucatan Strait, 640 fms. Also Antillean.
rostrata Redfield, Cat. Marg., Am. Jour. Conch., 6, p. 246,
@iotelwirs 70. Dall Bull Me CZ. 1 8)ypuitia7 | 805 re) Wie
SA ahs 5O,k SOO:
oblonga Sowerby, Thesaurus, pl. 76, f. 106, 108. j0N
oblonga of Swainson.
Distribution.—Gulf of Mexico, Lat. 23° N., Lon. 89° W.,
in 84 fms. Pliocene, Caloosahatchie beds.
(Volvarina) avena Valenciennes, in Kiener, Coq. Viv. arginella,
PW plow tl. 24.) 1841: Sowerby, WaeswConchyi1) p, sor,
DEO te1Zo0, 49>) Dall) Bull) 27) Ws Suen Me ps) 108; 180:
Dall & Simpson, Bull. U. S. Fish Com., 1, p. 394, 1901;
Wauchans Publ. No. 133, Carn. inst pez 1910:
Distribution.— Key West to Colon. Also Texas region,
Gulf of Mexico. Pliocene to Recent. Pleistocene, La-
bella, West Fla.
(Volutella) ovuliformis d’Orbigny, Moll. Cuba. 2, p. 101, pl. 20,
feasts. 6845 Dall Bull 37.) Usa Say NGmMiNs {ip TOS... 289;
Dallesa Simpson Bull Wes) hishh) Comets n)31'395,) Look:
Vaughan, 2nd Ann. Rept. Fla. G. 5., p. 148, I910.
Distribution.—Cape Fear, N. C., to Guadeloupe Isl. Also
West Florida. Pleistocene, Manatee, West Fla.
(Volutella) hadria Dall, Bull. M. C. Z., 18, p. 142, 1889; Bull.37,
Oe SONG MEN pastoS.))7380,
SI BULLETIN 38 $x
lacrimula Dall, Pr. U. S. N. M., 6, p. 324, 1883. Not lacrimula
Goulds) Pr. Bost. Soe Ni El) Si pi 2dn L8O2:
Distribution.—Cedar Keys, West Fla. Collected by Hemp-
hill on mud flats. Also Charlotte Harbor, West Fla.
Genus VOLUTA Linnzeus
virescens Solander, Dillw. Desc. Cat., 1, p. 562, Sowerby, Thes.
Lplis2) fH 78 Ey Ony Mane Aa PanOqwph 2a uiiaa sn ieSZe
Dally Balli37. Us So ING Me panos Son
Distribution.—Texas to Cartagena, Colombia.
Genus SCAPHELLA Swainson
junonia Hwass, Chemnitz, Conch. Cab. 11, p. 16, pl. 177, f. 1703,
L7 Oar pip wr 7Os5s Dall Bal VCH ZT Sak. AS. pl aann te
5ae-en SO; Bulls 27. (Ul S NN A Men Onno sla gau sts Cem
1889.
Distribution.—North Carolina to Nassau. Also Gulf of
Mexico!) Wat! 25° Ni. Lonij82-. Wat 26) fms:
Genus AURINIA H. and A. Adams
dubia Broderip, Zool) \Jour.,) gipe wemupliai ft.) 1, 1828)).(as
Voluta); Reeve, Conch. Icon., Voluta, pl. 22, f. 59, 1849;
Tryon, Man., 4, p. 90, pl. 27, f. 77, 81, 1882; Dohrn, Jahrb.
Mal! Ges.;)'6,,p: 150, pk. ait eaeai 7G Dall) Bull. Mi CZ
CS Ps USL. SOs BU 27. KU Eisai NNO. TTO;) Sore ra Wase
NE 24D. Soa.) pli zoutiinramoo2)
Frusus tesellatus Schubert & Wagner, Suppl. Bd. Mart. und
Chemn., Conch. Cab., 1829; Kiener and Reeve; but not of
Zekeli and Pictet, 1852.
mutabilis ‘Tuomey & Holmes, 1856. Not of Conrad, 1841.
Distribution.—Hatteras to Cape Florida. Gulf of Mexico
between Mississippi delta and Cedar Keys, at 111 fms.
ropusta Dall Bally My CZ yrs apt 53, ples, f) 24)’ so. Bulle
BT De LLO Mp.) 250 Lua sOE
82 MOLLUSCA OF THE GULF OF MEXICO 52
Distribution.—Gulf of Mexico, Lat. 28° N., Lon. 86° W.,
at 280 fms. Also off Cuba, 119 fms.
Genus MITRA Lamarck
gemmata Sowerby, Thes. Conch., sp. 334, 1874; Tryon, Man.,
AMM Too Plu 5S) toss ross Oa Bully, 277 Ui SiN My
p. 110, ’89; (as var. of hkanleyz Dohrn): Dall & Simpson,
Balle WS) ish Com\, 0,’ pi 396) reer.
Distribution.—Charlotte Harbor, West Florida, to Jamaica.
straminea A. Adams, Pr. Zool. Soc., p. 132, 1851; Sowerby,
dthess Conch. t..561; Tryon, Many, 4. wae) pl. ar, ft. 183,
HSS OA sBull 7. Ui. 95) Nao Meio hO: .8Or
Distribution.—Gulf of Mexico, west of Florida.
wandoensis Holmes, Post-Plioc. Fos. S. Car., p. 77, pl. 12, f. 10,
10a, 1860, (as Volutomitya); Dall. Bull. 37, U. S. N. M.,p.
LOMAS OMT NY olla)... 34) 9! O21 LSGe,
Distribution.—Cape Hatteras to Florida Straits and West
Florida, 14-80 fms. Very closely allied to and probably
identical with J7. rusht Dall, (Conch. Exch., 2. p. 9, 1887;
Bull. M. C. Z., 18, p. 160, ’89) from the Gulf of Mexico, 27
fms., U. S. Fish Com. Station 2372.
SiviiaeWallk Bull Mi CiiZ.. 18, Pp. 150) plenmse eon 1880 Bull,
BRO ONG Me ips Tro) plu Tes dOmses
Distribution.—Gulf of Mexico, west of Florida, 19 fms.
Also Antillean,
(Conomitra) blakeana Dall, Bull. M.C. Z., 18, p. 163, 1889; Bull.
By On Sa Nine Mere Ds Isto}, 80)
Distribution.—Recent, Yucatan Strait, 640 fms. Closely
related to the fossil species, stamzzea Conrad and fusoides
Wear
Genus MITROMORPHA A. Adams
pygmaea Dall, Tr. Wagner Inst. Sci., 3, p. 95, pl. 10, f. 3, 1890.
53 BULLETIN 38 83
Distribution.—Type from Caloosahatchie marl, Fla., Plio-
cene. Also Bascom No.1 well, near Mobile, Alabama, at
1500-1556 feet. Miocene. Not in the Recent fauna.
Genus FASCISLARIA Lamarck
gigantea Kiener. Icon. Coq. Viv., p. 5. pl. 10, 11, 1840; Tryon,
Man 2. p75; (pls GO. tama Slam atl eb unl Ieee sy IN eave
PMT? Sora NV LS.) 2p mon OoOu time leven tiny Acmns
Rept: Tex. ’G.'S.;)p. 234,92; Walliéz/Simpson, Bull) U27s:
Fish: Com.))1,\p)207,)) Teo Waughaniiadi Ann. Rept.vElas
G. 3., p. 149, 1909; Johnson, Nautilus, p. 44, Oct., 1919.
Distribution.—Hatteras to Brazil. o-1Io fms. Gulf coast;
West Florida; Point au Fer. La.; Corpus Christi and Mat-
agorda Isl., Tex. Pleistocene, New Orleans pumping Sta-
tion No. 7, North Creek and Manatee, West Fla. Pliocene,
Caloosahatchie River marl.
tulipa Linneeus, Syst. Nat. ed X, p. 754, 1758, (as Murex);
Kienenr. Cog) Vive, ps2) pla mz s Ao.) dvnyon\) Manis eau:
FAN DIAS ON te hon) ron) Oa MeN iS oe si. SOTA IUN Alte
iGO; Nanatta, SPraial uN e Samaalan nis 5) Duis Tal OOgr
Johnson, Nautilus, p. 46, Oct., 1919.
Distribution.—North Carolina to Colombia and Antilles.o-10
fms. Gulf coast: Cedar Keys, St. Joseph’s Bay and St.
Marks, West Fla., Texas coast. Pleistocene, North Creek,
Fla. Pliocene, Caloosahatchie beds.
distans Lamarck, An. s. Vert., 7, p.119, 1822; Tuomey & Holmes,
PLoS SCH ph 150 plesowena tos 7) iiloardy) Houselok
Reps ix) MOC.) 1, peep esomenonn Dall, lic Well Seaicnmny
102, pl. 7, f. 10, ’90; Singley, 4th Ann. Rept. Geol. Surv.
Tex.,'p. 334, ’92; Mitchell,List. Tex. Sh., p. 16; Vaughan,
Publ. No. 133, Carn. Inst., p. 171, I910; Johnson, Nautilus,
DN47 NOCE TOTO)
Distribution.—Hatteras to Yucatan o-54 fms. Gulf coast:
Cedar Keys, St, Marks, Fla.; Cameron and Chandeleurs,
54 MOLLUSCA OF THE GULF OF MEXICO 54
La.; Galveston, Matagorda, Corpus Christi, and Mustang
Isl., Texas. Pleistocene, New Orleans well of 1856, and
pumping station No.7; Orient and Labelle, West Fla.
‘The Pliocene shell of the Caloosahatchie beds is the related,
but slenderer, /. apicina Dall.
(Mesorhytis) meekiana Dall, Bull. M. C. Z., 18, p. 172, pl. 36, f.
Fin WSSOS) EUG oe Oo IN IE soy, si shzA iol aXe). aha ale cXG),
Distribution.—Gulf of Mexico, Blake Stations 16 and 20, at
292 and 220 fms. Also off Cuba, 400 fms. ‘The first living
species of Mesorhytis known, the type being the Cretaceous
species, gvacilentts Meek.
Genus FUSINUS Rafinesque
timessus Dall, Bull. M. C. Z., 18, p. 166, 1889; (as Fuses); Bull.
COE NEE ep 112.02 89.
Distribution.—Gulf of Mexico, between Mississippi delta
and Cedar Keys, 60 fms.; between Cedar Keys and Dry
Tortugas, 27 fms. Also near Cuba.
eucosmius Dall Bull MeeC. Z)) 18) pit 67 plans ,it.) 5, 18805
IBLUDLS B75 \Oy sha FON eae Ns eluls cshay,
Distribution.—Gulf of Mexico, west of Florida. Also Bar-
bados, 27-111 fms.
couel Petit, Journ de Conch, 4, pr 2torsplly 8) fim) 18535) Dall,
Balle Ne CZ e168) ps 167, 789)
Distribution.—Gulf of Mexico, between Tampa and Dry
Tortugas, 26 fms.
alcimus Dall, Bull. M. C. Z., 18, p. 170, 1889.
Distribution.—Gulf of Mexico, 100 miles north of Yucatan,
g5 fms.
amiantus Dall, Bull. M. C. Z., 18, p. 169, pl. 15, f. 11, 1889; Bull.
Boy NUIG Sio AN OM GAN ove esi zae Wo) Me ell/adids Uhksyay,
Distribution.—Gulf of Mexico, Blake station 2, at 805 fms.
O15) BULLETIN 38 85
aepynotus Dall, Bull. M. C. Z., 18, p. 169, 1889.
Distribution.—Gulf of Mexico, 84 fms. Also Antillean.
amphiurgus Dall, Bull. M. C. Z., 18, p. 171, 1889.
Distribution.—Gulf of Mexico, Blake station 45, at ror fms.
Genus LEUCOZONIA Gray
cingulifera Lamarck, 7, p. 107, 1822, (as Zurbinella); Reeve,
Conch. Icon., 4, pl. 3, f. 17, 1847; Dall and Simpson, Buil.
We) Ss) hish? Come. 1) 7p: 208) r901
Distribution.—Sarasota, Fla., to Texas and Brazil. Also
Antillean.
ocellata Gmelin, Syst. Nat., p. 3488, 1792, (as Buccinum) ;Reeve,
Conch: Icon.; 4, pl. 8,°f 28, 18475) Dall and Simpson; Bull?
Weis wish Com:;) 1; p.208, gor
Distribution.—Cedar Keys, Fla., to Guadeloupe, W. I.
Genus BUSYCON Bolten
pyrum Dillwyn, Desc. Cat. Rec. Sh., 1, p. 485, 1817; Dall, Bull.
Bre Wise ON Mp) 2 Sonia bei Md Sil c2\ Tse TOO Or
Singley, 4th Ann. Rept. G. Sex) \p. 334,92; Vanatta,
Pre VAL ING S,) Phila p. 757 roos i aughan: : ulblamnacs
Carn, Inst., p. 171, 19Io.
Distribution.—Hatteras to Gulf of Mexico. Upper Miocene
to Recent. Gulf coast: Cedar Keys, Fort Barranca, St.
Mark’s, Fla.; Cameron, &c. La.; Galveston, Matagorda,
Corpus Christi, &c., Texas. Pleistocene of Louisiana at
Grand Chenier and New Orleans pumping station No. 7.
A high spired variety isfound in abundance at Cedar Keys,
Point au Fer, Cameron and Galveston, and in the Pleistocene
of Grand Chenier, La., North Creek, Labelle, Manatee and
Orient, West Fla.
canaliculatum Linnzeus, Syst. Nat., p. 753, 1758, (as Wurex);
iolmes,Post- Pl) Hos) Si) Chip) /66.;npluann his) 18 5oe ale
86
MOLLUSCA OF THE GULF OF MEXICO 56
gaid, House of Rep., Ex. Doe., 1, pt. 2, p, 886, 78; Dall,
Balke OL Sua Meni pnt? pla 7 akin toon Clark Plers-
tocene Md., p. 180,pls. 46, 47, 48, 1906; Johnson, Occ. Pa.
BOSH SOC NE Ee i7,.) Dal sOQninOiliss
Distribution.—Cape Cod to West Florida and Texas.
Pliocene (Carolinas). Pleistocene, New Orleans well of ©
1856 (Hilgard).
perversum Linnzeus, Syst. Nat., p. 1222, 1766, (as Murex);
ditionmey,&) Elolmes, Pl. Kosi 'SCiipy 145; pl. 204 f: 3.1856;
Hilgard, House of Rep. Ex. Doc. 1, pt. 2, p. 886, ’78; Dall,
ree S38 0)) 116,\\/o07)) Vanatta wer ay NES. bhilas,
SDaTsie Loos N auchant PubitinegatCarnainst sn pi ij,
IQIO.
Distribution.—Hatteras to Cuba. Miocene (Carolinas) to Re-
cent, o-3 fms. Gulf coast: Cedar Keys, St. Marks, &c.,
Fla.; Point au Fer, Cameron, La.; Galveston and Corpus
Christi, Tex. Exceedingly abundant in the Pleistocene of
Grand Chenier, La., and found by Hilgard in the Lake Bor-
gue borings; also of Manatee, Orient, and Labelle, West Fla. ;
Pliocene, Caloosahatchie beds, Fla.
carica Gmelin, Syst. Nat., p. 3545, 1792, (as Wurex); Holmes,
RostsPiihos, 5). Cy p65, pli 11, Pemaso. Dall Bullies,
NEE Ma con 2) olla. £1.) 7897). Clarkebletstocene ss Mid= jp:
179, pls. 43-45, 1906; Johnson-—Oec. Pa. Bost. Soc. N. H.,
7) P- 139, 1915.
Distribution.—Massachussetts to the Antilles. o-10 fms.
Gulf coast, West Florida and Galveston, Tex.
eliceans Montfort, Conch. Syst. 303; Dall, Bull. 37, U. S. N. M.,
Os, BE, SO)
Distribution.—South Carolina to Campeche. o-5 fms.
Genus MELONGENA Schumacher
corena Gmelin, Syst. Nat., p. 3552, 1768; Tryon, Man., 3, p.
LOSMAD OI 1OD-2O2 18Sry Dalla Mla Seige 12 TOO);
57 BULLETIN 38 87
Vanatta,Pr Au iN) SS) Rinilal iss). 7757.) ko033) Vatenany 2k
Ann. Rept. G. S. Fla., p. 148, I909.
Distribution.—Florida Keys to Yucatan and the Antilles.
Gulf coast: Cedar Keys, St. Marks, St. Joseph Bay, Fla.;
La.; and Tex. Pleistocene, North Creek, Orient and La-
belle, West Fla.
melongena Linnzeus, Syst. Nat. ed XII., p. 1220; Tryon, Man.,
Bu PLO7s Platt. LOZ. TOS moon Dallumb illus 7A: Sie Nte
Me peirr2)). 805) Harris Bulli AqM ealeavole i lipsnt Or nOuys
Distribution.—Florida to Grenada. o-50 fms. Gulf of
Mexico, Texas region. Also Upper Miocene, Galveston
well at 2448-2465 feet (Harris).
Genus CANTHARUS Bolten
tinctus Conrad, Pr. A. N. S. Phila., p. 25, pl. 1, f. 9, 1846, (as
Pola) Dall) Bull. 27 Us SaNeRMeNn point oy So) Case fizz
tonidea); Dall and Simpson, Bull. U. S. Fish Com., 1, p.
399, 190T.
Distribution.—Hatteras to Vera Cruz. Gulf coast: Cedar
Keys, Fla., and Matagorda, Texas. Pleistocene, North
Creek, West Fla.
cancellaria Conrad, Pr. A. N.S. Phila., p. 25, pl. 1, f. 12, 1846;
Dalry WL0S. IN) Mover comige mess omeley.” athiAcmr,
Reptyvbexs G.'S.) p. /334wozsanaialbvaset. An) Ni...) elder
Soup 574.1903; Vauchany 2daAnm Rept... bla. (GA Se iis:
148, 1909.
Distribution.—Florida to Isthmus of Panama. Gulf coast:
Cedar Keys, Fla.; Horn Isl., Miss.; Cameron, Point au Fer,
Chandeleurs, La.; Galveston, Tex. Pleistocene, Grand
Chenier and New Orleans pumping station No. 7, and of
Orient, West Fla.
orbignyi Payraudeau, Cat. Moll. de Corse, p. 159, pl. 8, f. 4-6,
To20\ (as Buccnun)|:) Dally Bulle 7 iiss Nis HVE spa menGs
MoLLUSCA OF THE GULF OF MEXICO 58
os)
es)
"89; Dall and Simpson, Bnll. U.S. Fish Com., 1. p. 4oo,
IQOL.
Distribution.—Florida to Yucatan. 22-25 fms. Also Medi-
terranean.
limbatus Philippi, Abbild, 1, 3, pl. 1, f. 9, Tryon, Man., 3, p.
HO MPU M257 TOSd Dall wll ar UE Sen INi WOW i
TOMS Oe
Distribution.—Gulf of Mexico to Cuba.
Genus PHOS Montfort
eandeil d’Orbigny, Moll. Cuba, 2, p. 129, pl. 21, f. 23-25, 1845,
(as Cancellaria); Arango, Fauna Mal. Cub., p. 201, 1878;
Dall, Bull. 37, p. 116, 1889; Dall and Simpson, Bull. U. S.
Fish Com., I, p. 401, 1goI.
Distribution.—Hatteras to Colon and the Antilles. Gulf
of Mexico, between Mississippi delta and Cedar Keys, at 88
fms. Synonyms, fide Dall, are antillarum Petit, 1853 and
grateloupensts Petit, 1853.
parvus C. B. Adams, Contr. Conch., p. 59, 1850, (as 7rzton) ;
Dall, Bull. 37, U. S. N. M,. p. 116; 1880.
LLUGICALUS WA IPT IU S.IN. MES Gye 225Miply TO mio os.
Distribution.—Charlotte Harbor, West Fla., and Texas.
Also Antillean.
galvestonensis Harris, Bull. Am. Pal. vol. 1, p. 102, pl. 4,f. 4, 5,
1895.
Distribution.—Upper Miocene, Galveston well at 2158-2871
feet:
Genus NASSARINA Dall
columbellata Dall, Bull. M. C. Z., 18, p. 182, 1889.
Distribution.—Cape Catoche, Yucatan.
grayi Dall, Bull. M. C. Z., 18, p. 183, pl. 32, f. 12a, 1889; Bull.
STO Se Ne MEE Di TIO) pln aoh aha i QeNee On
59 BULLETIN 38 89
Distribution.—Arrowsmith Bank, Yucatan, 130 fms. Also
Antillean.
bushi Dall, Bull MCA 1s. 182. oluusniayn 2) Uneso.
Distribution.—Gulf of Mexico, Blake station 5, 152-229 fms.
Also off Sand Key, Fia. and Barbados.
Genus ALECTRION Montfort
vibex Say, Jour. A. N. S. Phila., (1), 2, p. 231, 1822, (as WVassa);
Ann} Coneh. pli 57, 10/25), Dalla ull iar aes Se eINMEe Go
116, pl. 50, ¢; 8, 1889; Singley,athWwaAna Rept, Mex! GaSe
Ps335, Fso2: Johnson, Occ: PayBosty socsiNi inl 7. ps nade
IQI5.
Winetens7s Perkins, Pr) Bost.) SoC. 1 3upy L741 SOO.
Distribution.—Cape Cod to Colon. Gulf coast: Cedar
Keys and St. Mark’s, Fla., Point au Fer, La., Galveston,
Corpus Christi, Tex. Pleistocene, New Orleans pumping
station No. 7; Manatee, Orient and Labelle, West Fla.
Pliocene, Caloosahatchie.
ambiguus Montagu, Test. Brit., pl. 9, f. 7, 1803, (as Buccinum);
Dall, Bull. M. C. Z., 18, p. 184, ’89; Not amdigua Dunker
(West African), which is zzcrassata Strom.
alba Say, 1826; candidissima C. B. Adams, Krebs Cat., p. 32.
Distribution.—North Carolina to West Florida and Barbados.
Yucatan Strait, 640 fms. Variety antillarum d’Orb., Ca-
loosahatchie Pliocene.
acutus Say, Jour. A. N.S. Phila., 2. p. 234, 1822, (as Vassa);
Holmes, -Post-Pl. Kos SiC mow 2hplen 12) 01.) 3. ut SOOM Cas
Buccinum); Hilgard, House of Rep. Ex. Doc. 1, pt. 2, p.
886.4757) Dall Bulli27 Wes umNn ME i pe TLON oom S™
Bully pA, | Pails voll pmo2nOsn wanattay peta aeiN wp
Phila., 55, p. 757, 1903; Deussen, Water Supply Pa. 335, p.
77, 1914.
Distribution. —Carolinas to Barbados. Gulf coast: Recent,
St. Joseph’s Bay and Crooked Isl., West Fla.;Point au Fer,
go MOLLUSCA OF THE GULF OF MEXICO 60
Cameron, La.; Galveston and Corpus Christi, Tex. Pleisto-
cene, Knapp’s wells, Terrebonne Parish, No. 1 at 1600-2443
feet, No. 2, 1050-1842, No. 3, 58-1839 feet, New Orleans
ar.esian well of 1856, Lake Borgne borings. Pliocene, New
Orleans Gymnasium Club well at 1200 feet. Upper Mio-
cene, Galveston well, 2871 feet. Miocene, Gilbert well No.
10, Bateson, Texas at 323 feet.
consensus Ravenel, Pr. A. N.S. Phila., p. 43, 1861; Dall, Bull.
VEC Z iS DATS 5s: Sor
Distribution.—North Carolina to Florida. 8-49 fms. Gulf
coast 20 miles off Charlotte Harbor, W. Fla., at 13 fms.
hotessieri d’Orbigny, Moll. Cuba, 2, p. 142, 1845, Atlas, pl. 21,
f. 40-42, (as Vassa); Dall, Bull. M. C. Z., 18, p. 185, 1889;
Dally s) Simpson) Bull. U.S) Fish) Com) p. 402) 1901r:
Distribution.— Gulf of Mexico, Blake station 36, at 84 fms.
Also Hatteras and Sombrero, 30-85 fms.
galvestonensis Harris, Bull. Am. Pal., vol. 1, p. ror, pl. 4, f. 2,
1895.
Distribution.—Upper Miocene, Galveston well at 2410-2871
feet.
trigalvestusiilaris: bull) Am?! Pal. volume) py mony iplaia. sp:
3, 1895.
Distribution.—Upper Miocene, Galveston well at 2158-2871
feet.
obsoletus Say, Jour. A. N. S. Phila., (1) 2, p, 232, 1822, (as
Wassa) Gould, Inv: \Mass., pl. 15, f))210;), Dall) Bull: 37,
UA Se ONGEMe pr nO) ply So. f you mesonmoluson\ Oce! Pax
BOSts SOCHNE EE 74) Ds TA Shir Or ss Wal buble rere WSN).
Mis en Oss LOZ.
Distribution.—Nova Scotia to Tampa, West Fla. Also San
Francisco Bay, (introduced with oysters from the Atlantic).
61 BULLETIN 38 Or
beaumontensis Aldrich, Bull. No. 1, Univ. Texas, July, 1901;
Nautilus, p. 74 and figs., Nov. 1901, (as Wassa).
Distribution.—Pleistocene. Spindle Top Hill oil well, near
Beaumont, Texas, at 390 feet.
Genus STROMBINA Moerch
gibberuia var. galvestonensis Harris, Bull. Am. Pal,, vol. 1, p.
103; pli iauifi6. 1895.
Distribution.—Upper Miocene, Galveston well at 2410-2871
feet
Note—The typical form of gtbberula Sowerby, (Pr. Zool,
Soc., p. 115, 1832), is recent on the Pacific coast of Central
America southward to Paiti, Peru.
Genus ANACHIS H. and A. Adams
avara Say, Jour. A. N.S. Phila., 2, p. 230, 1882, (as Columbella) ;
uomey and Holmes) Pl. skosiS.4@ap. 139) iphi28. i 12
1357) Dall Bull. 37. Ui. Sa iNe wep sore) ple son ty mons o:
NManatta, Pr.) (ALUN. S20 Philaswssenpsi574 £6035, Oonnsons
OeciPat Bost, Soc.) Nye ipa 122M Toms)
Distribution.—Recent and Pleistocene. Massachussetts to
Florida Keys. (Gulf coast: Crooked Isl., West Fla., and
Texas. Pleistocene, Lake Borgne borings, La., and North
Creek, West Fla.
avara var. semiplicata Stearns, Pr. A. N. S. Phila., p. 344,
1873, (as Columbella); Tryon, Man., 5, p. 159, pl. 55, f. 70,
7iewos: Dall Bulls 275 Ue S.eNeME Moe nT Od 8o.
Distribution.—Cedar Keys and Charlotte Harbor, West Fla.;
Louisiana; Corpus Christi, Texas.
avara var. similis Ravenel, Pr. A. N. S. Phila., p. 41, 1861, (as
Columbella; ‘Tryon, Man., 5, p. 159, pl. 55, f. 69, ’83; Dall,
Bull 7h1 4) TOs); Sor
QZ MOLLUSCA OF THE GULF OF MEXICO 62
Distribution.—Carolinas to West Florida, Galveston and
Yucatan.
avara var. translirataRavenel, Pr. A. N.S. Phila., p. 42, 1861;
dinyon,Mani5, p. 159. 783; Wally Bulliaz7 pp.) 1L6,.)'so.
Distribution.—New York to West Florida, Louisiana and
Yucatan.
catenata Sowerby, Pr. Zool. Soc., p. 52, 1844; Conch. Icon., 11,
DMe2nety IO, 158") Deyouy Mani suis m7, a ipley 5S.) ss) Sis
i824) Dall and Simpson, Bull) UW. Ss) Pish\\Com:) 1, p. 404,
IQOl.
Distribution.—Jamaica to Porto Rico. A shell closely re-
sembling this species, and probably identical with it, is in
our collection. As far as I know, this is the first time that
catenata has been found on the Gulf coast.
obesa C. B. Adams, Pr. Bost. Soc. N. H., 2, p. 2, 1845, (as
Buccnum) iryon, Man, 5, p. HOON piniszh\tai7 (aud 20, (33.
Dalle uil s7 i p.ar18, 280: Singley,.q4th Anne) Rept. dex:
Cacompr 335.925 Evartis, Bulli Amy Pal ivoliam ip: O25, 0)5);
Wanatta=Pr. A. N. Si Phila., 55, p: 758, 1903:
ornmata Ravenel, cancellata Gaskoin. <A. ostretcola Melvell is a
dark brown or black variety.
Distribution.—Hatteras to St. Thomas, W. I. and Vera
Cruz. Pliocene to Recent. Gulf-coast: St. Joseph's Bay,
Crooked Isl., West Fla.; Point au Fer, Cameron and Chan-
deleurs, La.; Galveston and Corpus Christi, Texas. Pleisto-
cene, Grand Chenier, Teche No. 1 well, Jennings, La., at
1158-1199 feet, Knapp's well, Terrebonne Parish, No. 1 at
1050-1800 feet, No. 3 at 1150-1440 feet, Galveston well at
440-446 feet. Pliocene, New Orleans Gymnasium Club well
at 1200 feet.
amphisella Dall, Bull, M. C. Z., 9, p. 91, 1881; 18, p. 188, pl.
1OM Me LOG SOs Bull 2 72 0 S..N weep aero ple TOnpia LOG,
89.
Ore BULLETIN 38 03
Distribution.—Yucatan Strait 43—640 fms.
hotessieriana d’Orbigny, Moll. Cuba, 2, p. 138, pl. 21, f, 37-39;
‘Tryon, Man), 5) p.lrgasnpl sods 2unneses Dall «pitti aE
POLLS. OO:
Distribution.—Tampa, West Fla., to Guadeloupe, W. I.
Very closely allied to the following species.
pretri Duclos, Chenu, Conch: Ill., pl, 16, f. 7, 8; Tryon, Man.,
Rep A MAAN DL (52a ONTOS
Note.—C. B. Adams’ eldella [Conch. Contr., p. 63, 1850,
(as Pleurotoma); Dall Bull. U.S. N. M., p. 118, 1889] an
unfigured species, is 7#de Dall and Simpson (1901) merely a
mutation of prefrz, as shown by cotypes from Adams.
Distribution.—Variety albella, Cape Fear, N. C., to West
Florida and Jamaica.
Genus COLUMBELLA Lamarck
mercatoria Linnzeus, Syst. Nat. ed. X, p. 730, 1758, (as Voluta);
Kiener, Coq. Viv., Columbella, p. 23, pl. 5, f. 1, 1841; Dall,
Bull. 37. p. 116,89; Dall and Simpson, Bull: U.S. Fish
Comat, p4'403) 1901.
Distribution.—Cape Lookout, N. C., to Barbados and West
Florida. Pleistocene of Costa Rica.
rusticoides Heilprin, Tr. Wagner Inst. Sci., 1, p. 81, pl. 8, f. 9,
1887. Pr VAC ONS. Philaliip is 2mploia. f..20)' 87.) Dally ain
WET SM ia) ip. 125; oouNauohan i eubl. Nose. tC@artsae
nSty pl 7 Ti TOO;
Distribution.—Cedar Keys and St. Marks, West Florida to
Cuba. Pleistocene, Labelle, West Fla. Pliocene, Caloosa-
hatchie beds, West Fla.
Genus ASTYRIS H. and A. Adams
lunata Say, Jour. A. N.S. Phila., 5. p. 213, 1826, (as Massa )
Tuomey aud Holmes, Pl. Fos. S. C. p. 136, pl. 28, f. 6,
O4 MOLLUSCA OF THE GULF OF MEXICO 64
1857, (as Buccinum); Hilgard, House of Rep. Ex. Doc., 1,
PEN) Ps S8ou1s73> Walle Bull27) 5 Ue SAIN MES (ps 118, pl:
SOM TOME SOs OMnMsom. Occ: Pai BostanSocun Np bln) 7.0.
133, I915.
Bnecinum zonalis Linsley, Am. Jour. Sci., (1); 48, p. 285,
1845.
Distribution.—Canada to Florida Keys. 0-12 fms. Gulf
coast: Cedar Keys, Fla.; Chandeleurs, La.; Galveston and
Corpus Christi, Tex. Pleistocene. Lake Borgne borings,
La., Knapp’s wells, Terrebonne Parish, No. 2 at 1050-1790,
No. 3 at 258-1375 feet. Also of North Creek, Manatee and
Labelle, West Fla. Pliocene of the Caloosahatchie, Fla.
duclosiana d’Orbigny, Moll. Cuba, 2 p. 136, pl. 21, f. 31-33, °45.
(Not of Sowerby, 1847, which is a Javan shell); Dall, Bull
37, U.S.N.M.,p. 118, ’89, (as dunata var.); Dall and Simp-
son, Bull. U. S. Fish Com., I, p. 405, 1901.
Distribution.—Hatteras to Barbados and Tampa, West Fla.
0-63 fms.
diaphana Verrill, Tr. Conn. Acad., 5, p. 513, pl. 58, f. 2, 1882;
Dalla ME CHZ ITs p. TOL plenaisinewloun So; “bth 27°
Oe SHUN MES Da mrs ple 25. fi. Op SOE
Distribution.—Rhode Isl. to Gulf of Mexico between Mis-
sissippi delta and Cedar Keys, 64-487 fms.
Genus NiTIDELLA Swainson
parvula Dunker, Zeitschr, Mal., p, 64, 1847, (as Buccinum);
Mnyvony Man Si. p.022.\ pl. 48a te Sa Wall Dialling,
We Sa NG Mes pe 183) 30:
Distribution.—Gulf of Mexico, Texas region, to Barbados.
Genus AESOPUS Gould
stearnsi Tryon, Man. Conch., 5. p. 179, 1883, (as Semznella) ;
Dah pBulliMeyC.\ Zi 18) pi) 194. ply Zou tisn SOs mba 3 7,
PALLIS Hi le Lous 52 SO:
65 BULLETIN 38
a
ea]
jilosa Stearns, Pr. AUN S) Bhilai pais asics mig ime 7a4 (as
Nitidella). Not Aesopus tilosus Angas, 1867, cf. Columbella
peculiarvts Guppy, Geol. Mag., London, pl. 18, f. 20, 1874.
Distribution.—Type locality, Tampa Bay, Fla. Also off
Cape Fear, N. C. 15-17 fms. Pliocene of the Caloosahat-
chie, Fla.
Genus MUREX Linnzeus
beaui Fischer and Bernardi, Jour. de Conch., 5, p. 295, pl. 8, f.
TiTos5os Dall Bulls Mo CZ a mee wemnO sien Soi a llama,
We SING pS) 78O:
Distribution.—Gulf of Mexico between Mississippi delta and
Cedar Keys, at 111 fms. Also Florida Reefs and Guade-
loupe. 82-183 fms.
cabriti Bernardi, Jour. de Conch., 7, p. 301, pl. 10, f. 3, 1858;
Dall Biull 13700. 9S. NE UMES iospene m7 So.
Distribution.—Hatteras to Barbados, Gulf of Mexico, Blake
station 36, at 84 fms.
messorius Sowerby, Pr. Zool. Soc., p. 137, 1840; Conch. Illust.,
Murex, £. 93, 1839; Reeve, Conch. Icon., Murex, f. 90, 45;
Dalit W. Li S., 3; pwineo. 160. Dalland) Simpson) Bull
UMSirish Com, 1, \p:) 407) 19e1
Distribution.—Gulf coast, Cedar Keys, Fla. and Texas, to
Colon. 2-30 fms. Pleistocene, New Orleans pumping
station No. 7. Pliocene of the Caloosahatchie River, Fla.
(Chicoreus) rufus Lamarck, Anim. s. Vert., ed. I, 7, p. 162,1882;
Apyonuk) Man) 2, yp) oo) uplimmsnatent4s) 1.30.) (CAttenuNeceyculsy
Avo) Dally Bull 27, Up SaNe Vern Si SO:
sexcostata Emmons, Geol. N. Car., p. 248, f. 106, 1858.
Distribution.—Cape Fear, N. C., to Cartagena, Colombia,
and Antilles. Also West Fla. 5-30 fms. Pliocene Ca-
loosahatchie beds, Fla.
96 MoLLUSCA OF THE GULF OF MEXICO 66
(Chicoreus) brevifrons Lamarck, Anim. s. Vert., ed. I, 7, p. 161,
1822; Reeve, Conch. Icon., 3, pl. 3, f.13, 1845: Tryon, Man.,
PPOs tb 72) Toso; Dalle sBull 27. Ue Sa NEMO. Os. 01S,
"89; Tr. W. I.S., 3, p, 140,’90; Dall and Simpson, Bull. U.
Sadish Com: 15.p..407, LOL:
Distribution.—South Carolina to Cartagena, Colombia. Gulf
coast, West Fla. Pliocene, Caloosahatchie beds.
(Phyllonotus) pomum Gmelin, Syst. Nat., p. 3527, 1792; Reeve,
Gonchs eon: 3) pl. 9, f.-35; 1845: sbryons) Many-2.. p. 97,
piezo. f. 132, 80; Dall) Bull. 27, U2 Ss NaMe, pp 120, pl 16,
fee? e280" br. Wi. I. S., 3, p: 42), 007 Dalljand) Simpson;
Bull. U. S. Fish Com., I, p. 408, I90t.
oculatus Reeve, 1845; asperrimus d’Orbigny, 1853; mextcanus
Petit, 1852; zmperialts Swainson; globosa Kmmons, 1858.
Distribution.—North Carolina to Venezuela. Gulf coast at
various localities, West Fla.; Cameron and Chandeleurs,
La., and Texas. Pleistocene, New Orleans pumping station
No. 7. Pliocene, Caloosahatchie River, Fla. Lower Mio-
cene, Antilles.
(Phyllonotus) fulvescens Sowerby, Conch. Illust., f. 30; Kobelt,
Nalin Mal. (Gesell), p.155,. 77; Dalit Bullaa7. ULo5 Ne ME,
perzo7s8o7) Vanatta, Pr: A. NaS Phila 55 apa 75011903:
Distribution.—Hatteras and the Gulf coast at Indian Pass,
West Fla.; Point au Fer, Chandeleurs, La.; Galveston,
Corpus Christi and Quintana, Texas. Pleistocene, Grand
Chenier, La., Knapp’s wells, Terrebonne Parish, No. 1 at
2000-2150 feet, No. 3 at 790-830 feet.
Genus FAVARTIA Jousseaume
cellulosa Conrad, Pr. A. N.S. Phila., 3, p, 25, 1846, (as AZurex);
DalleBull Me Ce: Zo18, p. 210, pli monteuns Go (as Oczzclra):.
Bully.3 eae SNe Ms ps 120, pla Tomine SO ambi WEL noe,
Beant 50s 90:
nuceus Moerch, Cat. Kierulf,p. 14, pl. 1,f. 9, ’50; Cat. Yoldi, p.
95, 1852, (as Ocinebra).
67 BULLETIN 38 O7
Distribution.—Cape Lookout, N. C., to the Antilles and
Wera Cruz. Gulf coast, Tampa Bay (type locality), Cedar
Keys and Texas. Pliocene of the Caloosahatchie.
cellulosa var. levicula Dall, Bull. M. C. Z., 18, p, 211, 1889; Bull.
27, Py 120% (89. (as Ocweora):
Distribution.—Cape Lookout, N.C., to West Florida and
Yucatan. 22-37 fms.
intermedia C. B. Adams, Contr. Conch., p. 60, 1850, (as Murex);
Dall Bull) 27, U.S) NOM. pait2onSoii(as) Oczeb7a) Dally
and Simpsou, Bull. U. S. Fish Com., 1, p, 408, 1901.
Distribution.—Key West to St Thomas and Texas region of
the Gulf. Variety a/ta Dall, Caloosahatchie Pliocene.
Genus UROSALPINX Stimpson
cinereus Say, Jour. A. N.S. Phila., 2, p. 236, 1822, (as Fusus);
Amer./(Conch., 3, pl. 209, 1831; Gould, Inv. Mass:,p.'370)) £:
637, (as Buccinum); Tryon, Man., 2, p."152, pl. 39, f. 3,80;
Dall Pr iOS SiN MeO np eiseOumnsse btu 27500 an suiNE
M., p. 120, pl. 50, f. 6, ’89; Johnson, Occ. Pa. Bost. Soc. N.
AYipyire2) Tor5:) Dall. Bulleri2 Wen SsoNii Mi. p.TOo MN TO2.
Distribution.—Nova Scotia to Florida. Gulf coast, Cedar
Keys (Dall); Pleistocene, Lake Borgne borings, La., (Hil-
gard). Also on the Californian coast (introduced with
oysters).
perrugatus Conrad, Amer. Jour. Sci., p. 397, 1846, (as Fusus);
DallBall Mi Ce OZ isi py 2mwimson Bull) 27 pei 2ONWso.
Vaughan\2d' Ann. Rept. /HlanG.iS.))p. 148. 909.
Distribution.—West Florida from Cedar Keys to Key West.
Type locality, Manatee River, West Fla. Pleistocene,
North Creek and Orient, West Fla.
tampaensis Conrad, Pr. A.N.S. Phila., 3, p. 25, 1846, (asMurex);
Dall Bull. Me iC. Z. 18) pw 214 8on sBull37,) Ui) Si Noe,
DIE 2 SO;
g8 MOLLUSCA OF THE GuLF OF MEXICO 68
Distribution.—West Florida from Cedar Keys to Sarasota.
Pleistocene, North Creek, West Fla.
Genus MURICIDEA Swainson
multangula Philippi, Zeitschr. fur Malak, 5, p. 25, 1849, (as
uss) eX pid: ind: Beschr) (33) pel Lay plan 244i.) O18 50%
DAB aN a7 UL SON Mi pent 20) Olan 7ONton tense Oats | Ve
SMa E49) T890%)) Pry Ua Oo INN 2AR DE SO5 i Bl) 20)
ite) 1G) CKO.
hegeprele Dall. Pro Us SiN .\M.) 6; pelza7i 1833)
Distribution.—Cape Fear, N. C., to Yucatan and the North-
ern Antilles, Gulf coast, Cedar Keys and Texas. o-g5 fms.
Pliocene, Caloosahatchie beds.
ostrearum Conrad, Pr. A. N.S. Phila., 3, p. 25, 1846, (asAZurex);
Dalley Oi SNe ME. 240.) 505.0 pla goin Z LOO.
Muyjomdana Conrad, Am, jour. Conch.) )5)) pi 206, (pl. 12, £2145
T8oo9u (as) Crosalpznx):\Dall, Pr. Us S MNAME ow p. 326) 183.
lle CZ TS oni 20 Bon ra Wilh Sait DW AON)) GO}
Vaughan, 2d Ann. Rept. Fla. G. S., p. 148, 1909.
Distribution.—East Florida south of St. Augustine and Gulf
of Mexico, Cedar Keys, Fla., to Yucatan. 0-13 fms. Rocky
shores. Pleistocene, Manatee and Orient, West Fla. Plio-
cene, Caloosahatchie beds, Fla. Resembles Uvosalpinx
perrugatus Conrad, but is slenderer and has a very different
operculum.
hexagona Lamarck, Anim. s. Vert., 7, p. 169, 1822, (as Murex);
Reever Conch Micont)(2.)\pl4) 27 tuiTzonmaqs Mable iil 27.
We So Ne OM. p20, 789; Dall & Simpsonm/Bully U0. SiFish
Com., I, p. 409, I9OI.
Distribution.—Gulf of Mexico, Texas region to St. Thomas,
W.1I1.25 fms. Genotype.
philippiana Dall, Bull. M. C. Z., 18, p. 213, 1889; Bull. 37, U. S.
ING OM ape 20% SOs) ert. Ole.) UNE) Meh 2ani mn S@unnmle Doi fic)
1902.
69 BULLETIN 38 99
Distribution.—Off Cape Catoche, Yucatan, and also Key
West, Fla. 20-25 fms. Coral sand.
Genus EUPLEURA H. and A. Adams
caudata Say, Jour. A. N.S. Phila., 2, p. 236, 1822, (as Ranella);
Stimpson, Am: Jour!) Conch. ai.) 158.) ple 8s te Sun oO:
iilsard, Mouse of Rep. Ex. Docr 1, pt-2) sp..886. 47.33) Dalle
BU VS) AN Mi Oli pass 2On Hn San sbi tac mila SemiNen MeN moe
TZOM Pl.) 50 te LT, SOx} Ounson) Oces Par aBost Soc iN Eley
Fey, (Oui KO EO) Hse
Distribution.—Massachussetts to Florida. 1-3 fms. Gulf
coast: Cedar Keys, St. Mark’s, Charlotte Harbor, Fla.;
Chandeleurs, La. Pleistocene, Lake Borgne borings and
New Orleans well of 1856 (Hilgard), and of Labelle, West
Fla.
caudata var. sulcidentata Dall, Tr. W. I. S., 3, p. 144, 1890.
Ranella clathrata ‘‘Gray’’ Calkins, Pr. Dav. Acad. Sci., p.
234) 1878.
muriciyormis Viyon, | Man.) 2.) palnss.. (nm part) Noor
Broderip.
Distribution.—Gulf coast of Florida, Pleistocene of South-
west Fla.
Genus TROPHON Montfort
(Boreotrophon) lacunellus Dall, Bull. M. C. Z., 18, p. 205, pl. 15,
iq rsso; }bull).37.) Ue Sa Ne Meeps c 20. (poll 1 Satis Or
Distribution.—Cape Fear, N. C., to Dominica. Also West
Florida. 227-769 fms.
Genus THAIS Bolten
patula Linnzeus, Syst. Nat., ed. X, p. 739, 1758, (as Buccinum) ;
Reeve, Conch. Icon., 3, pl. 1, f. 3, 1846; Tryon, Man., 2, p.
150, Pls) £4) 19-22) 807m all nB alle 7 Amis. Ne vines
OO
MOLLUSCA OF THE GULF OF MEXICO 70
M22 SOM (AS A UUpuUnd) wall Pr Wa Sum Ne vie 3751s 221,
IQIO.
Distribution.—East and West Florida to Texas, the An-
tilles and Brazil. Also Pacific, Gulf of California to Pana-
ma and Peru.
haemastoma Linnzus Syst. Nat., ed. XII, p. 1202; Reeve,
Conehyicon., 3, f. 21, 1846; Tryon, Man.\)2) :p.)167,) pl. 40,
Paso, 83 1880; Dall, /Bull.)327 0 U2ssy Ne Mes p22 .\pliaa:
is. 4, pl. 46, Ta-2b, 89°) Singley ‘athyAnn. Rept, dex.) G:
Se pessow oor WVanatta,) Pir) AVENE Sei Phila sic pag 5o,
1903. (All as Purpura).
Distribution.—Cape Hatteras to Trinidad Isl. Gulf coast
Cedar Keys and Ft. Barranca, Fla.; Horn Isl., Miss.; Point
au Fer, La.; Galveston, Corpus Christi, Tex. Pleistocene,
Grand Chenier, La., New Orleans pumping station No. 7,
Knapp’s No. 3 well, Terrebonne Parish, at 1400-1440 feet.
haemastoma var. floridana Conrad, Jour. A. N.S. Phila., 7, p.
Z2OceapnZOwta 2h. Iker ry On, Miadei2 epsnno7a splerd@u ft:
85, pl. 50, f. 86; Dall and Simpson, Bull. U. S. Fish Com.,
InP: Gul, Loon.) CAllias Purpura):
Distribution.— Florida to the Antilles, Gulf coast, West Fla.,
Cameron, Chandeleurs, La.; Matagorda Bay, Tex. Pleisto-
cene of Grand Chenier, La., (very abundant and showing
the same mutations as the living shells).
deltoidea Lamarck, Anim. s. Vert., 7, p. 247, 1822; Reeve,Conch.
liconsusn plan fis, 1S4o> Muster mC onchaCabet pla 2.) fh
De Mirror, MEL Boy mda gollan Mya is Us. lacctsroe! IDEN Is i000 |.
BE OM ol UNE) Men pe 122) So. Dally andtSimpsony Bully U):
SwhishiComyyis py 411, Toor. | CAlWasw aarp na.
Distribution.—Florida to St. Vincent, W. I., Gulf coast,
West Fla. and Tex.
Has BULLETIN 38 LOL
Genus SISTRUM Montfort
nodulosum C. B. Adams, Pr. Bost. Soc. N. H., p. 2, 1845, (as
Purpura) rvons Nani (2) pi TOOn Dien 5 One n27 Saelooor
Dall and Simpson, Bull. U. S. Fish Com., 1, p. 411, Igor.
Distribution.—Cape Romano, southwest Florida, to Colon.
Also Antillean.
roseum Reeve, Conch. Icon., sp. 46, 1846; Dall, Bull. 37, U. 5.
IN VMs p.)/ 122.) 1830:
Distribution.—Gulf of Mexico, Texas region, to Barbados.
The rosy color is thought to be caused by an encrusting hy-
drozoon.
Genus CORALLIOPH!{LA H. and A. Adams
abbreviata Lamarck, Enc. Me’th., pl. 435, f. 2a-b, 1795, (as
Haynie) AX mans.) Vientuni emule on ene 22)
galea Chemnitz.
Distribution.—Cape Fear, N. C., to the Antilles. Gulf
coast, West Florida. Pliocene, Caloosahatchie beds.
lactuca\; Dall, | Bull. M. iC. Z.\18))p1)220, pl! 16,/f. 6, 1880:
Distribution.—Gulf of Mexico, Blake station 5, in 152-229
fms. Also off Fernandina, Fla.
deburghiae Reeve, Pr. Zool. Soc., p. 208, pl. 38, f. 3, 1857, (as
Riizochilus)\:\ Mnyon, Mane 2iinla Gon) fs) 283.) TS soOuncas
Watzax7s)i2)\ Dall, Bull WA SNe Men p22. pla TONE Li siiso:
Distribution.—Cape Hatteras to Barbados. Also West Fla.
56-878 fms.
Genus EPITOMIUM Bolten
angulatum Say, Amer. Conch., No. 3, pl. 27, (two upper figs)
1831, (as Scalaria clathrus var.); Hilgard, House of Rep.
Bx DOC. WT)" Pts) 2) (Passive cso ne oO allen Dall 27. Oks kINe
M:)) (py) 122,89; Singleya Ath Pano iRept Mex CaiSii pe
236,792; Johnson) \\Occ mea Bosts SOC4 UN). al...) 7.npIntOs"
IQI5.
LOZ MOLLUSCA OF THE GULF OF MEXICO 72
Distribution. —Connecticut to West Fla., Point au Fer., La.,
and Galveston, Tex. Pleistocene, Lake Borgne, La., bor-
ings.
sayanum Dall, Bull. M.C. Z., 18, p. 313, 1889, (as Scala); Bull.
SPO SUN VEN Dana pl.) SO,) Mutou Sonmebt WN lS. 3,
Piss) 90; Singley, 4th) Ann.) Repty Tex. G.'S.))p. 336,
1892.
cdathrum Say (in part).
Distribution.— Virginia to Florida Keys, West Florida, Cam-
eron, La., Corpus Christi and Galveston, Texas. Pleisto-
cene, Grand Chenier, La. Pliocene, Caloosahatchie River,
Florida.
tenue Sowerby, Pr. Zool. Soc., p. 110, 1844, (as Scala); Dall,
Bulle a 7e Wi SiN iM. pi 522) t8so:
Distribution.—Gulf of Mexico, Texas and Cameron (?) La.,
tole sLnomas, WT.
nitidellum Dall, Bull. M. C. Z., 18, p. 314, 1889, (as Scala); Bull.
TORRONE: Wen pink). 8: Er O en oamiNn Mest 245) 6 SO,
DI BOK ta.8.) 1902:
Distribution.—Hatteras to Cedar Keys, Fla. 32-63 fms.
Seinen Wall Bull..May/C./Z.) 18) p. 210 n88q,n bull 375) Uso. N
NRT PY 12407 BOs) (aS Sala) 5) (REAM E ROU MING IME N24: Ds N5O6)
PLE ZOR TOM TOO2:
Distribution.—Hatteras, N. C., to Texas and Vera Cruz,
Mexico. Io-16 fms. Pink with white varices.
denticulatum Sowerby, Thes. Conch., p. 87, pl. 32, f. 25, 26, °47;
miyon Mani. Os pls) 13. f. 100, (e875) DallyyBuhlhy i237 (Us) Ss
INE py n2qn Soe
Distribution.—Hatteras to West Florida and the Bahamas.
lineatum Say, Jour. A. N. S, Phila., 2, p. 242, 1822, (as Sca-
laria); Hilgard, House of Rep. Ex. Doc. 1, pt. 2, p. 887, °78;
Mnvonw Mani Ope 79. ply 16, cf kmorisnu Oa lair Nis be se.
73 BULLETIN 38 103
20 pe 5S, QO; JohnsonaiOce wha OSes SOC. IN auelna aap:
HOPE Akay els
Distribution.—Massachussetts to Charlotte Harbor and
Cedar Keys, West Fla.; Cameron, Point au Fer, La.; Gal-
veston and Corpus Christi, Tex. Pleistocene. Grand
Chenier, La., and Knapp’s No. 2 well Terrebonne Parish,
at 1550-1570, 1731-1739 feet. Pliocene Caloosahatchie
beds, Fla.
turriculum Sowerby, Thes. Conch., 1, p. 92, pl. 33, f. 61, pl. 34,
f. 88, 1847; Singley, 4th Ann. Rept. Tex. G. 5S., p. 336,
1892.
Distribution.—Cape Hatteras to Haiti. 16-32 fms. Gulf
coast at Galveston (Singley). Pliocene of the Caloosahatchie
River, Fla.
mitchelli Dall, Nautilus, 9, p. 112, Feb. 1896; Pr. U.S. N. M.,
24, p. 506, pl. 30, f. 3, 4, 1902.
Distribution.—Matagorda Island, Texas. Collected by J. D.
Mitchell.
cochlea Sowerby, Thes. Conch., Scalaria, p. 103, pl. 35, f. 142
(only) 1847;Moerch, Jour. A. N. S. Phila., 8, p. 205, 1876;
Mayon Man), (9. 81, :pln7 ink 2400 8o7 Walls PEW. Saunt
MONZA D500. ply; 30, taj, oo2r
Distribution.—Hatteras, N. C., to Tortola. W.I. Gulf of
Mexico near Cedar Keys, Fla., at 25 fms. Sowerby’s refer-
ence to its occurrence at Loando, Portuguese West Africa,
is questionable.
galvestonense Harris, Bull. Am. Pal., vol. 1, p. TO3) Wplena iter e7pe
1895.
Distribution.—Upper Miocene, Galveston well at 2552-2871
feet.
Note.—The following species of Epztomium have been re-
ported by Singley from the Texas coast:—sericifila Dall,
104
MOLLUSCA OF THE GULF OF MEXICO 74
(Bull. M. C. Z., 18, p. 313, 1889); multistviatum Say (Jour.
ONES ve hilay nse pe 20S" 1520 DallveBulliisr We SauN. Mie.
pl. 50, f. 5, 1889) ;contorquatum Dall, (Bull. M. C. Z., 18, p.
318, pl. 18, f. 9, 1889); and modestum C. B. Adams (Jamaica
Shells). These species are not cited, as far as I am aware,
by other authors from Gulf coast. They are not in our col-
lections from the Gulf and are noted here solely fide Singley,
(Fourth Ann. Rept. Tex. Geol. Surv., p. 336, 1892).
Genus JANTHINA Bolten
janthina Linneeus, Syst. Nat., ed. X., p. 772, 1758, (Helix);
d.Oxrbigny, Voy. Am: Me’r); p.1413,\ pl: 61, f) 8-10), 13841;
Dal Wer Wyss Ne M327. p. 2220 neo sn) olnson. Occ. Pal
IBOSta SOCNE Een 7.) p. TOT Tor 5:
fragilis Lamarck, An. s. Vert., p. 89, 1801; Tryon, Man., 9,
p. 36, pl. 9, f. 4, 1887; Singley, 4th Ann. Rept. G. S. Tex-
ASD son 1892. bartsch, Bull. orgwUaySe Ney Mie. ‘py (65;
IQI5.
Distribution.—Nantucket, Mass., to Colon; and West Fla.
to Quintana and Galveston, Texas. Pelagic. Also Pacific
Ocean, Lat.42°N.to 36° S.,and Indian Ocean, Port Alfred, 8.
E. Africa.
pallida Harvey, Thompson’s Ann. Phil. and Nat. Hist., 5, p. 96,
Dieta Sat nbyon.. Man. Opec ep lmTOntN tS) .T S870
Distribution.—Europe, the West Indiesand Gulf of Mexico,
Cameron, La. (washed ashore). Also tropical waters of the
Pacific Ocean. Pelagic,
globosa Swainson, Zool. Illust. (1), 11, pl. 85, 1822; Tryon,
Mano) ple ton tb2,)87-) singley Athy Ann Rept mMexa) p.
336, ’92; Mitchell, List Texas Shells; Bartsch, Bull. 91,
Se Ne Me pos sro T's:
Distribution.— Gulf Stream, Pelagic. Washed ashore at
Quintana, Tex. and West Fla.Also Port Alfred, S.E. Africa.
communis Lamarck, An. s. Vert., ed, Desh., IX, p. 4; Tryon,
BULLETIN 38 Io5
=
Cr
Manitou plio, ti ooumesre Dall Butiia 7.) MUL S.. iNet VME os
126, 80; Mitchell, List. Tex. Sh.; Bartsch, Bull. 91, U.S.
INS pL 05.) LOLS:
Distribution.—Nantucket to Coion, West Florida, Matagorda
Bay and Mustang Isl., Tex. Pelagic. Also Indian Ocean,
Port Alfred, S... Africa.
Genus MELANELLA Bowdich
(Eulima) conoidea Kurtz and Stimpson, Pr. Bost. Soc., N. H., 4,
Pulses all wr WSs IN IM Nowe 22ONTsS2.) Miao
Mani Sipe 27i2ipl. (40 Nao oo moally Mira Vieni 2. hpe
159, pl. 5, f. 11, 1890; Singley, 4th Ann. Rept. Tex., p. 336,
192: )OonsSon, Occ.'Pa: Bost: sSocwiNetrts 7.) pol TO 15.
Distribution.—Hatteras to the Autilles. Gulf coast: Cedar
Keys, Fla.; Galveston and Corpus Christi, Tex. Pliocene,
Caloosahatchie beds, Fla.
(Eulima) gracilis C. B. Adams,Contr. Conch., p. 110, 1850; Dall,
Balla OL SiN Mo) pitE26. 780%
Distribution.—Hatteras to St. Thomas, W. I., West Fla.
and Yucatan. Pliocene, of the Caloosahatchie.
(Eulima) intermedia Cantraine, Diagn, Moll., Bull. Ac. Brux.,
Pata HL8353) dnvon, Mamiisinmz74, ple hoo, £43.80; Dally
Bully 270 Ul) S-Ni MoS puoi 28h ia) So. i
Distribution.—England to the Canary Isls., New England
to West Florida and Barbados.
(Eulima) jamaicensis C. B. Adams, Pr. Bost. Soc., N. H., 2, p.
Gi ns453/\Dryon (Mans ipii2 73s pli Go. tS vain) i807) Dalit
Balle Me Cy IZ i) raMipal 227i oSo!
Distribution.—Gulf of Mexico, Blake station 20, in 220 fms.
off West Fla. Also Barbados.
(Eulima) subcarinata d’Orbigny, Moll. Cuba, p. 217, pl. 16, f. 4-
6. Dryon Man.) 18) \p.) 273. 4plst6o) t447)/1886;) Dall) Bulltizin:
ONS SNM ip IziG ear
106 MOLLUSCA OF THE GULF OF MEXICO 76
Distribution.— West Florida to Haiti.
(Eulima) gibba De Folin, Les Me’le’agrines, p. 64, pl. 6, f. 4,’67;
sbi OnwViane nS pv27 20 plow tiacwe 7 ballyn Bulla 7)
Ws Sa INIA WEY Fo AIS Key
Distribution.—Cape Hatteras to Campeche.
(Liostraca) bilineata Alder, Moll. Northumberland and Durham;
sirevonti Man .8) p.i279, (oli ZO. Wge-7a Tees Dal Ball,
BRD E 20%) SQ.
Distribution.—Norway to Mediterranean, and West Florida
to Haiti.
(Liostraca) hemphilli Dall, Pr. U. S. N. M., 6, p. 330, pl. to, f.
Perso BU 3 7S. IN: | Mapai 2onplcaSs of. Tn. SO.
Distribution.—Cedar Keys to Marco, West Fla.
(Liostraca) fusus Dall, Bull.M. C. Z., 18, p. 329, pl. 19, f. 11b,
SOR IBiCldls Aq NOheioy | ONO Cais ope caLS)y sFelliadeyy seyiieueloye Meso)
Distribution.—Fernandina, East Fla., to St. Kitts, W. I.
Gulf of Mexico, Yucatan Strait, 640 fms. Also off Hava-
na, 400 fms.
Genus NiISO Risso
splendidula Sowerby, Pr. Zool. Soc., p. 6, 1834, (Zudimia); A.
Indams) viness(Conch 7/250) Pp. Som NOMAzanplamt7Os mS) 2 54%
siyon Man iS. Pal2o7) Dl. 7 Le emoOs a ball Bill i2i7)
We SeeN yee ose 80) SO: Pr. MU EEO sN EN aie Tila N 2215
IQIO.
Distribution.—Cape Fear, N. C. to Grenada, and Gulf of
Mexico, between Mississippi delta and Cedar Keys in 111
fms. Also Pacific Ocean, Sta. Elena, Guayaquil Bay, Ecua-
dor (Type locality), in 6-8 fms. (Cuming).
aeglees Bush, Rept. U. S. Fish Com., p. 83, 1883-'85; Dall, Bull.
BOM Os NG Me plas si ple var fai TOa NS On CaS iZalenrupra
Sowerby var. ).
aa BULLETIN 38 lo7
Distribution.—Hatteras to Tampa, West Fla., 7-32 fms.
Pleistocene, Knapp’s No. 3 well, Terrebonne Parish, La.,
at 1150-1200 feet.
Genus PYRAMIDELLA Lamarck
delabrata Linnzeus, Gmelin, Syst. Nat. p. 3585, No. 113, (as
Trochus): (Lryon, ) Maney8\ ps (200 pl agen i772 alll
Bull 276 WSs IN Me oan 2. 8 eS or
Distribution.—Sarasota, West Fla. to Barbados.
(Longchaeus) crenulata Holmes, Post-Pl. Fos. S. C., p. 88, pl.
13, tw Alaa, 850!) (asiOve/esers) ew allie vill NI) Ce Zanersy
10 AL eSKstOe Ue AWE SL io. arin ome Wewoigeyy exe, Ja.
Ne SA ehila sp. 75S. loose
Obeliscus tesellatus Dall, Pr. U.S. N. M., 6, p. 330, 1883. Not
of Adams, 1855. For full synonymy see Dall, 1889.
Distribution.—South Carolina to St. Thomas, W. I. Gulf
coast, Cedar Keys, Tampa Bay. Fla., and Galveston, Texas.
Pleistocene, North Creek, West Fla., and of the Lake
Borgne borings, La. Habitat, grassy or muddy flats to 2
fms.
(Longcaaeus) candida (Meuschen) Moerch, Mal. Blatt., 22, p. 158,
(as Obelescus)):) Dall,” Biullaie7eOe. sae Ne OMe pil t2 Sens oor
ie Wei Sa 3 py 247, neece
Distribution.—Hatteras to Barbados and West Florida.
Pleistocene of Texas.
(Syrnola) smithi Verrill, Pr. U.S. N. M., 3, p. 380, 1881,(as Zur-
bonella)-) hr Conn: Acadasuap isco. pl 58, yf. 1 emuss:
Bartschs Pr.) Bost. (Soc: Ne WEleneae ps) gil. | pl. 1. Lenni:
1909; Johnson, (Occ!) Ra Bost) soc Nive, 7. 92,0) 15 NOt
of Tryon.
Eulimella unifasciata Dall (pars), Bull. M. C. Z., 18, p. 338,
PLMO, ft.) 11) 1889) Bulle enw See NLM spt 20. p leno yte
IIc, 89. Not of Forbes (Rept. Aegean Inv., p. 188, 1843)
which is a European shell.
x08 MOLLUSCA OF THE GULF OF MExICco 78
Distribution.— Massachussetts to Barbados and West Fla.
(Syrnoia) thelma Dall, Pr. U. S.N. M., 46, p. 254, pl. 20, f. 3,
IQi4.
Distribution.—Upper Miocene or Pliocene. Well near Al-
exandria, La., at 49 feet.
Genus TURBCGNILLA Risso
exilis C. B. Adams, Contr. to Conch., p. 74, 1850, (as Chemntt-
Ea ee Dalal 3700 Ss NG Maia 28h Soro. Wi. Meio.
Bee 250, 02:
Distribution.—Hatteras to Haiti and West Florida, 3-63 fms.
Pliocene of Caloosahatchie and Shell Creek, Fla.
belotheca Dall, Bull. M. C. Z., 18, p. 335, pl. 26, f. 7d, ’80; Bull.
CHAO OE NE OMee pr 28%) ol. 26) tad SOs
Distribution.—Gulf of Mexico, Blake station 32, at 95 fms.,
west of Florida, 50fms. Also Barbados, i100 fms.
interrupta Totten, Am. Jour. Sci., (1), 28, p. 352, f. 7, 1855, (as
engoicla Dalles) 27): VU: Sy INGiMeen py in28, pl.) 26h
Za Dessowm lr WWE le S43. pl 25oumo2umBanusch bia) Bost:
SOCwN A EEN 24h ps7 pl. 12) ft ToM2etTrooo +) johnson, Occ:
ae OSt 7 SOCe NN Ble 759 .) O15) OMG
wmaana Mal Pr. 10S: Ni: M), 62-0 i232 1882)
Distribution.—Nova Scotia to Barbados. 2-107 fms. Gulf
coast: Cedar Keys, Fla.; Galveston and Corpus Christi, Tex.
Pleistocene, Lake Borgne borings, La., Knapp’s wells, Ter-
rebonne Parish, No. 2 at 1050-1790; No. 3 at 1150-1839 feet,
also of North Creek, West Fila. Pliocene of the Caloosahat-
chie beds.
textilis Kurtz, Cat. Rec. Mar. Shells, N. and S. Car, p. 8, ’60,
(ash@hemnzizza)\ Dall, ir. We) IeSeet pa 257" 1S O2-
Distribution.—Charleston, S. C., to Marco, West Fla.
Pliocene, Caloosahatchie beds.
79 BULLETIN 38 109
obeliscus C. B. Adams, Contr. Conch., p. 72, 1850, (as Chemnit-
Ze) NOM M SHUN eva IOL a SWINE AVIS op iqeken staan Bren INI A Ile Sic 4
BNO ZOTe Oa"
Distribution.—Hatteras to Jamaica. 12-63 fms. Gulf coast,
West Fla. and Galveston, Tex. Pliocene, Caloosahatchie
beds, Fla.
Virga; Dall; Pry Ue SiN. MAG Osipsz2Q ns Sar iBall 27. Unis sine
Mie OuI2 5) 7 8o.
Distribution.—Hatteras to Florida Keys and Cedar Keys,
Fla. 2-15 fms. Pliocene, Caloosahatchie River.
punicea Dall iPr, SiN aM IG ips agioms8arlin.) Wi lene:
Bean Zon ESO2h
Distribution.—North Carolina to the Bahamas. 2-31 fms.
Gulf coast, Cedar Keys, West Fla. Pliocene, Caloosa-
hatchie beds.
curta Dall, Bull, M. C. Z., 18, p, 337, pl. 26, f. 7c, 1889; Bull.
WET SONG Mp. 28. pl 26h ies,
Distribution.—Yucatan Strait, 640 fms. Also off Cape Hat-
teras and Antilles.
speira Ravenel, Pr. Elliott Soc. Nat. Hist, 1, p. 280, 1859, (as
Chemnitzia) Tolmes:) Post kiiHossnou ©. p./82, pliant
LAWLOsSor Mal k ar. WW ise wDe25S.) TOO2.
Distribution.—Pleistocene, S. Carolina, and “de Hilgard, of
the Lake Borgne, borings, La. Pliocene, Coloosahatchie
beds (?). Not reported in the recent fauna.
reticulata C. B. Adams, Contr. Conch., p. 75, 1850; Dall, Tr.
WE Seip: 2OOn 1O2.
Distribution.—North Carolina to Jamaica, o-60 fms. Re-
ported by Singley from Galveston, Tex. Pliocene, Caloosa-
hatchie beds, West Florida.
conradi Bush, Vanatta, Pr. A. N. S. Phila., 55, p. 758, 1903.
{IO MOLLUSCA OF THE GULF OF MEXICO 80
Distribution.—Cited by Vanatta from Calhoun Co., North-
west Florida.
(Parthenia) cedrosa Dall, Pr. U.S. N. M., 6, 1883; Bull. 37,
WES NGM Dai ZO pl Asi fa) 4m Sao.
Distribution.—Cedar Keys, West Fla.
(Careliopsis?) styliformis Moerch, Malak,Blatt., 22, p. 169, 1874,
DAS we MN COZ.) 18. pi i338.) TSsoq Bulle aan ss MIN:
Wile. Oe IeeXON \eloy.
Distribution.—Hatteras to Sarasota Bay, West Fla. Also
St. Thomas, W.I. 2-52 fms.
Genus ODOSTOM!IA Fleming
acutidens Dall, Pr. U.S. Nat. Mus., 9, p. 331, 1883.
Distribution.—Hatteras to Cedar Keys, West Florida, 2-107
fms.
Note.—In 1892 (Tr. W. I. S., 3, p. 250), Dall regarded this
shell as.a variety of the European Miocene to Recent species,
O. conoidea Brocchi (Conch. Foss. Subapp.,2, p. 659,pl. 16,f.
2, 1814). ‘The latter species is recorded by Harris from the
Pleistocene of the Galveston well at 440-458 feet.
impressa Say, Jour. A. N. S. Phila., 2,p. 244, 1822; Dall, Pr.
WeeSUNGEMe.) 6. pH 2am! T882) Bulbas MWe Sa Ni iMEn D130)
PeNS2 it bso Sineley, 4th pAnnivRepty lex.) Gi Si)p.
BOT RO2 i banrtsch ahr Bost. SOCAN nen azn Ml 20it oS
LOO: oOnnsSonh Occ! Pay Bost. SocwNe Mee Te Dp osn LOLs:
Distribution.—Massachussetts to Florida. Gulf coast:
Tampa and Cedar Keys, Fla., Corpus Christi, Tex. Plio-
cene, Caloosahatchie beds, Fla.
seminuda C. B. Adams, Jour. Bost. Soc. N. H., 2, p. 280, pl. 4,
Tana S Zo (asi /azzenza) > DallsBuallai aa Wo SsuNi) Mes p
MZOM pS 25) £410,994) Bartsch, Pry Bost Soe. Ni ME 24.\p
T}
Leon}
BULLETIN 38 Tit
97, pl. 13, f. 45, 48, 1909; Johnson, Occ. Pa. Bost. Soc. N.
H., 7, p. 97, 1915.
Distribution.—Massachussetts to Florida Keys and West
Florida.
Genus PERISTICHIA Dali
toreta Dall, Bull. M: C. Z., 18, p. 340, 1884- Bull. 27, U.. SY Ne
Mei piso; ply 42 hOnsa)
Distribution.—Cape Lookout, N. C., to Key West and
Charlotte Harbor, West Fla. 2-22 fms.
agnia) Dall) Bull. M. (C. Z.)\-18) op avowrsso- Bulli27, U.S, Ne
MEN pst ZON Bo.
Distribution.—Marco, West Fla. o-2 fms. Also Key West
and off Hatteras.
Genus ATLANTA Lesueur
peroni Lesueur, Jour. de Physique, 85, p. 390, pl. 2, f. I, 2,
isi7 Gd ,Onbisny,) Voy: Amey Metis pr) 171, Pl.) t2.ite reise
nSZon Wall “Bally 27 (Us OMeNenIMe api zoOn plana a maaaian
Ple166. it) MItoa, 1889; (RrAOM Sanwa 637.) 1. U2 2 54uno nor
Buller 1 2, Wi S. Nev pags toon.
Distribution.—West Atlantic, Lat. 42° N. to Tropics. Gulf
of Mexico, West Fla. and Texas. Also eastern Pacific.
Pelagic.
Genus OXYGYRUS Benson
keraudreni Lesueur, Jour. de Physique, 85, p. 391, pl. 2, 1817;
H. and A. Adams, Genera Rec. Moll., 2, p: 92, 1858, 3, pl.
Gout, 6. 6b: Vern) \Transy Conny yAcad.) (6, "p14 20) 18 85¢
Dally Bull: 27. WU. Ss NM ep raoucso. johnson.) Occysrar
Bost Soc uN: Eas 7. pu argoxmmoms:
Distribution.—West Atlantic, Lat. 39° N.to Tropics. Gulf
of Mexico, West Florida and Texas. Pelagic.
X12 MOLLUSCA OF THE GULF OF MEXICO 82
Genus SIMNIA Risso
(Neosimnia) intermedia Sowerby, Spec. Conch., No. 1, p. 9, f. 32,
33, (as Ovulum), Thes. Conch., Ovulum, No. 40, p. 479, pl.
HOO HM. O02, 1848") Reeve, Couch Neon Ovrlane pl. 7.)t.
Bet OWensoS lity on Man. 57) 7))i py \25Iq\ oleae nee S5-S0 as
Ovnla); Dall, Bull. M. C. Z., 18, p. 235, 1889; Bull. 37, U.
SePNee Me ps 134), '8o.
Distribution.—Hatteras to Brazil. Gulf coast, Vera Cruz.
15-170 fms.
(Neosimnia) uniplicata Sowerby, Thes. Conch., p. 478, pl. roo, f.
30-32, 1848,(as Ovulum); Reeve, Conch. Icon., Ovulum, pl.
OMA Loos dinvon,) Mani hon 2isah polls.) cb. ao) Ao,
coo WalieBulleiz7 py 134, 89: Bullemra Oe SN. Meo:
LZOm OT.
Ovulum subrostratum Sowerby, 1848, antillarum and arcua-
tum Reeve, 1865. carolinensts Moerch, 1877.
Amphiperas canadiensts Moerch, 1882 (typ. err. for carolinen-
SAO) e
Distribution.—Hatteras to Barbados and West Florida. 12-
121 fms. Also Pacific coast, San Pedro, Cal. to Lower Cal-
ifornia.
Genus CYPHOMA Bolten
gibbosa Linnzeus, Syst. Nat., ed. X, p. 726 1758, (as Bulla);
Incevies Conch!) eons, )15, pl.) 7.) fangeni sos. (as Ovzz/e72):-
igor Many 7) 01 250, ple 3, t-Sl, S2se5en Dally Bulla 27,
U. S.N. M.,p. 134, ’89; Dall and Simpson, Bull. U. S. Fish
Com., I, p. 420,1901, (as Ulizmus).
Distribution.—Georgia to Trinidad, 15-50 fms. Also West
Florida.
83 BULLETIN 38 Itz
Genus PEDICULARIA Swainson
decussata Gould, Pr. Bost. Soc., N. H., 5, p. 127, 1855; Dall,
Bolle Me Cy 2008. p 237. h plo vite Oa pus Soruis tt llaiai7.
We SaN MIS pe ra. pling. tabs!
alozda) Dall Bully MOC) Z. on piso mos:
Distribution.—Georgia to Barbados, and Yucatan Strait
640 fms.
¥
Genus CYPRAEA Linnzeus
spurca Linnzeus, Syst, Nat., ed. X, p. 724, 1758; Reeve, Conch.
liconi 2) splot,” £68. 18415 :) dnyonee Manin 9. .0O'5 1) ple mon
BELOW T 7.) T9e5>, Dall Bulle 27 Ura aiNe MeN D136) 780)
Distribution. —Cedar Keys, West Fla., to Barbados, 0-25
fms.
exanthema Linnzus Syst. Nat., ed. XII, p. 1172, 1767; Reeve,
Conch. Icon., 3, pl. 5, f. 16,/1848;) Tryon, Man., 7, p: 164,
pl. 1, f.4, 5, 1885; Dalland Simpson, Bull. U. S. Fish Com.,
Hea 20%) LOOK.
Distribution. Hatteras to Colon, West Florida and Texas.
0-10 fms.
cinerea Gmelin, Syst. Nat., p. 3404, 1792; Reeve, Conch. Icon.,
Biiply220t) 124) 1846:) Pryous Nan 7,)) 106, pl \/2)) ters.
ROW 85: Dall Ball a7) Ue Sune iNn pst 20) 30:
Distribution.—Hatteras to Guadeloupe, West Florida and
Texas. 0-163 fms.
Genus TRIVIA Gray
pediculus Linnzeus, Syst. Nat., ed. X, p. 724, 1758,(as Cypraca);
Reeve, Conch. Icon., 3, pl. 23, f, 131, 1846; Tryon, Man., 7,
Py2or pl! 20/6. (94,95, Moose Mali Ta Ve MIU Sais DE
168, ’90.
IY4 MOLLUSCA OF THE GULF OF MEXICO 84
Distribution.—St Augustine, Fla., to Barbados and West
Fla. Pliocene, Caloosahatchie beds.
suffusa Gray, Descr. Cat. Cypr. p. 16, 1832; Tryon, Man., 7, p.
ZOT MMU 2T se Tao TSS sy Dalle Vi Cu ZS. 24.0%
1880.
Distribution.—Cedar Keys, West Fla., to Barbados. 1-3
fms., on reefs. Pliocene, Caloosahatchie beds.
candidula Gaskoin, Pr. Zool. Soc., p. 200, 1835; Tryon, Man.,
PED 203) Die) 2218) 23) 2a) 785 DalliBall ME CZ 8,
ZAQwIOO: Billy 372 Ul S. IN. Moy Diz Ooo.
olorina Duclos, approximans Beck.
Distribution.—Hatteras to Barbados. 18-140 fms. Gulf of
Mexico, Yucatan Strait, 640 fms.
globosa Gray, Descr. Cat. Cypr., p. 14, 1832; Tryon, Man., 7,
OMEZOON ple ama tay O2i F920 85;)) Dall iBall MeN CZs. 0p.
2S Or wll 27 Oe Sa ON.) Mp E20 eOr
pilula Kiener, sphaerula Mighels.
Distribution. —Gulf of Mexico, U. S. Fish Com. station 2373,
west of Fla., at 23 fms. Also Key Westand Antilles.
Pliocene, Caloosahatchie beds.
Genus ERATO Risso
maugeriae Gray, In Sowerby, Conch. Illustr., p. 17,pl. 7, f. 47,
Eee2ueryon Many.) 5.) p. (9). pli /asuty aawiaaw Sac alll. ili.
WW Ala S24 3 Da LOS LOOO:
laevis Emmons, N. Car. Geol. Rept., p. 262, f. 139, 1858.
Not of Donovan.
Distribution.—Hatteras to Antilles and Colon; West Fla.
and Vera Cruz. 0-63 fms,
Genus BURSA Bolten
aifinis Broderip, Pr. Zool. Soc., p. 179, 1832, (as Ranella);
85
BULLETIN 38 LY)
Mnyon. (Mani 2.) pudom rola anita a Senn oS i) ally ie aillemezn
WSUS NS ME pine 24 y Son (asi Gy renee)
Distribution.—Hatteras to the Antilles. Varietv cubaniana
d’Orbigny, dredged on Arrowsmith bank, Yucatan, also off
Key West and Cuba.
Genus DISTORTIO Bolten
reticulatus Link, Beschr. Rostock Samml., 3, p. 122, 1807, (as
Wistorines)); Mall Bully 27) (0 Sean vik 122.) So. Dall
and Simpson, Bull. U, S. Fish Com., 1, p. 416, 1901; Mitch-
elinniist mlesc! Sells. cm arm
Distribution.—Hatteras to Barbados, West Florida and
Mustang Island, Texas, 22-154 fms.
reticulatus var. clathratus Lamarck, An. s. Vert., 7, p. 186,1822,,
Inc. Me’th., pl. 413, f. 4, (as Z7ztov); Dall and Simpson,
BullesUs Sy) ish Com! pya16, noon:
Distribution.—Porto Rico and Gulf of Mexico near Key
West (dredged by U.S. Fish Com.), also at Chandeleurs,
La. Pleistocene, New Orleans pnmping station No. 7.
Genus COLUBRAR!A Schumacher
lanceolata Menke, Synopsis, p. 87, 1828, (as Ranella); Reeve,
Conch: Leon)! 2,) plus Wi opens 4m n(asi 77722072) Kad Stee
Conch Cabs; spl.65,) £08) myonuneViamet 2) pL 27h plemuomtes
1o2, Test: Dall Bully s7.(UemowNn vic "Pp: 132.0080 Meal
and Simpson, Bull. U.S. Fish Com., 1, p. 416, 1901.
Distribution.—Hatteras to Barbados and Vera Cruz on the
Gulf coast. Genotype
testacea Moerch, Yoldi Cat., 107, (asZ77zton); Mal. Blatt., 24, p.
25 lnyon, Man:i(2 ol) ijn eesrssr. Dall Bulle sar Oe
SH NEEM pe 24 "SoMa isu Nile S25. 35s LOS OO-
Distribution.—Hatteras to Sombrero, and Texas region of
the Gulf of Mexico. Pliocene, Caloosahatchie River, West
Florida.
1:26 MOLLUSCA OF THE GULF OF MEXICO 86
Genus CYMATIUM Bolten
olearium Linnzeus, Syst. Nat., ed. XII, p. 1216, (77zton); Reeve,
Conchilicon aiuig2 yt sa4cMrvon Vlaneii aN Osim. DIAS) h.
27a Sie Dal oBalleya7 OS. IN Mey pale 2h SO (asia ane-
QuUsia) ww pareschs, Bully Ue Ss: Ne Me, py O4 i 1oms.
Distribution.—Hatteras to Cartagena, Colombia, and the
Texas region of the Gulf of Mexico. Also Indian Ocean,
Port Alfred, South Africa.
chlorostomum Lamarck, An. sans Vert., 7, p. 185, 1822, (as 777-
tomeKeiener,. icon, Coq. -Viv., p;) 19), pl.) 12) 4.123, Pryon,
Mammen ie pla Teka 48 Heer Dalle Wales 7 WES.
INGEN ps132) 895) Dall’ andi Simpson Bully \Us (S) Bish
Com., 1, p. 417, 1901,) as Lampusza).
Triton pulchellus C. B. Adams, Contr. Conch., p. 60, 1850.
Distribution.—East Florida to Barbados and Gulf of Mexico
along the Mexican coast.
graci'e Reeve, Conch. Icon., 77cton f. 58, 1845; Tryon, Man., 3,
DiEuee-o7 4 1SSi Dall. Bull. MCHA ae Teihpy227, plei2oy te
PSs ules UL S\N). Mi. Tae Mole aoivater 26nS ou (as
Lampusia).
Distribution.—Cape Catoche, Yucatan, 24 fms., U. 5. Fish
Com. Also Colon and Barbados. —Max. depth Ioo fms.
cynocephalum Lamarck, An. sans Vert., 7, p. 184, 1822, (as 777-
ton); Reeve, Conch, Icon., 2, pl. 8, f. 26, 1844; Tryon, Man.,
Be {Os WO, ile iiny 0 On ole uscpe igs. oes ION ie rodblkM eye Ole
SPNCOMei pr ii22)?89-) Dall and) Simpson, Bull. o.bishi
Com), 1, p. 417, 1901, (as Lampusza):
Distribution.—Florida Straits to Margarita Isl. Texas re-
gion of the Gulf.
Genus NYCTILOGCHUS Gistel
femorale Linnzeus, Syst. Nat., ed. X, p. 749, 1758, (as Murex);
87 BULLETIN 38 1l7
Reeve, (Conch. i lcom ian olay ite 22 tensa ae (asmminetos)):
Aryon,; Many iasip. Towiplnon Zon nse Dalla Billa iw
S. N. M., p. 132, ’89; Dall and Simpson, Bull. U.S. Fish
Comin i) p47", Toe), (as) Zotoz.21270))):
Distribution.—Cedar Keys, West Fla., to Guadeloupe.
Genus ASPELLA Moerch
scalarioides Blainville, Faune Francaise, p. 131, pl. 5,f. 5, 6,1826,
(as Wurex); Dall and Simpson, Bull. U. S. Fish Com., 1,p.
410, IQIO.
Distribution.—European and Antillean.
scalarioides var. paupercula C. B, Adams, Contr. Conch., p.
604 1850,) (as J7uvex) Wall Bali Zi 8. . (2ZOSsnsg:
Billa e TU Se IN) Mee oat 2Ou aoe
Triton cantrainet Recluz, Jour. de Conch., 4, pp, 246, 418, pl.
8, f. 10, 1853; Kobelt, Jahrb. Mal. Gesell., 4, p, 244, 1877,
(as Ocinebra).
Distribution.—West Florida, at 50 fms.; Texas and the
Antilles.
scalarioides var. obeliscus A. Adams, Pr. Zool. Soc., p. 260,
HeGIn iryon, Man. 2ipylmzotmple a8.f)\/467)) t88Os. Dall
Buallkys7 US Si. UNM Spm Ou So!
Distribution.—Vera Cruz to St. Thomas. Texas region of
the Gulf.
Genus OOCORYS Fischer
abyssorum Verrill and Smith, Tr. Conn. Acad. Sci., 6, p. 177,
(not the fig, which is O. sulcata), 1884, (as Benthodolium);
Dall Balls MI COZ Ts pez ou eso pull 27.7 Un nse IN,
JOE Ga em tidroxeyy
Distribution.—Northern part of the Gulf of Mexico, U. 5.
Fish Com. station, No. 2400, in 169 fms. Also off Caro-
lina. Max. depth 2221 fms.
MOoLLUSCA OF THE GULF OF MEXICO 88
Genus CASSIS Lamarck
cameo Stimpson, Silliman’s Am. Jour. Sci., p. 443, 1860; Tryon,
Mame hor 27 olen vten SOs 3 Sau aliM alana) Use IN:
Met Da lsan SO:
Distribution.—Hatteras to Barbados and West Fla.
tuberosa Linnzeus,Syst. Nat., ed. X, p. 735, 1758, (as Buccinunz) ;
Reeve Conch, Icon.) 5) pli 3) 7 f) 74848; “iryon, Man.
Conch 7 (pe. 27 Pl. 2h f)50.0 795 moat Bulk seo. IN
M., p. 134, ’89; Dall and Simpson, Bull. U. S. Fish Com.,
PP Ww4Loy LOOL.
Distribution.—Hatteras to Barbados and West Fla.
(Cypraecassis) testiculus Linnzeus, Syst. Nat., ed. X, p. 736,
MAG SUAS OeccLTUMe\\ Neeve. | ConchuwMliconme V5) plyiants TO}
HOS wLEVvon | Manis 7p) 272. ple 2iwteoa pls Anite O25) (85)
Dall, Bull. 37, U.S. N. M., p. 134, ’89; Dall and Simpson,
Bulle WS) Mish Coma) p: 418) Tro]8n
Distribution.—Hatteras to Trinidad, W. I., and the Texas
region of the Gulf.
(Semicassis) inflata Shaw, Nat. Misc., 5, p. 22, pl. 959, 1812;
ivony Man 7 pura pla ihe OS es Dall Bulli 37. U:
S. N. M.,p. 134, ’89; Dall and Simpson, Bull. U.S. Fish
Com., I, p. 418, tg0I; Mitchell, List Tex. Shells, p. 6; Van-
atta Ao NaSa Phila: |) 55...) 758 0rOO3:
Distribution.—Hatteras to Brazil, Gulf coast: Calhoun Co.
West Fla.; Point au Fer, Chandeleurs, Lost Island, La.;
Galveston, Tex. Pleistocene, New Orleans pumping stze-
' tion No. 7,Knapp’s No. 2 well, Terrebonne Parish, at 1791-
1842 feet.
Genus ONISC {DIA Swainson
dennisoni Reeve, Pr. Zool. Soc., p. 21, 1842, (as Onzscia); Tryon,
Manne p 252, plato: i 20;17 85-4 Dally) Bulls MeiC. Ze yrs,
Pa Zensoe
D
BULLETIN 8 Es)
w
OQ
Os
Distribution.—Arrowsmith Bank, Yucatan, at 130 fms.
Also Guadelonpe Island. (Type locality).
Genus TONNA Brunnich
galea Linnzeus, Syst. Nat., ed. X, p. 734, 1758, (as Buccinum);
Reeve, Conch. Icon., 5, pl. 1, 1848; Tryon, Man., 7, p. 261,
pl. 1, f. 3, 7°85; Dall and Simpson, Bull. U. S. Fish Com., 1,
Pp. 419, 1901. (Allas Dolzzm).
Distribution.— Hatteras to Trinidad, West Florida, Louisia-
na, Galveston and Matagorda, Texas. Also Mediterranean.
perdix Linnzeus, Syst. Nat., ed. - S 734, asBucciuune) ;
Reeve, Conch. Icon., 5. pl. 6, , 1849, (es Dok cm); ‘Tryon,
Man., 7, p. 264, pl. 3. £8 15, ie wa 23-25, 785; Dall, Balk:
s7.0Ul: SN. M., p. 134, "So:
SG
Distribution.—West Florida to Brazil.
Genus PYRULA Lamarck
papyratia Say, Jour. A. N. S. Phila., 2, p. 238, 182
Man... 7, p..266, pl. 6, i 35565 > mall) Tr. W_ 2.
£63, °90; Vanatta, Pr. A. N. S. Phila... 55, p-. 7
on
YD
fed ;
Oo
(@)
Oo wo)
Distribution.—North Carolina to the West Indies. Gulf
coast, Calhoun Co., Fla. and Texas. Pleistocene, New Or-
leans pumping station No. 7 and of Fla. Pliocene, Caloos-
ahatchie River.
Genus STROMBUS Linneus
pugilis Linnzeus, Syst. Nat., ed. X, p. 744, 1758; Reeve, Conch.
Icon., 6, pl. 16, f. 39-41, 1851; Tryon, Man., 7, p. 109, pl. 2
f. 13-15, °85; Dall, Tr. W. I. S., 3, p- 177, °90; Singley, 4th
Ann. Rept. Tex., p. ou "92; Dall ae Simpson, Bull. U.
S. Fish Com., 1, p. 422, - Vanatta, Pr. A. N. S. Phila.,
L20 MoLLUSCA OF THE GULF OF MEXICO go
DAS on 1903; Vanghan, Publ, No, 123) Carn. Inst.,)-p:
sifu) Woe),
Distribution.—Hatteras to Colon. Gulf coast; Cedar Keys
and Calhoun Co., West Fla.; Point au Fer, Chandeleurs,
Cameron, La., Galveston and Corpus Christi, Tex. Pleisto-
cene, Grand Chenier, La., New Orleans pumping station
No. 7; North Creek, and Labella, West Fla. Pliocene, Ca-
loosahatchie River.
costatus Gmelin, Syst. Nat., p. 3520, 1792; Tryon, Man., 7, p.
Os 1So57 Dall, Bull. 37, U. So iNe Men ps 136; 307) Dall and
Simpson, Bull. U. S. Fish €om., 1, p. 422, 1901.
accipitrinus Reeve, Conch. Icon., 6, pl 7, f. 12, 1850; Tryon,
Vie munp LAN Tents, 75.
Distribution.—St. Augustine, East Fla. to Guadeloupe, W.
I. Also West Fla.
Genus SEGUENZIA Jeffreys
monocingulata Seguenza, Boll. Comitato Geologico, 7, p. 188,’76;
Watson, Chall. Rept. Gastr., p. 105, 86; Dall, Bull. M. C.
Zon, US iO, Ay, SOP io Leos, UNG INC. Go, iMiay Fall Kors sic
88, 89, 18869.
formosa Jeffreys, Rept. Valorous Cruise, p. 200, 1876; Dall,
SGN WL (CL AL) JO, Ale texe=
Distribution.—Western North Atlantic and Bay of Biscay.
Gulf of Mexico, West Florida and Texas regions. 100-2023
fms.
trispinosa Watson, Jour. Linn. Soc., 14, p.. 591; Chall. Rept.
Cashes Plants 14s 01Ss0;” lnyont\ anny sO. uple
Cai 7 Osoo, 1so7) Dally Bully 27a Uni Sm Ne Mien. 42)
1880.
Distribution.—Hatteras to Pernambuco, Brazil and the Tex
region of the Gulf. 294-675 fms.
No)
i)
BULLETIN 38 Tat
iconica Watson, Jour. Linn. Soc., 14, p. 589, 1879; Chall. Rept.
Gastr 15.) po io7. plaza i ats So7 ml nyonul Man yiouape
Mohs sollyntswin dane lain crest rectyel SO ANOUN 1 BoUUls Wl) (Ce Aat ies, yoy. BOG).
1880.
Distribution.—Gulf of Mexico, Florida Strait region, Culebra
Isl., W. I., and off the Azores. 390-1568 fms.
Genus TRIPHOGRA Blainville
perversa var. nigrocincta C. B. Adams, Jour. Bost. Soc. N. Jel Bs
p. 286, pl. 4, f. 11, 1839, (as Cevzthzum); Gould, Inv. Mass.,
Pens23,ih- 1592, 013707 DalliPr Ue Sena MOM pag anoge
Ballis7 WS. IN Me) pln 38, 78osnoimeley, 4th, Anni iepe.
CUS Mext op. 238i oz Mallia NN ES Muis Dl 2OganO 2k
Johnson: \Oces Pal Bost) Soc. MNAiHe 7) pe T2O. rons:
Distribution.—Massachussetts to Barbados. o-30 fms. Gulf
coast: Cedar Keys, Florida, Corpus Christi, Texas. Plio-
cene, Caloosahatchie beds, West Fla.
decorata C. B. Adams, Contr. Conch., p. 177, 1850, (as Cerith-
Zee Dally Dr Wey iS ase a2 Om Aulsoe2e
Distribution.—Gulf of Mexico, north and east shores, and
the Antilles. Also Lower Miocene of the Chipola marl,
West Florida. Variety oltvacea Dall, (Bull. M. C. Z., 18,
Pw24AN TS8o7 Bull 275 USING ON ep hl 84) 8O)) i Gltlisot
Mexico, west of Florida, in 50 fms. Also Key West and
Antilles.
colon Dall Bull). Mo CZ. vol RSGOMs ers ES) p. 247.) pli zou.
T2NMS Oe Bulla 7 We So Ne Vien ane sen ple) 2Os) te 2 SOE
Distribution.—Yucatan Strait, 640-1002 fms. Also off
Havana, Cuba, 450fms.
triserialis Dall, Bull. M. C. Z., 9, p. 84, 1881; 18, p. 246, pl. 20,
122 MoLbLusca OF THE GULF OF MEXICO 92
Mesa Oa 18897 (Balls U1 Ss New perasapl20, 6.054,
6a, 1808.
Distribution.—Yucatan Strait,640 fms. Also off Barbados,
154 fms.
bigemma Watson, Jour. Linn. Soc., 15, p. tot, 1880; Dall, Bull.
Me CH Z anon ps Si.) 18er. Watson, Chall Repr.,Gastr. yp:
Roermieageh. 6. 785". Dall Bully MiCa Zins) p.243,) 789.
Distribution.—Vucatan Strait, 640 fms. Also off St. Thom-
ase Wel) 200 fms;
hmcus all Bull. M- C. Z., 9, p. $3, 18813) 18, p. 249, pl. 20, f.
ISOM OU a7 Oi Sc Ne Mi ps T3Z8s pl 20%) fo iI SO,
Distribution.—Yucatan Strait, 640 fms. Perhaps a variety
of dzgemma Watson.
abrupta Dall, Bull. M. C. Z., 9, p. 84, 1881; 18, p. 249, pl. 20, f.
©, "BOs Iebly Sar Wk) See ING IME jon 1624S), Olly BO, 3s Cy evo
Distribution.—VYucatan Strait, Cape San Antonio, 640
fms.
torticula Dall, Bull. M. C. Z., 9, p. 82, 1881; 18, p. 249, pl. 20,
Pore so Biull aes 5. No NE pains se pl 20Mfin tb;
1889.
Distribution.—Yucatan Strait, 640 fms.
inflata Watson, Jour. Linn. Soc,, 15, p. tor, 1880; Dall, Bull.
MICA eno pi Si L881; Watson, \ChalliiRepey CGasth., ‘p:
FO plow it 285: Dall, Bulli Mi CunZe ss nps i240,
1889.
Distribution.—Yucatan Strait, 640 fms. Also off Georgia,
St. Augustine and Culebra Isl. 294-640 fms.
inflata var. ibex Dall, Bull. M. C. Z., 9, p. 86, 1881; 18, p. 249,
(Ol, BOS wig alo esky!
Distribution.—Yucatan Strait, off Cape San Antonio, in 640
fms. Also off Havana, 450 fms.
93 BULLETIN 38 123
cylindreila Dall, Bull. M. C. Z., 9, p. 83, 1881; 18, p. 250, pl. 20,
PON. So.) Bulls 27 aU mou INE MEMO. MsSiunli aor talGai2oO.
Distribution.—Cape San Antonio, Yucatan Strait, 640
fms.
Genus CERITHIOPSIS Forbes and Hanley
greenei C. B. Adams, Jour. Bost. Soc. N. H., 2, p. 287, pl. 4, f.
Ly 1820. (as) Gerzihzum)) Dallas mila Men on N on Evie aoe
ESS pl 52 hl2</18Or) Blt NVA OMuen DA ZOO" 7O2- i) Ohl Soms
OccewPaisBosts Soc. Ne ee .7.) pale On mols
Distribution.—Prince Edward Island to Dominican Repub-
lic and west of Cedar Keys, Fla., Corpus Christi and San
Antonio, Texas. 3-10 fms. Pliocene, Caloosahatchie River,
Florida.
crystallina Dall, Bull, M. C. Z., 9, p, 89, 1881; 18, p. 254, pl. 20,
fh SO; Hell 3 7 UL Se7 NEM ON tS. lin2Oumtin 2Ommea
°89; Dall and Simpson, Bull. U.S. Fish Com., 1, p. 424,
IQOl.
Distribution.— Washed ashore dead, Cedar Keys, Florida,
dredged alive west of Florida in 50 fms. Common in An-
tillean dredgings. 50-805 fms.
martensi Dall, Bull. M. C. Z., 18, p. 255, pl. 20, f. 2, ’89; Bull.
27 UL SE INE ME pe 28 plea oust h iSO:
Distribution.—Gulf of Mexico, west of Florida, at 1181
fms.
vanhyningi Bartsch, Pr. Biol. Soc. Washington, 31, p. 135,
1918.
Distribution.—Tampa Bay, West Fla.
burkevillensis Dall, Pr. U. S. N. M., 46, p. 231, pl. 22, f. 5,
IQI4.
Di stribution.—Upper Miocene or Lower Pliocene, Burke-
ville, Texas.
124 MOLLUSCA OF THE GULF OF Mexico 94
(Eumeta) subulata Montagu, Test. Brit. Suppl., p. 115, p!. 30,
PVG WTSOSMiCasidu7ex) Dall Balle MAC i Zar Bi) ps) 252i:
ZONA SO UM a 7n MOL Sa Ne Me De bao PLY 2x ti. )i4)
PINS 2 hei Sor jonnsou,\|Ocel Pa Bost) SOC IN.) El.) 7), p-
LZ0 VOWS:
emersoni C. B. Adams, Jour. Bost. Soc. N. H., 2, p. 284, pl. 4,
f. 10, 1838, (as Cerzthtum) Verrill, Inv. Vineyard Sound, p.
OAs ple24nnt. 151.1873.
punctatum Philippi, 1848. Not of Bruguiere, 1789.
Distribution.—Massachussetts to Grenada, W.I. Gulf of
Mexico, Cedar Keys, Texas and Yucatan. 2-15 fms. Plio-
cene of the Caloosahatchie.
Genus SE/LA A. Adams
adamsi H. C. Lea, Tr. Amer. Phil. Soc. 2d ser. 9, p.. 42, 1845,
(Gs Cerethzm)< Dally: We TS. 2p i2o7.018o2)\)Vanatta
Pi eae ONS. eladlasiis 5.) ps) (758, LOOgsmloumsons Occ.) Par
BOSHESOCH ING (ela 7\ py 127. TOs:
terebralis C. B. Adams, Jour. Bost. Soc. N. H., 3, p. 320, pl.
aut. )74 1840 (as Gertthium): De) Kay, IN2)Y.) Moll) p. 130,
OIoe ie ae. DalluBulls MIC. Ze isapaz5ou soul:
SeoONE Mig pins 8) pole 52).°£.15,)) (SO NOt Cemsrerume icercunale.
Lamarck.
Distribution—Massachussetts to Samana Bay, Dominican Re-
public. o-20fms. Gulf coast: Cedar Keys, Fla.; Chande-
leurs, La.; Galveston and Corpus Christi, Texas, 0-20 fms.
Pleistocene, Gulf coast. Pliocene, Caloosahatchie River,
Fla. Lower Miocene, Chipola marl, West Fla.
Genus CERITHIUM Bruguiere
floridanum Moerch, Mal. Blatt:, 23, p. 114, 1876; Dall, Bull. 37,
WAS TONG Me os 240) 18. Lr Welle Siete oS 2nn pla Tale ts
ROW so2- 1 Vauoham Publ Now nagar last) pa) Tn
IgIo.
Distribution-—Hatteras to Cuba and West Fla. Pleistocene,
Labella, West Fla. Pliocene, Calaosahatchie beds.
95 BULLETIN 38 L25
algicola C. B. Adams, Pr. Bost. Soc. N. H., 2, p. 5, ’48; Tryon,
Mant9, pa l2o.pl22 i380) So. 8o- Dall ire Nia elaa sia ae
Py2825));92.
Distribution.—Tampa and Charlotte Harbor, West Florida
to Jamaica. Between tides. Pliocene, Caloosahatchie
beds.
litteratum Born, Mus. Cees. Vind., p. 323, pl. 11, f. 14, 15, 1780;
Dall and Simpson, Bull. U. S. Fish Com., 1, p. 425, 1901.
Distribution.—East and West Florida, Antilles and
Europe.
muscarum Say, Amer. Conch., 5, pl. 49, f. 1, 1832; Tryon, Man.,
Ook ahead ener alee 3D ent. Iie NW Ih Sun iy) 0, Oe,
Vaughan, 2d Ann. Rept. Fla. G. S., p. 148, 1909.
Distribution.—Florida to Jamaica. Gulfcoast: Cedar Keys
and St. Mark’s, Fla.; Chandeleurs, La. Pleistocene, Mana-
tee, Labelle and North Creek, Fla. Pliocene, Caloosahat-
chie beds.
variable C. B. Adams, Pr. Bost. Soc. N. H.. 2, p. 5, 1848; Sow-
ery, Conch! icon... 15 ply igiieronamso5: Dall) Bille yore
U. S. N. M., p. 140, 1889; Mitchell, List Texas Shells.
Distribution.—Tampa, Fla.; Galveston, Corpus Christi,
Carancahua and Espiritu Santo Bays, Tex. South to the
Antilles and Curacoa.
minimum Gmelin, Syst. Nat., p. 3564, 1792, (as Murex); Tryon,
Man., 9, p. 167, pl. 34, f. 9-11, 1887; Dall and Simpson,
BallU.S. Fish Com: 174260 100%.
Distribution.—Tampa, Fla., to Guadeloupe Isl.
minimum var. nigrescens Menke, Synopsis, p. 85, 1828; Dall,
Boll27. 00! SouN. Mee pamnao a So)
Distribution.—Tampa to Venezuela.
eburneum Bruguiere, Ency. Me’th., pl. 442, f. 1a, b; Tryon,
126 MOLLUSCA OF THE GULF OF MEXICO 96
Maite won pat Zou plole tn 7 lA 72 Nn noo 7m Dalla tila a7 sins.
N. M., p. 140, 1889; Singley, 4th Ann. Rept. Texas Surv.,
PaissOn L892!
Distribution.—Florida to the Swan Isls. Cited by Singley
from Corpus Christi, Texas.
galvestonense Harris, Bull. Amer. Pal, vol. 1, p. 104, pl. 4, f. 9,
ga, 1895.
Distribution.—Upper Miocene, Galveston well at 2236-
2920 feet.
Genus CLAVA Martyn
chipolana Dall, Tr. Wagner Inst. Sci., 3, p. 290, pl. 22, f. 8,
1892.
Distribution.—Lower Miocene of the Chipola marl, Alum
Bluff, Fla., and of the Bacom well, Mobile, Ala., at 1241 ft.,
Chipola horizon.
Genus CERITHIDEA Swainson
costata Wood, Mal. Blatt., 23, p. 90; Dall, Pr. U. S. N. M., 6,
Pees Toos bw a7Ul 2 IN) MEM peniAoNTso:
Distribution.—Tampa, West Fla., to St. Thomas and Ja-
maica.
scalariformis Say, Jour. A. N. S. Phila., p. 128, 1825; Tryon,
Mansons tos) plage. t1. 73.) S70 Walilanby ull ras TNC
MEF) Pad, So.
Distribtion.— Georgia to Cedar Keys, West Fla.
varicosa Sowerby, Genera Shells, No. 42, f. 5; Dall, List Came-
ron Shells; Bull. M. C. Z., 18, p. 259,’89
Distribution. —Texas, Breton Island, Chandeleurs, La., and
south to Cuba and Jamaica.
turrita Stearns, Pr. Bost. Soc. N. H., 15, p. 24, 1872; Tryon,
Manving.cp. Tod ply sau hk 82.0787 Dalla dm We TenSss 20 pe
290,1892.
Oy BULLETIN 38 I27
Distribution.—Cedar Keys, Fla., to the Bahamas. Pliocene
of Shell Creek, Fla.
hegewishi Philippi, Mal. Blatt., 23, p. ’89; Zeitschr. fur Malak.,
Pp) 19,1841; Singley, 4th Ann.) Rept!) Dex.) pi aso, \18o2e
Mitchell, List Tex. Sh.
Distribution.—Cited by Singley and Mitchell from Matagor-
da, Carancahua and Corpus Christi Bays, Texas.
Genus POTAM!IDES Brongniart
matsoni) Dally) Pr. S, .Ne OM taonpayzennpl. 2a.) fo) Tonge
1914.
Distribution.— Well near Alexandria, La., at 49 feet; well
at Pine Prairie, La., at 1540 feet; surface near Burkeville,
Texas.
Note.—The brackish water molluscan fauna of Burkeville,
Texas; Alexandria and Pine Prairie, La., was referred in
1914 by Dr. Dall to the Pliocene. Dr. W. D. Matthew thought
the mammalian remains (tibia of a rhinoceros and upper
molar of a horse, either Protohippus or Merychippus) indicat-
ed a Late Miocene or Early Pliocene age. For discussion of
the Burkeville beds (lower member of the Fleming clay) see
Dumble, Univ. Tex. Bull. No. 1869,pp. 224-225, 1918. The
stratigraphic relation of the Burkeville beds rather sug-
gests an Upper Miocene horizon.
matsoni var. gracilior Dall, Pr. U.S. N. M., 46, p. 231, 1914.
Distribution.—Well near Alexandria, La., at 49 feet; well
at Paririe Bluff, La.; near Burkeville, Tex.
Genus ALABINA Dall
adamsi Dall, Bull. M. C. Z., 18, p. 258, ’89; Bull. 37, U.S. N.
IME pro, S77 MVM Oe namo Nr2/70.. 7.92) 1 (UAUllanas
Bittium) .
128 MoLLUSCA OF THE GULF OF MEXICO 98
Distribution.—Hatteras to Haiti and West Florida, moder-
ate depths. Pliocene. Caloosahatchie River and Shell
Creek, West Fla.
cerithioides Dall, Bull. M. C. Z., 18, p. 258, ’89; Bull. 37, U.S.
INEM OIA: So: Nebr Wl S03 conn 765) Digimon su 7O2:
Dall and Simpson, Bull. U. S. Fish Com., 1, p. 427, 1901.
(All as Bztttum cerithioide).
Distribution.—Cape Lookout, N. C.. to Samana Bay, Do-
minican Republic. Singley lists with a question a shell from
Corpus Christi, Tex. Pliocene, Caloosahatchie River, Fla.
Genus BITTIUM (Leach) Gray
varium Pfeiffer, Arch. fur Naturg., p. 256, No. 139, I840, (as
Cerithium); Tryon,Man., 9, p. 152, pl. 29, £. 86, °87; Dall,
a VS Te Seale. 27/4. 92:) Dall andi Simpson, Bulle Us, S:
Fish Com., 1, p. 426, 1901.
wnoocrulum C.. Bs Adams, Pr. Bost. SociNe (HY pis, 184.5;
Sowerby’s Thes., p. 876, f. 210, 211, (as Cerzthium).
Distribution.—Chesapeake Bay to St. Thomas, W. I. Gulf
coast: Cedar Keys, Fla.; Chandeleurs, La.; Corpus Christi,
Galveston and Laguna Madre, Tex. Pleistocene, North
Beach. Fla. Pliocene, Caloosahatchie and Shell Creek,
Florida.
Note.—&ittium nigrum Totten, synonym of &. alternatum
Say, has been erroneously reported from Cedar Keys, Fla.
and Chandeleurs, La. These specimens were probably
varium. ‘True alternatum, tide Dall, does not extend south-
ward of New York Harbor.
DoiplexDall, Tr Woale'S.))3, p.1275)) pls arta 92:
Distribution.—Lower Miocene, Chipola marl, Fla., and of
the Basom No. 1 well, Mobile, Ala. at 1500-1556 ft., Chipola
horizon, Aldrich’s collection. A varietal form.
99 BULLETIN 328 129
galvestonense Harris, Bull. Amer. Pal.,vol. 1, p. 104, pl. 4, f. 8,
1895.
Distribution.—Upper Miocene, Galveston well at 2550-287¢
feet.
Genus MODULUS Gray
modulus Linnzeus, Syst. Nat.. ed. X, p. 757, 1758, (as Trochus);
Dall, Bull. 37, U. S. IN. M:, p: 142,/1889; Singley, 4th Aun:
Rept. Tex. G. S., p. 339, ’92; Dall and Simpson, Bull. U.
SH ish Comp 4 27) ar ook
lenticularis Chemnitz, Tryon, Man., 9, p. 261, pl. 48, f. 91,92,
1887.
Distribution.—Hatteras to Brazil and to Cartagena, Colom-
bia. Gulf coast: West Florida, Chandeleurs,La.; Galveston,
Corpus Christi, and Espiritu Santo Bay, Tex. Between
tides. Pliocene, Caloosahatchie and Shell Creek, Fla. Up-
per Miocene, Galveston well at 2552-2871 feet (identifica-
tion doubtful).
floridanus Conrad, Amer. Jour. Conch., 5, p. 107, pl. 12,f. 6, ’70;
ulgard, House of Rep ex Woc.y 1 pt. 2))\p) S87. ase
Dall Pr OS! IN NEO on ees oe. Bull’.37 Un SauNe
MO pada 24( 789; Tr OW IS eae ip 205.) \92° Vaugdaanmy
Bubly No. 133) Carn. Inst) joao no:
Distribution.—Hatteras to St. Thomas, W.I. Gulf coast,
Cedar Keys and St. Mark’s, West Fla. Pleistocene, Lake
Borgne borings, La., and North Creek and Labelle, Fla.
Pliocene, Caloosahatchie beds, Fla. and of the New Orleans
Gymnasium club well at 1200 feet.
Genus PACHYCHEILUS Lea
anagrammatus Dall, Proc. U. S. N. M., 46, p. 232, pl. 21, f,
SOLO LA.
Distribution.—Upper Miocene or Pliocene. Well near
Alexandria, La., at 49 feet. Also near Burkeville, T cxas,
130 MOLLUSCA OF THE GULF OF MEXICO 100
surface exposure. Brackish water formation.
satillensis Aldrich, The Nautilus, 24, pt. 11, p. 132, pl. 8, f. 1,
HA-CmMOl was eoramcaes)):) Wall ProvUis SN Mis p22)
1914,
Distribution.—Upper Miocene or Pliocene. Satilla River,
Ga., and Burkeville, Tex. Brackish water formation.
Suavismoale roc. We) S. NM: 46) \p.' 232). pl!) 2) £16) 9; 2614.
Distribution.—Upper Miocene or Pliocene. Well near Al-
exandria, La., at 49 feet, and well, Pine Prairie oil field,
La., at 1540 feet. Also surface exposure, Burkeville, Tex.
Genus CAECUM Fleming
floridanum Stimpson, Pr. Bost. Soc. N. H., 4, p. 112, 1851;
Mayon Man, /Se pi2n7)\ ply 66,4) 620786; Dally Bully a7.
WHRSHEN Moura Boe dir Wet Omak mao sn 72.
tmnegmlare de Kolin, Jour. de Conch.) 15))\p)) 46) pl.3,/.f. '6,
1867.
annulatum Emmons, Rept. Geol. N. Car., p. 274, f. 190,’ 58.
Not of Brown, 1844.
Distribution.—Hatteras to Bahia, Brazil. Also West Fla.
2-18 fms. Pliocene, Caloosahatchie, Fla.
instructum de Folin, Fonds de la Mer; Dall, Bull. 37, U.S. N.
NV De aLhAZ SO:
Distribution.—Hatteras to Tampa, West Fla. Pliocene to
Recent.
bipartitum de Folin, Fonds dela Mer; Dall, Bull. 37, U.S. N. M.,
Pi 142, 80.
Distribution.—Hatteras and West Florida.
carolinianum Dall, Trans. Wagner Inst. Sci., 3, p. 300, pl. 22, f.
Zs rso2s itis 37. WY iS.) Ney Mee haa NES so. (maine
only).
IOI BULLETIN 38 13k
Distribution.—Hatteras to Tortugas and to Egmont Key,
West Fla. 2-63 fms. Pliocene, Caloosahatchie River,
Florida,
glabrum Montagu, Test. Brit., 2, p. 497,1803, (as Dentalium);
‘Anyon,| Mani Sap.) 215.0 pliiGO, Foy rSSor mDallnu baller.
WEE SOUND V4 2a So.
Distribution.— North Carolina to Tampa, West Fla. Also
European. Pliocene of the Caloosahatchie and of England.
Miocene of Yorktown, Va.
cooperi Sanderson Smith, Ann. & Lyc. N. Hist. N. Y., 7, pp-
L544 LOS T8025..0,) (Py BOs ues 7 Oslin Oi. ania Sap:
22TH DU O7, ik SA Sse alltguAerNN ein SiS. ZOOM MS OZ.
Wanatta, Pro Al Ni S) imine igs. 7158.) 1903; Johnson,
OceiPa. Best. Soc) INTE omet2 An he 15)
costaium Neri) (Aa jour Setangs)i snips 253) pl: 0, tee
1872.
Distribution.— Massachussetts to the Antilles. Gulf coast:
Crooked Island, Calhoun Co., West Fla. Pliocene of the
Caloosahatchie and Shell Creek, West Fla.
pulchellum Stimpson, Pr. Bost. Soc. N. H., 4, p. 112, 1851; Shells
of New England, p. 36, pl. 2, f. 3; Hilgard, House of Rep.
Bx Doc. \1,°pt:) 2) p. S8oumerouy anyon, Man. 'Siaosmiam
pi 661.63.) 786; Dalle mB allie anOretsy ON. Mi... wea orale
KOwit. 122.) So: \Vanattan | bana Neon EP hilal scum mages
1903; }ohnson, Occ) PaiBostasocweNey El.) (74). 2 ahnolsn
Distribution.—Cape Cod, Mass., to the Tortugas. Gulf
coast: Crooked Island, Calhoun Co., Fla. Pleistocene, Lake
Borgne, La., borings.
Genus MEIOCERAS Carpenter
deshayesi de Folin, Annales Soc. Linn. Maine et Loire, 11, ’69,
Dal B all 275) SOL SN leis nig 8 Oy
Distribution.—Tampa, West Fla., to Jamaica, W. I.
132 MoLLuscA OF THE GULF OF MEXICO 102
undulosum de Folin, Ann. Soc. Linn. Maine et Loire, 11, 1869;
Dall Balls Ueto NS VE pw 14.2030)
Distribution.—Charlotte Harbor, West Fla. to Jamaica.
Pliocene to Recent.
nitidum Stimpson, Pr. Bost. Soc. N. H., 4. p. 112, 1851, (as
Guecum)- Tryon, Man.. 8, p. 222) pl) 67, f. 73; 86; Wall,
Miva S30 ps 3O2, 68922 Vanatta brs AUN] on ehilas
55) P- 758, 1903.
Distribution.— West Florida, at Tampa and Crooked Isl.
(Calhoun Co. ), to Jamaica.
Genus BIVONIA Gray
exeriay Dall Bull. MM. C. Z:. Marvard Colli;o.-p. 39; 1881; 18,
pazo4") pl 26, f..6, °8o; Bull. 37,U) SaNyM:, p.) 144) pl. 26,
Te Ony OOr
Distribution.—Cape Lookout, N. C., to Barbados. Gulf of
Mexico, West Florida and Texas regions, deep water. 3I-
1002 fms.
Genus VERNICULARIA Lamarck
spirata Philippi, Arch. fur Naturg. 2, pt. 1, 224, pl. 7, f. 1, 1836,
(as Vermiculus); Tryon, Man., 8, p. 187, pl. 55, f. 99, 100,
oor Wall Balla WS. Ne Iv pan ra pl Siete SOs
MNES) sp) 204, 92; Singleya 4th Amiy Kept, Gi iS)
slkexa 9.) 340,792) Johnson, Occ) Pal /Bost/SocsN.)H., 7, \p-
122 1915.
Distribution.— Massachussetts to the West Indies. Gulf
coast: West Florida; Matagorda and Corpus Christi Bays,
and Quintana, Tex. Pliocene of Shell Creek, &c, Fla.
Note.—Moerch described as lV. melanosclera (Pr. Zool. Soc.,
p- 174, 1861) a form from Vera Cruz which Tryon regarded
as a variety of sfzvafa, and on the same page of the Proceed-
ings Moerch described V. guadrangularts from Yucatan. This
was also regarded by Tryon as a variety of spzvata (See
103 BULLETIN 38 133
Tryon’s Manuel, 8, p. 187, 1886). Shells from the Gulf,
Blake station 36, at 24 fms., have been referred by Dall to
V. lumbricalis Vann. (See Bull, M.C. Z., 18, p. 261,/ 789)
but the true Linnean /umbricalts is Oriental.
nigricans Dall, Pr. U.S. N. M., 6, p. 334, 1883, (as /umbricalis
Vat. Ball a7 So) Ni ME tos MAAN SOE
Distribution.— West Florida and Texas. 2-14 fms.
Genus PETALOCONCHUS Lea
irregularis d’Orbigny, Moll, Cuba, 1, p. 235, pl. 17, f. 16, 18,’42,
(as \Vermetus); Dall, sBullea7 iia su Ne Meo p. 144. 188o.
Cra We S23 spaigonaiogr
Distribution.—Gulf of Mexico between Mississippi delta and
Cedar Keys, Fla., at 27 fms. Also Antilles to Guadeloupe.
Pleistocene of Fla. Pliocene, Caloosahatchie beds.
varians d’Orbigny, Voy. Ame’r Me'’rid., p. 456, pl. 54, f. 7-10,
reas nyon! (Mani: Som 7Omi pli gon hy 22.) 220786 allle
Cree Sas DAO smmoz.
Distribution.—Southwest Florida to Rio de Janeiro. Plio-
cene, Caloosahatchie beds. Oligocene, Tampa silex beds.
erectus Dall, in Agassiz, Three Cruises of the Blake, 2, p. 71, f.
Z2O7 SS) oulls Mei. Zins apyneo2znpla 2S st.) An SOM taille
BUM So NEM p44 LSM ineEAN eG.
Distribution.—Antilles and West Florida. 37-805 fms.
Pliocene, Caloosahatchie River, Fla.
Genus SIPHONIUM
nebulosum Dillwyn, 2, p. 1076, No. 19; Tryon, Man., 8. p. 184,
plisas it: S75 1886; Dall BullMia 7 Ue S) IN MAY py Ta an Sor
Distribution.—East and West Florida to Tortola.
Genus SILIQUARIA Bruguiere
squamata Blainville, Dict. des Sci. Nat., 49, f. 213; Tryon, Man.,
134 MOLLUSCA OF THE GULF OF MEXICO 104
SMPALOOMIpLY SS ai aG SOR Mall ys mile Mar MOMSEN en VIO) to:
144, 89.
Distribution.—Sarasota, West Florida, to Barbados. 2-163
fms.
modesta Dall, Bull. M. C. Z., Harv. Coll., 9, p. 39, ’81; Agas-
siz, Three Cruises of the Blake, 2, p. 71, f£. 296, 788; Dall,
PEEON LS IOS Oe AGH aokein Moy iPAetoyiy o) Us waXey) yea Lan PesKo\e | J8yal Ula Ne yriAl Bitsy
INOS) Peord4. ple) 26. f)4 8o.
Distribution.—Off Cedar Keys, West Florida, to Curacoa
Isl. 94-805 fms.
Genus TURRITELLA Lamarck
(Haustator) variegata Linnzeus, Syst. Nat., ed. X, p. 767, 1758,
(asi7z7lo) Reeve, ‘Conch: Icon!,; 5) plas.)f.)19)) 49" bryon,
Mian Ss ps TOS. pl. Ol. tf. 5857 SOs Wall Bulla Os) 5.1 Ne
M., p. 144, ’89.
Distribution.—Matagorda and Espiritu Santo Bays, Texas
to Cartagena, Colombia, and Antilles. Pleistocene Knapp’s
wells, Terrebonne Parish, La., No. I at 1660-1700, 2250-
2450 (?), No. 2 at 1434-1800, No. 3 at 1330-1375 feet.
(Haustator) yucatecana Dall, Bull. M. C. Z., 9, p. 93, 1881; 18,
D205, 0pl 20h 2 oo. bull. 27 UL om NGnMEN Dua Agy ple
BORE ann OO.
Distribution.—Gulf of Mexico, Yucatan Strait, 640 fms.
(Toreula) exoleta Linnzeus, Syst. Nat., ed, X, p. 766, 1758, (as
Turbo); Reeve, Conch. Icon., 5, pl. 6, f. 22, 1849; Tryon,
Mani \3\ 9p. 205,) pluio8: ‘99, 18386; DullkiBullaa77 Us S. NE
M., p. 144, ’89; Dall and Simpson, Bull. U.S. Fish Com.,
TDs AZOu TOOT
Distribution.— West Florida, in 50 fms.; Texas coast. An-
tilles to Barbados. 45-170 fms.
(Torcula) acropora Dall, Bull. M. C. Z., 18, p. 264, 1889; Bull.
BOR SHON: Means TAA SO.
105
BULLETIN 38 135
Distribution.—Hatteras to Grenada Isl. Gulf of Mexico,
West of Florida, at 14-50 fms., and the coast of Texas, Vera
Cruz and Yucatan. 3-413 fms. The Pliocene, closely relat-
ed form is 7. subannulata Heilprin of the Caloosahatchie
marl.
chipolana Dall, Trans. Wagner Inst. Sci., 3, p. 312, pl. 22, f. 24,
1892.
Distribution.—lLower Miocene of the Chipola beds, West
Florida, and of the Bascom No. 2 well, Mobile, Ala., at
1241 feet, Chipola horizon. Aldrich’s collection.
subgrundifera Dall, Trans. Wagner Inst. Sci., 3. p. 313, pl. 22, f.
22h TSO2.
Distribution.—Lower Miocene, Chipola beds, Calhoun Co.,
West Fla., and of the Bascom No. 2 well, Mobile, Ala., at
1241 feet, Aldrich’s collection. A varietal form was record-
ed by Harris from the Upper Miocene, Galveston well, at
2552-2920 feet.
terebriformis Conrad, Cat. Mio. Foss., Pr. A. N. S. Phila., p.
5034863; (Name only) Dall ir Wry le Ss.) 2enpa) amen
1892.
Distribution.—Lower Miocene, Chipola marl, Fla., and of
the Bascon No. 1 well, Mobile, Ala., at 1500-1556 feet.
Aldrich’s collection. Also Chesapeake Miocene of Easton,
Md. (Conrad’s type locality).
sajilla Dall, Proc: U.S) NM peieaa\pl 22) f 6.1914.
Distribution.—Upper Miocene or Pliocene.. Well near Al-
exandria, La., at 49 feet. Brackish water formation.
Genus MATHILDA Semper
yucatecana Dall, Bull, M. C. Z., 9,p. 90, 1881, (as Bzttzum); 18,
p1/266, pl. 20, £217.77 807 nulla amon ING Mie iplen20wcts
ae) rexOye
Distribution.—Gulf of Mexico, Yucatan Strait, 640 fms.
136 MoLLUSCA OF THE GULF OF MEXICO 106
Also off Georgia and Fernandina, East Florida. Least
depth, 294 fms.
Genus LITTORINA Ferussac
ziczac Gmelin, Syst. Nat., p. 3587, 1792, (as 7rochus); Reeve,
Conchwlcon (ron) ply iii £) STS 572 yon viata 70)! 9,
AT pAAS ie) 5 TSo7s) Dall Bullia7 Wl iS.) Ne M.D L460;
’89; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 429,
Igol.
Distribution.—Florida Keys west to Texas and South to
Barbados.
augulifera Lamarck, Anim. s. Vert., 7, p. 54, 1882, (as Phaszan-
Cia) Dally Bull 37.40. (S) Ney Mei pa i464) 80;) Dall and
Simpson, Bull. U. S. Fish Com., 1, p. 430, Igot.
Distribution.— West Florida and Texas to the Antilles and
Cartagena, Colombia.
mespilum Muhlfeld, Verh, Berl. Ges. Nat., 1, p. 219, pl. 8, f. 8,
no24ni(as: Aelia nyon,.. Man. Ovi ps 252s spl-4'5, 1h. 16)
18S7- Dall Bulle 37. U.S. Ne MES pain4oy So.) Dally and
Simpson, Bull. U.S. Fish Com., 1, p. 430, 190I.
Distribution.—Texas to Barbados.
inrorata. Say, Jour. A. N. S. Phila., (1); 2, p: 230; 1822, Tuo-
mey and Holmes, Pl. Foss.czSCar:, p: i109, pl. 26, f. 5,
L857); Bost.—Fl. Hoss. S, Car, por plait. 5, 1860;
ayon, Man.9,p. 246, pl. 43) £48) 7375) Wally Bull 37, U.
SUNCOM po 146 ole 69. fu Sonia Wen OH 2D. 13205
’92; Singley, 4th Ann. Rept. G. S. Texas, p. 340, ’92: Van-
atta ema ING Sa Plata: (9553) pa 75Sulnooss jounson, Occ.
Pan BOSta SOCeINGIEL 7. Pp. LZ LORS:
Distribution.— Massachussetts to Jamaica. Culf coast:
Cedar Keys, and Calhoun and Washington Cos., West Fla.,
Chandeleurs, Weeks Island, Point au Fer, La.; Matagorda,
Espiritu Santo Bays and Galveston, Tex. Pleistocene, New
Orleans pumping station No. 7.
107 BULLETIN 38 137
Fenus LITIOPA Rang
bombix Kiener, Ann. des Sci. Nat.,p. 203, 1829; Ann. des Sci.,
BOM DA 222 NLA iT aan MMT SMa Sea uMultay: Olntnl MN lealkiey
OQ PNZSL;) Pal sste 74s Mos WalluuB ull tac nSH NG yIN IGE!
148, ’89; Singley, 4th Ann. Rept. Tex., p. 340, ’92; Dall,
Bullen.) We Se Ne Mp Hinssui room lohnson, |Occy bas
BOSE SOC WIN Mie 7 Mei LO Munon 5
melanostoma (Rang) Sumner, Bull. U. S. Bur. Fish., 31, pt.
Zep ZO LOL:
Distribution.—New England to Brazil. Gulf coast at Cam-
eron, La.; Corpus Christi and Galveston, Tex. Also Pacific,
California coast. Pelagic on floating Sargassum.
Genus TECTARIUS Valenciennes
muricatus Linnzeus, Syst. Nat., ed X, p. 1758, (as 7urbo); Reeve,
Conch Icon.,) 10) pla. aera aSs7e(as “ztior77a)- Tryon
Man',9, p.258, pl. 48, £:°68,\1887>))Dall, Bull. 37; Ul S2iNe
M., p. 146, ’89; Dall and Simpson, Bull. U. S. Fish Com.,
WP WABT LOOT:
Distribution.—Hast Florida and West Florida and Antilles
to Colon.
Genus ECHINELLA Swainson
nodulosa Pfeiffer, Tryon, Man., 9, p. 258, pl. 48, f. 75, 1887,
(Es<cliisyn.): Dall Bulk WaiSiwiNe Wwe) po) 146.) ?8o: Dall:
and Simpson, Bull, U. S. Fish Com., 1, p. 431, 1901.
Distribution.— North Carolina to Barbados and Texas.
Genus ISAPIS H. & A. Adams
ebsoleta)) Dall (ProciyU) Sa uNiiiMneinA ON ol 222) umplliiao titi oe
IQT4.
Distribution.—Upper Miocene or Pliocene. Well near Al-
exandria, La., at 49 feet.
138 Mou.usca OF THE GULF OF MEXICO 108
Genus ALASA A. Adams
tervaricosa C. B. Adams, Pr. Bost. Soc., N. H., 2, p. 6, 1845,
(as Rissoa); Moerch, Malak, Blatt., 23, p. 57, 1876; Dall,
Bull. U. S. N. M., p. 146, ’89; Dall and Simpson, Bull. U.
Swish Comet pyaar.) T9OT.
Distribution.—Tampa, West Fla., to Haiti.
Genus ARCHITECTONICA Bolten
granulata Lamarck, Anim. s. Vert., 7, p. 3, 1822; Ency. Me'th.,
PMA. tf Sab om792)) (as Solazzum) Wall ire NV .5° 3,0)
Beo) T8920) Bo Ss. INU My) 1374p) 232n nolo; Mlauny. Bull:
mers Paley NO4)295) (ps1 Dl. 23, tsa onli
perspectiva Tuomey and Holmes, Pl. Foss. S$. Car., p. 120, pl.
26, 1857. Not of Linneeus nor Lamarck.
Distribution.—Hatteras to the Antilles. Gulf coast at
Chandeleurs, La., Matagorda and other localities, Tex.
Also Pacific Ocean, Lower California to Peru. Pleistocene,
New Orleans pumping station No. 7; Upper Miocene, Gal-
veston well at 2158-2871 feet. Lower Miocene, Chipola
marl, West Fla., and Dominican Republic.
Genus TORINIA Gray
cylindrica Gmelin, Syst. Nat., p. 3572, 1792, (as 7vochus); Tryon,
Manion pai7pluisd. 82, 18875 Dalle BallniaynnWy is! iN.
M., p. 148, ’89; Dall and Sampson, Bull. U. S. Fish Com.,
We pase, TOOL,
Distribution. —Texas to St. Thomas, W. I.
canalifera C. B. Adams. Dall, Bull. 37, U. S. N. M., p. 148,
1889.
Distribution.—Texas to Guadeloupe.
Genus RISSOA Fre’minville
nrecipitata Wall, Bull) Me C1207 18) psi27O ply tow f.) 1 e889:
Bulle VU SNe OME pe 14S plerronp terion SO.
Log BULLETIN 38 13G
Distribution.—Gun Cay, East Fla., 498 ims., Yucatan
Strait at 640 fms.
acuticosiata Dall, Bull. M. C. Z., 18, p. 280, pl. 19, f. 10, 1889;
Bell a7 US wNG ME pe SOM ply ToM tno MG Se:
Distribution.— Yucatan Strait, 640 fms., and between Mis-
sissippi delta and Cedar Keys, 32 fms. Also off Hatteras
and Barbados. Probably a variety of R. xanthias Watson
(Chall Gastr., p. 588, pl. 44, f. 5, 1885).
Genus RISSOINA d’Orbigny
decussata Montagu, Test. Brit., p. 399, 1803, (as TZurbo);
Schwartz von Mohrenstern, Denkschr. d. Kais. Akad. d.
Wissensch. Wien. 19, p. 80, f. 12, 1860; Tryon, Man., 9, p.
ro HOMIE AU ain tore IORNIL IRONS AVON (Cu ZAR eS Sun io), DSi,
1SOn MOL WIWis TS. 2p miaaan moo!
Distribution.—Cape Fear, N. C., to Haiti and to West Fla.
2-17 fms. Also Huropean. Variety plazata Dall, Caloosa-
hatchie Pliocene.
laevigata C. B. Adams, Contr. Conch., p. 14, 1850; Schwartz
von Mohrenstern, Denk. Kais. Ak. Wien., 19, p. 111, f. 79,
Hg60;)Tryou, Man.) 0) palig@ootmlunsouiy 943.0787 2) Dallaire
Wel S208 ne aa2 nook
Distribution.—Cape Lookout, N. C., to St. Thomas, W. I.
Also West Florida. o-22 fms. Pliocene, Caloosahatchie
beds. Lower Miocene, Chipola beds, West Fla.
chesneli Michaud, Descr. des Cog. Nouv., p. 17, 1832, (as Rés-
50g) bryon, Man. oul paiesomplamsOntn 7 3) S75 lDallenmerst
WIS. 3p: 343) o2-otueleyanatiay Anim. Rept lexenaos
BAM Mn O2:
Distribution.—Hatteras to Guadeloupe, Gulf coast: Cedar
Keys, Fla.; Corpus Christi, Tex. Also Mediterranean and
Mauritius. Pliocene, Caloosahatchie beds, West Fla.
browniana d’Orbigny, Moll, Cuba, 2, p. 28, pl. 12, f. 33, 35, ’42;
r40 MOLLUSCA OF THE GULF OF MEXICO TIO
nyouyy Manes oi ipisgo, pl.) Sour 45. « S7uaniattal, et Ie.
ING Selatan ss. DMT Sou OOS.
Distribution.— West Indies and Gulf coast, Crooked Island,
Calhoun Co., West Fla. (Vanatta).
elegantissima d’Orbigny, Moll. Cuba, 2, p. 26, pl. 12, f. 27-29,
Hose Vl myo) Mata iG, 09s 274) pls 50. tai thy Loo
Distribution-—West Indies and Gulf coast, Chandeleurs,
ae Cally):
Sagraiana d’Orbigny, Moll. Cuba, 2, p. 25, pl. 12, f. 4, 5, 1852;
atnyout Man wionpanssa) pl. 578 ti SOn mosTe: Dalley Bully) 27%.
Wao NMS Dp. 1/50; 780:
Distribution.—Florida Strait, to Martinique. Also Gulf
coast at Vera Cruz, Mex.
Genus BENTHONELLA Dall
nisonis, Wall, Bull’ Mi C.'Z., 18, p. 282, 18809; Bull) 27, U.S:
NEMEC TDs SOMO}
Distribution.—Gulf of Mexico between Mississippi delta and
Cedar Keys at 940 fms.
Gaza Dalal My Ci Z.. 18. p. 2825) 8o: Ballas aWians. IN. Me,
Ds 2 SOn) jolly gay sen ee soy
Distribution.—Georgia to Cuba,-West Florida and Texas
regions of the Gulf. 6-463 fms.
mschen Dall Bull, MaiCZ.)\ 18) 1282) So, Bulla Us S. NG
INR JOy SAN exo),
Distribution.—Gulf of Mexico between Mississippi delta and
Cedar Keys at 940 ims., bottom temperature 39° F. Also
off Cuba.
Genus SKENEA Fleming
planorbis Fabricius, Fauna Groenlandica, p. 394, 1780; Gould,
ity AMiaSsie ps 206s) f4563) iS 7Onwo alu alla sau ShoNG
Mi’, p. 150, pl. 52, f. 18, 1889; Bush, Tr. Conn, Acad. Sci.,
anit BULLETIN 38 TAL
LO, \p. Too: pls 22) f:'5.'8. 09-1900; Johnson, Occ) Pay Bose:
SOC INE JE 177 apis TO Me hose
flelix depressus Montagu.
Distribution.—Greenland (type locality) to Charlotte Har-
bor, West Florida. This species is the genotype.
Genus AMPULLARIA Lamarck
depressa Say, Dall, Bull. 37, U.S. N. M., p. 150, ’89.
Distribution.—Georgia to West Florida, Texas and Mexico.
caliginosa Reeve, Conch. Icon., Mon. Ampullaria,{. 118, pl. 25,
T8567) Mall, Procs) Us Ss NGWMeon ip. lah s.r Soy: itliaia ze
WET SEN Mei pet SOM ms Sor
Distribution.—Cedar Keys, West Florida to Texas, Mexico
and Nicaragua. In freshwater swamps.
pinei Dall, The Nautilus, p. 75, Nov. 1898.
Distribution.— Homosassa River, Fla.
Genus PALUDESTRINA d’Orbigny
aldrichi Dall, Proc. U.S. N. M., 46, p. 234, pl. 22, f. 7, 1914.
Distribution.—Upper Miocene or Pliocene. Well near Al-
exandria, La., at 49 feet. Brackish water formation.
planawAddrich, Nantilus) 24) ptmen|ps) 220, pl: 8) 1) 2amonm:
Dall wer U.S. (NM AGiipmezas Tora.
Distribution.—Upper Miocene or Pliocene. Well near Al-
exandria, La., at 49 feet. Also Satilla River, Ga., (type
locality).
curva, Dall. Proc. Uz) SyyNaMe 46.235, (pis 22,44 nomar
Distribution.—Upper Miocene or Pliocene. Well near Al-
exandria, Ga., at 49 feet.
cingulata Dall, Proc. U. S. N. M., 46, p. 235, pl. 22, f. 1, 1914.
Distribution.—Upper Miocene or Pliocene. Well near Al-
exandria, La., at 49 feet.
142 MoLLUSCA OF THE GULF OF MEXICO 12
turricula Dall, Proc, U. S. N. M., 46, p. 235, pl. 22. f. 9, 1914.
Distribution.—Upper Miocene or Pliocene. Well near Al-
exandria, La., at 49 feet. Brackish water formation.
milium Dall; Proc. U. S. N. M., 46, p. 235, pl. 22, f. 2, 1914.
Distribution.—Upper Miocene or Pliocene.. Well near Al-
exandria, La., at 49 feet.
Genus ASSIMINEA Leach
auberiana d’Orbigny, Moll. Cuba, 2, p. 8, pl. 10, f. 6, 7, 1842,
(as\Paludestrina): Wall) Pr: U.S.) Ne WMEs) 6) p. 335, 1883;
ables WS) INE Met.) TSO Son Miles nus" De SAT,
1892.
Distribution.—Cedar Keys, West Florida, and the Antilles.
Pliocene, Caloosahatchie and Shell Creek, West Fla.
Genus TRUNCATELLA Risso
caribaeensis Sowerby (MSS.); Reeve, Conch. Syst. 2, pl. 182, f.
7, 1842; Binney, Land & Fr-Water Shells, N. A. Smith.
Mise. Coll., No. 144, pt. 3, p. 98, pl. 198, 1865; Dall, Bull.
BRE) Ne NE pe ons52,8o;) Dall Sa Simpsont Bully U.S:
BiShiCom! wn 436, ple 53), £475) LOOM
Distribution.— West Florida, Alabama, Texas (?) and Mex-
ico. Also Greater Antilles.
bilabiata Pfeiffer, Wiegm. Arch., 1, p. 253, 1840; Binny, L. and
Bie We Shu iN Al pt.) 3, p:/ 991, t)/L99%) L865 Dall, Bull, 37;
We SeNe ME. wip 152), 89):
Distribution.—Sarasota, West Florida, to Honduras and
Cuba.
pulchella Pfeiffer, Wiegm. Arch., 1, p. 356, 1839; Binney, L. and
RmenWe Shi Ne AG pt) 3p) oot a2oommsog) Dall, Bully a7.
Was IN Mvp. 152, So; Dall /éz)Simpsony) Bulls U.S. Fish
Comin aps 4.304 LOOM:
Distribution.—Florida to St. Thomas, W. I. Gulf coast:
113 BULLETIN 38 I4
i)
Tampa, West Florida; Galveston, Texas. (Dr. Gurley).
subeylindrica Linnzeus, Syst. Nat., ed. XII, p. 1248, 1767, (as
Fleless);\ (Gray, in) Murtom, "Man p! 295) i472, 1185.7; pinueys
Bandvand Er. Wess sine AGM pts 25 pr OO, am ahZOR noo be
Dall and Simpson, Bull. U.S. Fish Com., p. 436, 1901.
Distribution.—Tampa, West Fla., to St. Thomas, W. I.
Genus CHEILEA Modeer
equestris Linneeus, Syst. Nat., ed. X, p. 780, 1758, (as Patella);
ayon (Man 8) paag7 pl.) ai hese 264 S38 oruDall lie. VE
TAS.) 2419) 248) so2' as wideciarca) rile Gael SEWN» IVES aris
Pal232. TOTO:
Distribution.—Hatteras to Barbados, Gulf of Mexico, Blake
station 2. Also Pacific Ocean, Mexico to Chile. Pliocene,
Caloosahatchie beds, Fla.
Genus CAPULUS Montfort
intortus Lamarck, Anim. s. Vert., 6, pt. 2, p. 18, 1822: d’Orbig-
mys Moll, (Cubal 1 (pilnS6 plea isd22) 2301842 idiom,
Mane 8 ip. 13a ipl zoet aig San oor OAc mB willy 27) 1 Wena iN
ME pal 545) (80:
Distribution.—Gulf of Mexico, Blake station 5, at 229 fms.
Also off Florida reefs, Hatteras and Bahamas.
Genus CREPIDULA Lamarck
fornicata Linnzeus, Syst. Nat., ed. XII, 1, p. 1257, 1767, (as
Patella’. Wilgard, House of Rep! Ex. Doc., 1, pt. 2, p. 8387,
19797 Dall,) Bull 37 (Un Sa aN iis 2) DliAS) ite ype
BOE.) 235) 2A.) TSSO-) Mid Ee! Sean). 35 03 OZ eoleLeye
ApwAnn. Rept. exp anime 2| WVanatta,: bt.) Aen Ss
Bhilays5)) 7.58; noes wjolusonhOce. Pa. Bost. Soc). vrEen
TPE LOO) LOLS:
Distribution.—Granada to Cartagena, Colombia. Gulf coast:
Cedar Keys, St. Mark’s, Ft. Barranca, &c., West Fla., La.,
Galveston and Corpus Christi, Tex. Pleistocene, Lake
LA4. MOLLUSCA OF THE GULF OF MEXICO 114
Borgne borings, New Orleans well of 1856, and New Orleans
pumping station No. 7. Pliocene, Caloosahatchie beds.
Lower Miocene Chipola beds, West Fla.
plana Say, Jour. A. N.S. Phila., (1), 2, p. 226, 1822; Tuomey
andmolmes)) PIN EKossS. (Car. p. Tima please ta 12.0857
Dalia lee ze UM Ss Ne MeN piso) pl yAsaten ta lin 5Onn ts
BOM SOn i WV lei Ss.) sh DYE 5 ol O22) Wanabbasiate Au VN) S)
Phila., 55, p. 752, 1903; Johnson, Occ. Pa. Bost. Soc. N. H.,
Fy [Da Meaty vere) HiGy,
Distribution.—Canada to Trinidad, W. I. Gulf coast: Cedar
Keys, St. Mark’s, Fla.; Chandeleurs, La.; Galveston, Corpus
Christi and Matagorda Bays, Texas. Pleistocene, New Or-
leans pumping station No.7 and North Creek, Fla.
Pliocene, New Orleans Gymnasium Club well at 1200 feet,
and Caloosahatchie beds, Fla. Lower Miocene Chipola beds,
West Fla. and of the Bascom No. 1 well, Mobile, Ala., at
1500-1556 feet, Chipola horizon.
aculeaita Gmelin, Syst. Nat., 7, p. 3693, 1792, (Patella); Tryon,
Mamie Da 2oh pl. 2o)1t 61-62) 1886. DalllBull Mes .9Z,
ES W260 1SSOn Vanatta,, Pr. ANS iehilaa S50 pel 758:
noosy Walk Buallyr1 200). (S.NEMEs pro 2n oom
Distribution.—Cape Lookout, N. C., to Barbados. Gulf
coast: Indian Pass, West Fla.,-and Texas. Usually 0-25
fms., but dredged in Gulf of Mexico, near Tortugas at 539
fms. Dwarfed and white at that depth. Also Pacific Ocean,
California to Chile. Pliocene, Caloosahatchie beds, West
Florida.
convexay say. Nour ANS. Philas (1) 402.) pl2o7iins22 J Dall,
JEU 7h. Who SR INIS IME NG os aula mole Evo). w3t,/ is. sis O ea Ate EN,
Ws Slop, Sy hOse Sven Ve
Distribution.— Nova Scotia to Florida. Gulf coast: West
Fla.; Chandeleurs, La.; Galveston, Tex. Pleistocene, North
Creek, West Fla.
115 BULLETIN 38 r45,
Genus CRUCIBULUM Schumacher
auricula Gmelin, Syst. Nat., p. 3694, 1780, (as Patella); Dil-
ways Cat.) 2, tonya Lowi all Bulls Nien Cs 72am es ar
234 /8O;buUlMia 7 UL ony INAIME OES 2 NmS oe
planatum Schumacher, Essai, p. 182, 1817.
scutellatum var. auriculatum (Chemn.) Tryon, Man., 8, p. 118,
DI B2n/ fi 24) i258 Sox
Distribution. —Cedar Keys, West Fla., and Texas, south to
Barbados and Northern Brazil, 25-111 fms. Pliocene, Caloosa-
hatchie beds, West Fla. The analogous Pacific species is
C. spinosum.
striatum Say, Jour. A. N. S, Phila., 5,:p. 216, 1826; (as
Calypiraca); Tryon) Man.) (8) sp. 118; pli 33, £144, 45,-86:
Dall Boils Mi Ca Zor pe 2 oA eo Ohnson.)) Oecumaar
IBOSst-.S0ci INGE 7. oul OO NNO 5:
Distribution.—Nova Scotia to Florida. Cited by Dr. Dall
from the Gulf of Mexico at Vera Cruz.
Genus CALYPTRAEA Lamarck
candeana d’Orbigny, Moll. Cuba, 2, p. 190, pl. 24, f. 28, 29,
1842, (as /nfundibulum); Dall, Pr. U.S. N. M., 6, p. 335,
493, 1(as Galerus) Vlnyon Mane no.) pala. ply a Ak itaeon
Tie rose: Malley Bull iowa one ies lie: 52h SOP
parvulus Dunker,1875.
Distribution.—Hatteras to Brazil. 6-52 fms. Gulf of Mex-
ico, Cedar Keys, West Florida, and Yucatan Strait, 640
fms., dead shell.
Genus XENOPHORA Fischer de Waldheim
conchyliophora Born, Index Mus. Ces., p. 333, 1778, (as Tvochus) ;
snyon, Mant, 68) pO plo, te SOON oon Dalla walla.
WAS OUN Mis p. 547 So lr Vial Sia Da1 30048 O2nIiole
AP TAO, 0a. +.92) BiUlliGowOs SiN Vea Ose ape siusnntelhl,
14.5 MOLLUSCA OF THE GULF OF MEXICO 116
SH SeeMiatinven, BillcAmen Pals INOMiZOMhpNmEaaiipl.W2 3. ie
7) IQIt7.
agglutinans Lamarck.
Distribution.—Hatteras to the Antilles. Gulf of Mexico,
West Florida and dredged at various stations, 14-229 fms.
Usually not so deep as X. carzdaca and more heavily loaded.
Pliocene, Caloosahatchie beds, West Fla. Lower Miocene,
Chipola marl, West Fla. Oligocene, Tampa silex beds, Fla.
Upper Eocene, Jackson, Miss. Eocene, Wood’s Bluff, Ala.
Upper Cretaceous, (Ripley) of Prairie Bluff, Ala. The
most ancient species of the Gulf Mollusca.
caribaea Petit, Jour. de Conch., 5, p. 248, pl. 10, f. 1, 2, 1856;
iyo Vian. 8, pw162,.pl..474 f Go Soa Dalles Bull Merc
ZS ps 261, 89° Dall) and Simpson, Bullanuey Ss Fish
Comes 438.) TOOL.
Distribution.—Hatteras to Barbados and Gulf of Mexico, .
various stations, 14-274 fms.
Genus NATICA Scopoli
livida Pfeiffer, Wiegm. Arch., 6, p. 254, 1840; Moerch, Malak.
Bate 24 pod gi) Dally Bulls Nie CZ knenn pn 20210780):
Bll 7a Why Sa NMG pet's) 1480:
jamaticensts C. B. Adams, Contr. Conch., p, 111, 1850.
proxtma Sowerby, Thesaurus, atica, pl. 8, f. 111. Not of
C. B. Adams.
Distribution.—Hatteras to Barbados. Gulf of Mexico, be-
tween the Mississippi delta and Cedar Keys, 26 fms.
maroccana Chemnitz, Conch. Cab., 5, p. 270, pl. 188, f. 1905-10;
Dillwyn Cat. Rec! Sh. 2) \p. 983) (Nonna sinsi7) Dall’) Bull:
NM Cre Zen ieee 202. 188Os) Billi27 Wun SweN ey VCs Dil s 4.
1889.
Distribution.—Hatteras to Barbados and Gulf of Mexico,
west of Florida. Also European and West African.
Igy BULLETIN 38 , I47
canrena Linnzeus (in part) Syst. Nat., ed. X, p. 775, 1758, (as
Nerita); Reeve, Couch. Icon., 9, pl. 4, f. 14, 1855; Tuomey
and) Ivolines, Pil Moss. (Ss Caren odin is wap ll i> Sanne an7 inns Suz
DEVI SxeN ea ea OP INT IGA ou larcyaliieKoe: ACen Nin Ibs S4
p304.7 025 (Maunye (Bull Amen pales Niow 2 ous ome Amine
Ber.) 3s KOH KONIG
pucaccila Conrad, Prva NS! Phila wipe s64.) soa.
Distribution.—Hatteras to the Antilles, Pernambuco and
Cartagena. Gulf coast, West Florida and Galveston. Plio-
cene, Caloosahatchie beds, Upper Miocene, Galveston well
at 2158-2920 feet. Lower Miocene, Bascom No. 2 well, Mo-
bile, Ala., at 1241 feet, Chipola horizon.
alticallosa Dall, Trans. Wagner Inst. Sci., 3, p. 365, pl. 22, f.
Zon loo2e
Distribution.—Lower Miocene, Chipola marl, Northwest
Florida, and of the Bascom No. 2 well, Mobile, Ala. at 1241
feet, Chipola horizon.
Subgenus Crypionatica Dall
pusillaSay, Jour. AlN) S.) Phila’, 24) p.257)) 1822: shialeard
louse of Rep, Ex. Dochwpta2)) prssowers76., Wallaby
B 7p On UN EM oe ESAS On meet oO cl SLnipl ea uie El
Ann.) Rept.! lex. p:\ 240/92 anatear pen sy An Na Ss ieinilaas
55s D207159s) LO0Ss MOhnson Ocean Boosts: Socy ING ween zea.
TOS (TOLLS.
Distribution.—Maine to Florida Keys. Gulf coast: St.
Joseph’s Bay and Crooked Island, West Fla., Galveston,
Tex. Pleistocene, New Orleans artesian well of 1866, Lake.
Borgne borings, Knapp’s wells, Terrebonne Parish, La., No.
2y/at) 1150-1200, 1330-1470, 1500-1525 feet. | Pliocene
Caloosahatchie beds, and of the New Orleans Gymnasium
Club well at 1200 feet.
Genus POLINICES Montfort
Subgenus Euspira Agassiz
tenuis Recluz;) Jour. de! Conch) 1,\\p\)/388; pl: 12, £27, 1350;
143 MOLLUSCA OF THE GULF OF MEXICO 118
iyo Mane Sy pussy pl nav 254 88 aie stale Mie" Ce
EES PK ZO5 SO olay 27) Uk) Sy Neen DS a Logs
(All as Lunatia).
Distribution.—Gulf of Mexico, Yucatan Strait, 640 fms.
Also off Cape Florida, Cape Fear and Cuba, 84-640 fms.
Also Pacific Ocean, Valparaiso, Chile, Recluz’s type lo-
cality.
lestalea Watson, Jour. Linn. Soc., 15, p. 261, I880; Dall, Bull.
WER ZO pe Osh Sia Watson, | Challneot: | Gasth:.p.
AAO 27.7. Ss Wall Ball Gv Cea inow pai 215). 0.
(All as Lunatia).
Distribution.—Yucatan Strait, 640 fms. Also off Fernandi-
na, Fla., and Sombrero Isl,, 450 fms,
inngillae Wall Bally Me CZ, 9, p. 03) 18807) 18a pe 2o5) pl: 27,
fy) WO, SOs: LSDUlla ery LOS RON PIMC aL ialGyal soils aq ssauenuaa eexo).
(All as Lunatia).
Distribution-—Yucatan Strait, 640 fms. Also near Old
Providence, 382 fms. JL. radiata Watson, may be identical
with this species.
hemicrypta, Gabb; Jour, A. N. S. Phila (2).4)p.375, pl.
OVA 5.18003 Casi /Varzcea));) Dally Bullioqo Weis NE IM. p:
HOG, Oe Oy Es Cy anoint
eminwloiues Gabb,) jour, A. N.S.) Philas(2)18.p. 330,) pl:
Aas) t. 4, 1875.
Distribution.—Upper Miocene, Galveston well at 2465-2733
feet, and of the Jennings-Heywood No. 29 well, Jennings,
La., at 1960-1980 feet. Lower Miocene, Chipola marl, West
Fla. Oligocene, Tampa silex beds.
Subgenus NEVERITA Risso
duplicata Say, Jour..A. N. S. Phila., 2, p. 247, 1822, (as Natica);
Walls wlliy 7. We Se NOUN. ps) a54ae olsen i262 SO7 Clute
11g BULLETIN 38 I49
NUS OMS BE oinoa ors. (Ors) encima Jee. VAL IN, Sy Jelouilian se.
D759, 1003; Johnson,/Oce; (Pa Bost soc: Ny Ey 74 los.
IQI5.
canwpechiensis Recluz, fossata Gould, texastana Roemer.
Distribution.—Massachussetts to Mexico. Gulf coast: Cedar
Keys, and many other localities, West Fla., Horn Island,
Miss.; Point au Fer, Cameron, Chandeleurs, La.; Matagorda
Bay and Galveston, Tex. Pleistocene, Grand Chenier,
New Orleans well of 1856, New Orleans pumping station No.
7, Zigler, No. 15, Jennings, La., at 1350-1481 feet, Jennings-
Heywood No. 30 well at 1127-1169, 1169-1277 feet, Knapp’s
wells, Terrebonne Parish, No. 2, at 1150-1200, 1330-1375,
1443-1618, 1700-1839 feet. Pliocene Gymnasium Club well
at 1200 feet. Chesapeake Miocene, Maryland to Florida.
Upper Miocene, Galveston well at 2158-2920 feet. Miocene,
Crowley No. 25 well, Jennings, La., at 2468-2500 feet.
Subgenus Payraudeautia Bucq. Dautz. & Dollf.
nubifa Dall, Bull. M. C. Z., 18, p. 2094, 1889, (as everzia);
Dall and Simpson, Bull. U.S. Fish Com., 1. p. 439, pl. 58,
ENO TOO 'G:
Distribution.—Gulf of Mexico, Blake station 23, at 190 fms.
Also off Barbados, 140 fms, and Porto Rico.
Subgenus Mammilila Schumacher
uberina d’Orbigny, Moll. Cubana, 2, p. 31, pl. 17, f. I9, 1842,
(as Matica); Moerch, Malak. Blatt., 24, p. 60, 1877; Tryon,
Man., 8, pl. 16, f. 57, ’86; Dall and Simpson, Bull, U. S.
Bish) Com!) puidgou moor
Distribution.—Gulf of Mexico, west coast of Florida, 14-40
fms., living. Also off Sombrero 54-70 fms. and Porto
Rico.
factea Guilding, Tr. Linn. Soc., 17, p. 29, 733; (as Watcina);
iiyou, Man \'3: 0p: 49 pl 16h ie sooo, allman ai.
150
MOLLUSCA OF THE GULF OF MEXICO {20
U.S.N. M,, p. 156, ’89; Dall and Simpson, Bull. U. S. Fish
Comyn y P4126 IGOL
Distribution.—Florida Keys to Porto Rico and Brazil. Also
Texas region of the Gulf of Mexico. West Fla. (?).
brunnea Linck, Beschr. der Rostock Sammlung,2, p. 140, 1807;
Dat ull 27) WS. Ni Ma py 156, 80)
mamillarts Lamarck, Anim. s. Vert., 6, pt. 2, p. 197, ’22, (as
Wage). inyon, Mani, (8) p. 43, pli iSite 74860, Dall and
Simpson, Bull. U. S. Fish Com., I, p. 439, Igot.
Distribution.—Florida Keys to Trinidad Island and Texas
region of the Gulf of Mexico.
Genus SINUNM Bolten
prespectivum Say, Amer. Conch., p. 175, pl. 25, (as Szgaretus);
Holmes, Post-Pl. Foss. S. Car., p. 81, pl. 12, f. 16, ’60;Tryon,
Nine Se DNS 7 pleat Ol VO4u SO. Dalla iilema AOS.
NEEM ps r56. "So; \Vanatta, Pr. AY Ne Sa Pihilaynsise p. 758:
1903.
Distribution.—New York to Martinique. Gulf coast: Cal-
houn Co., Fla.; Chandeleurs, La.; Galveston, Mustang Isl.
and Pass Cabello, Tex. Pleistocene, New Orleans pumping
station No. 7, cited by Harris from Upper Miocene, Galves-
ton well at 2252-2600 feet.
maculatum Say, Amer. Conch., pacu7 On) plv25qmn@asiS7sarers) :
bison, Man.( 8 aps i50y pl. 24,0157 5S nOOmmOale Btnll a aia:
ese Ne ME pens On) SO:
Distribution.—Hatteras to Guadeloupe and West Florida,
shallow water. Closely related to martinzanum Phillipi,
GAbb: a) Beschrin,) py 144)) plait ha sa) 1 Say)
minor Dall, Bull. M. C. Z., 18, p. 297, ’89, (as Szgavetus); Bull.
Bry ORS NM pat 5Ou SO.
Distribution.—West Florida, Florida Keys and Sombrero
Isl., W. I. 54-84 fms.
12 BULLETIN 38 15x
Genus EUNATICINA Fischer
semisulcata Gray, Zool. Beechey’s Voyage, p. 137, (as Watica);
@iyon) Man! 8. piiz2) plight. ga 86 DalluBraile Mey Cia,
LS ON 200s) SOc Mir Helene snipe a SOnmozs
fordiana and fordit Simpson.
Distribution.—East Florida to Porto Rico. Gulf coast,
Sarasota Bay, West Fla., between tides.
Genus LAMELLARIA Montagu
rangi Bergh, Mer., p. 94, No. 8, 753; Dall, Bull. 37,U. S. N. M.,
DAU5O. 1 SO!
Distribution.—Texas region of the Gulf of Mexico. Ap-
parently never figured.
Genus MARSENINA Gray
amplaVerrill,: Pr U.S) NGM miara) 80-0. Conn:
NCA WATiS and Sei isi) pels usn plazh tae, (can So Dalle
Bully 37, U.S. NM, p.1156,)8o.),ohnson: Occ. Pay Bost:
SocwN He 7). TOO. KOs:
Distribution.—Maine to Sarasota Bay, West Fla.
Genus ACMAEA Eschscholtz
candeana d’Orbigny, In de la Sagra’s, Hist. Pol. y Nat. Isla de
Cuba; 2) p. 199) pl.25,) f) 1-3.) 1845,u(as-facela)): Dall Bull:
BG US.) ONE EME pr SON Sorin oll Mamie) Taosiae.
*91; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 440,
IQOl.
Distribution. —Florida Strait to Tobago, W. I. Also West
Florida and Texas.
punctulata Gmelin, Syst. Nat., 13, pp. 3705, 3717, 1792, (as
Ratela): PAlsbry,\eleny ota Mian nmieuiipAins7s) liDlis ature
1891; Dall and Simpson, Bull. U.S. Fish Com., 1, p. 440,
Igo!.
152 MOLLUSCA OF THE GULF OF MEXICO 122
Distribution.—Key West, Florida to Porto Rico, Guade-
loupe, and the Gulf of Mexico at Vera Cruz, Mex.
feucopleura Gmelin, Syst. Nat., 13, p. 3699, 1792, (as Patella);
Eusbryhnyony Man) 13.7.) 40) plats firo-2n on) Dall &
Simpson, Bull. U. S. Fish Com., 1, p. 440, 1901.
melanoleuca Dall, Bull. 37, U. S. N. M., p. 156, 1889. Not of
Gmelin.
Genus LEPETELLA Verrill
tubicola Verrill and Smith, Amer. Jour. Sci., (3), 20, p. 396,
pon Verll iraConn Acad) Arts andeseluisnqp. 534) ple
Some 2ou2qaue S25 Wall i Bull iM Ci Za arsip aan eile 2s
VOM Soe ubln a7 Uli Ss. Ne Mi ph ng eegaplen 25e)t..6,) 70%
Johnson, Occ. Pa. Bost. Soc. N. H., 7, p. 86, 1915.
Distribution.— Martha’s Vineyard, Mass., at 130-388 fms.,
to Gulf of Mexico between Mississippi delta and Cedar Keys,
at 324 fms.
Genus PHASIANELLA Lamarck
pulchelia C. B. Adams, Proc. Bost. Soc. Nat. Hist., 2, p. 7, 45,
(asi Yio) Aryon, Man) 10, p: (264.383 Walle Ball’) Me.
CIA re Os Sole oOs UBuUll raz.) sO Sa aiING Me Dae TSS.
1889.
brevis C. B. Adams. Not of d’Orbigny, 1842.
Distribution.—Cedar Keys, Fla., to St. Thomas, W. I. Plio-
cene of South Carolina.
Genus TURBO Linnzeus
castaneus Gmelin, Syst. Nat., p. 3595, 1792; Pilsbry, Man., 10,
pazos, pligsy fi88-8o) 7838. Dall) Bull. i27esUM SNE Mey p:
m5s- 780°) Dall) and Simpson! Bull Uy Ss-eishy)|Com:)\1,) p:
A441, 1901.
Distribution.—Cape Hatteras to Trinidad, W. I. Also West
Florida in shallow water. 25-295 fms, Pliocene, Caloosa-
hatchie and Shell Creek, Fla.
123 BULLETIN 38 5
a)
crenulatus Gmelin, Syst. Nat., p. 3575, 1788; Pilsbry, Man., 10,
DaZ2O2 pina si te on 7O8- Walle Bulla 7 Oe SlNeeN io: aa
(S08 (Gy VE ME Sia 2e DHi aS ouiio2™ ‘
Distribution.—Hatteras to Barbados. 2-30 fms. Also West
Florida at Crooked Island and St. Andrew’s Bay. Pliocene,
of the Caloosahatchie, probably a variety cf crenwlatus.
Genus LEPTGTHYRA (Carpenter) Pease
induta Watson, Jour. Linn. Soc., 14, p. 715, 1879 (as Turbo):
Challe Gastr.pi128) plo Oya t issn:
abide Mall, Bull. Me CUZ. 18 Hp es2nuple 23. tO, SO.
Distribution.—Hatteras to Martinique, 15-2805 fms. Yuca-
tan Strait, 640 fms.
Genus LIVONA Gray
pica Linnzeus, Syst. Nat., 10, p. 763, 1758, (as Zzrbo); Pilsbry,
Manta) pi 277.) olin pa 2a eo wiDaliane miles Morn Samet
M., p. 160, ’89; Dall and Simpson, Bull. U.S. Fish Com.,
LD: 443, 1901.
Distribution.—Charlotte Harbor, West Florida, to Colon.
37-160 fms.
Genus GAZA Watson
superba Dall, Bull. M. C. Z., 9, p. 49, ’81; 18, p. 354, pl. 22, f.
Aga, “895, Bulle g\ We SNe mea pA TOON Pls 22) iiayAletias
1889.
Distribution.—Gulf of Mexico between Mississippi delta and
Cedar Keys, 324 fms. Also Barbados.
fischeri Dall, Bull. Mus. Comp. Zool., Harv. Coll., 18, p. 355,
Pa 7et. 16.) Sor Bulls ah oMeNe NEEDS OO wOl. i2y7. mt.
1889.
Distribution.—Gulf of Mexico, west of Florida and St.
Lucia, W. 1., 423-426 fms.
154 MOLLUSCA OF THE GULF OF MExIco 124
(Callogaza) watsoni Dall, Bull. Mus. Comp. Zool., 9, p. 50, ’81;
Hoss FO DeNe24atenenTani plana anh. | iaNmcu plan 2duniini 2. 2a
18809.
Margarita jilosa Dall, Bull, M. C. Z., 9, p. 42, ’81.
Distribution.—YVucatan Strait, 640 fms. Also off Cuba and
Barbados.
Genus LIOTIA Gray
gemmia Tuomey and Holmes, Pleioc. Fos. S. Carolina, p. 118,
pli26,, tf. 4,156, (as 7vechus)-Dall and Simpson, Bull. U.S.
Fish Com., 1, p. 445, 1901.
tricarinata Stearns, Proc. Bost. Soc. Nat. Hist., p. 23, for ’72;
(siavchiiecionica) Wall, Bull. M. CC) Aiars8s i380, 789;
Bales 7 OW Su IN. Map. 166, 78:
Distribution.—Hatteras to Porto Rico. Also off Tampa,
West Fla., 15 fms. Pliocene, Caloosahatchie beds.
Hanoi DWally Bulls Mi CZ. 18, (p. 280, pl. 334k Bi SO:
Distribution-—Gulf of Mexico, Blake Station 2, at 805 fms.
Also off Carolina, Key West and Havana.
Genus CALLIOSTOMA Swainson
euglyptum A. Adams, Proc. Zool. Soc., p. 38,54, (as Zizyphinus) ;
Reeve, Moni Z7zy pi voll 2) t) 17.0085) Halles ule MeN Gs 174
ROM PA GOS TOO wellibiva Man. napa eyiAiuo le tegtohen 7a sp).
57, 1. 9, 89.
Distribution.—North Carolina to Florida, Texas and Vera
Cruz, Mexico. Erroneously referred by Reeve to Tasmania.
3-32 fms. Pliocene ancestor is C. philanthropus Conrad.
circumcinectum Dall, Bull. M. C. Z., 9, p. 44, ’81; 18, p. 364, pl.
22 eR Baio Billy 270 oe NGM peo Dl 22 uh. nae
3a, ’89.
Distribution.—Florida Keys, Antilles and Yucatan Strait.
640-805 fms.
125 BULLETIN 38 155
corbis Dall, Bull. Mus. Comp. Zool Harv. Coll., 18, p. 365, pl.
Bhai SOs MBM) 277 OSH iING VE De LOD) Une mune omen
18809.
Distribution.—Gulf of Mexico, west of Florida, to Jamaica.
220-450 fms.
tiara Watson, Jour. Linn. Soc. 14, p. 696, ’79; Rept. Chall.
Caste) ps 60,) plavOmt. vans suas Pivockiasyie Daly buleyarzs
We Sane ol nO2ti SO:
Distribution.—Gulf of Mexico, Texas region, to Dominica,
W.I1. 220-780 fms.
roseolum Dall, Bull. Mus. Comp. Zool, Harv. Coll., 9, p. 45, ’81;
Ices OA CLO 8) ea by ote (OMI ovel. \” sOyae BION. ALON use ING Me INE 70).
LOZ lI 2A.t.) GunGa aoe
Distribution.—North Carolina to Yucatan. Also Florida
Straits and Havana. 21-200 fms.
pulcherum C. B. Adams, Contributions to Conch., No. 5, p. 69,
n50;)\(as| Lvochus);\(Pilsbry,) Man) 11/4375) 89. (Malls Bull:
BT UMS Na Mien sp MOMS O!
Distribution.—Hatteras to St. Thomas, W. I. Texas region
Gulf of Mexico. 15-63 fms. ‘Type locality, Jamaica.
veliei Pilsbry, The Nautilus, p. 128, 1900.
Distribution.—Caxambas Pass, Southwest Florida.
(Euirechus) jujubinum Gmelin, Syst. Nat., p. 3570, 1792, (as
ivockus) ;) Pilsbry;,)Man.\\ 0, psi4o4. ply 40... 16) 7899) Dall
Bull 37, WU. S2Ne Mp a62 080; all and Simpson Bile
WS) ish, Com), psjaaa, (toon,
Distribution.—Hatteras to Cartagena, Colombia. Gulf coast,
West Florida, Texas and Yucatan. Pliocene, Caloosahatchie
beds, Fla.
(Eutrochus) jujubinum var. perspectivum Koch, Philippi, Abbild. u.
150 MOLLUSCA OF THE GULF OF MEXICO 126
BESeAh yl, Pt.)2), i Lrocwes, pl. Tt. 15))) 1843+) Eilsbry,, Man,
PP 4O5,) Dia OO 135,130,189)
Up men sts \COntAC, WatiAe) ING os) bila: ape Zon pl 2 tua.
Morn(asiiivockus)) Wally Bulliz7 Us Say Me ior 16260:
Distribution.—Hatteras to Honduras. Also West Florida,
shallow water.
(Eutrochus) jujubinum var. rawsoni Dall, Bull. M. C. Z., 18, p.
369, °89; Dall and Simpson, Bull. U. S. Fish Com., 1, p.
444, 1gol.
Distribution.—Cedar Keys, Fla. and the Antilles.
(Eutrochus) yucateeanum Dall, Bull. M.C.Z., 9, p. 47, ’81; 18,
DapszOy Pl 24ef.140 gas So.
Distribution.—Yucatan Strait, 640 fms. Also off Carolina
coast, 15-32 fms.
(Dentisty!a) asperrimum Dall, Bull. Mus. Comp. Zool., 9, p. 40,
78m, (as| Margarita): 18, p. 373, °89; Pilsbry, Man, 11, p.
Misi o Om Oa tll 27) UL) Se Niy MAC May mO2 nis:
Distribution.—Hatteras to Barbados. Also West Florida.
100-177 fms.
Genus SOLARIELLA S. Wood
amabilis Jeffreys, British Conch. 3, p. 300; 5, pl. 61, f. 6, (as
Trochus); Dall, Bull. M. C. Z., 18, p, 378, ’89; Pilsbry, Man.,
EDs Lan pl s7a he 52.) (Soy
Distribution.—Yucatan Strait, 640 fms. Also Antillean and
European (North Sea off Shetland Isls, 85-95 fms).
scabriuscula Dall, Bull. M.C. Z., 9, p. 41, ’8c, (as Wargarita);
HOMO) Pi 2u i. TO loads So.) Dilly a7 ny sn Nie Mie 5 1.
HOMME Dl 21h Le Om TOds, | 8On
Distribution.—Southern part of the Guli of Mexico, 539
fms., bottom temperature 39° F. Also off Cuba.
127
BULLETIN 38 : I57
aeglees Watson, Jour. Linn. Soc., 14, p. 704, ’79; Chall. Rept.
Gastr., p. 81, pl. 5, f. 10, 785; (as Margarita); Dall, Bull.
VE CZ Ok! AO. Ole Ss DE RYIOn OO cial Sbiayaa Via tle 1s
Bieby Holy nGfoyw peg csiay 10O).! | SX0),
Distribution.—Gulf of Mexico, 287-888 fms. Also off St.
Vincent, W. I.
aeglees var. clavata Watson, Jour. Linn. Soc. London., 14, p.
705, Chall. Rept. Gastr., p. 82, pl. 5, f. 8, (as Margarita);
Dali) Bulll Me CZ 85 pi280) 78omelsbry Mam. pike ps
ZOOM DL OO} fs OSs OO oO!
Distribution.—Yucatan Strait, 640 fms.; Culebra Isl., W. I.
350 fms.; Pernambuco, Brazil, 675 fms. Dall and Pilsbry
regard this as a variety of aeglees.
lubrica Dall, Bull. M. C. Z., 9, p. 44, ’81, (as Wargarita); 18,
D282. pl. 20h) O- 1789. bilsbrmyaiMiane atti. Qo4u ls isin ae
250/20. 8O- Wa lly Bally, sac nlel sy INI IVs TO: 1 s@ Aly) if0 en aeeaates
9, 9a, 89.
Distribution.—Cedar Keys, West Florida, to St. Lucia, W.
I. 116-805 fms.
lubrica var. iridea Dall, Bull. M. C. Z., 18, p. 382, ’89; Pilsbry,
Mani rn) pi22Q so. Dall mle 2 Tanase ON. Verano
1889.
Distribution.—Cape Florida, Gulf of Mexico, West Florida
region. 193 fms.
lissoconcha Dall, Bull. M. C. Z., 9, p. 41, ’81, (as Margarita);
WOR Remo, Aeiats an tekel. “oye JEU, OL Sal INE Ao}, 641, fol
ZINE.) 8.82, OO)
Distribution.—Gulf of Mexico, Lat 28° N., Lon. 88° W.,
at 331 fms.
Genus BASILISSA Watson
alta Watson, Jour. Linn. Soc. London, 14, p. 597, 79; Rept.
Chall) Gastrs) p.mooliplai7. i SieoMbilsb hy.) Mam ent ip:
158 MOLLUSCA OF THE GULF OF MEXICO 128
ATOMDInE SON ius OO sO allo Dalia Nisa Men Del TOA
1880.
Distribution.—-Gulf of Mexico, West of Cedar Keys; the
Antilles and Brazil. 339-1019 fms.
alta var.delicatulaDall, Bull. M. C. Z., 9, p. 48, 81, (as Seguen-
Ea i8.py 384) plaz2, t./2.0 80; Pilsbrayay Mantis, \p. 421
Desa tai 8O Dall, Ballin: Us SmiNtuMietep sn 64) ple
Domi 24 Zann So:
Distribution.—Gulf of Mexico to Tobago, W. I. 805 fms.
superba Watson, Jour. Linn. Soc. London, 14, p. 598, ’79; Rept.
ChalleiGartrp. tom pl: 7, 4h. 10,,/86s)etlsbrye Miami.) Ta |). ps
AQ ple Bout: O77) 789; Dall, Bulls a7 Us Sunnie. pi 164s
1889.
Distribution.—Cape York, Australia, type locality. Also
Gulf of Mexico west of Florida. 400-1400 fms.
(Ancistrobasis) costulata Watson, Jour. Linn. Soc. London, 14, p.
Goor Sou Rept Chall: Gastr. p:) 103). ple7eiia im) S011 Dalle
Bull MC) Z., 18, p:' 384, pl. 23, f.4; 4as8o; Pilsbry,; Man,
IDE AZO pla Bon a so oO: Dall Bale anWe i SeNi Mess
HAL, Ole Denes vale abaya tsKos
Distribution.—Georgia to Culebra, W. I.
costulata var. depressa Dall,Bull. M. C. Zool. Harv. Coll., 18, p-
3845180) bllsbiry,) Man. Tr p: 428. plu, Pian soon Wall:
Bulle Ue SNe Mos 164.) pli 2ecit4n an oor
Distribution.—Yucatan Strait, 640 fms. Also Antillean.
Genus UMBONIUM Link
bairdi Dall, Bull. Mus. Comp. Zool., Harv. Coll., p. 359, pl. 21,
Paonia) So. eilsbrys |) Mane) “ni pela ST ple sO@d tans. (6.
1889.
Distribution.— Yucatan Strait, 640 fms. Also Florida reefs,
200 fms.
12G BULLETIN 38 I5@
Genus COCHLIOLEPIS Stimpson
parasitica Stimpson, Proc. Bost. Soc. Nat. Hist., 6, p, 308, and
he 53./ Dall, BullyiNiiG. (ZeamS)\ 91260.) (So Bulla aa).
Sa aNG Me. p! 162) (Sorina Wem Schis 20 ans amon
Distribution.—Charleston Harbor, S. Car. to Florida Keys
and Tampa, West Florida. At Charleston this species lives
under the scales of a large annelid. Pliocene, Caloosahatchie
beds, Fla., rare.
striata Stimpson, MS.; Dall, Bull. M. C. Z., 18, p. 360, 1889.
Ballea7.0U. 6 S..,Ne MG) pinG2n) (8oruehr. Was ner mllmsts
Sei 3) pL 470) ple te TON O2h acot moun. Matchelt
SOC! 1p. 138) Keb. 192m:
Distribution.—Beaufort, N. Car. (Jacot); Egmont Key,
near Tampa, West Fla. (Colonel Jewett). Upper Miocene
of North Carolina.
Genus ADEORBIS S. V. Wood
adamsi Fischer, Journ. de Conch., p. 173, pl. 1o, f. 11, 1875;
Dal eer O.7S. NM VIMEO penser Sa meiisiiy alma Cotte
Acad. Sci., 10, p. 104, 1899-1900.
Distribution.—Cited by Dr. Dall from Cedar Keys, West
Fla.
Genus VITRINELLA C. B. Adams
mooreana Vanatta, Proc. Acad. Nat. Sci. Phila., 55, p. 758,
1903.
Distribution.—Crooked Island, Calhoun Co., Fla.
hemphilli Vanatta, Proc. Acad. Nat. Sci. Phila., p. 24, pl. 2, f.
iy Zip, Meas
Distribution. —Cedar Keys, Florida.
interrupta C. B. Adams, Monograph of Vitrinella, p. 6, 1850;
jot: de ,Conch:.6.\ps076,-DalluMbulla 7 MUGS: INGE Mer:
166, 1599;) Bush) itr Couns wNcadaisciny (10), ppsjLoom Loz.
*99-1900.
e]
160 MoLLUSCA OF THE GULF OF MEXICO 13
Distribution.—Tampa, West Fla., in shallow water, to
Jamaica.
Genus CIRCULUS Jeffreys
liratus Verrill, Tr. Conn. Acad. Sci., 5, p. 529, ’82, (Omalaxis);
6, Pp. 452, °85, (Skexea); Bush, of. ctt., 6, p. 464, °85; Bull.
VC eZ nz) Da 240. Oly MIT hal LO Oech GomnaniAcads
SCH TOM Prats.) pl 22tui7alnT2 Tab YOO 1oOO!
Adeorbis supranttidus var. orbignyt Dall, Bull. M. C. Z., 18, p.
POMS Os (AH partis lr. Wel Te Siig pmai24Anog Johnson,
WceyiPa) Bost: Soc. Ni), 7) pe rzommons) Not ov bze 722
nischer, Journ de Conch. 6, pp.) 173) 286), 157.
Distribution.—New England to Hatteras and West Florida.
Lower Miocene, Chipola beds, West Fla.
Note.—The true Czrculus orbignyi Fisher is West Indian and
very minute, only 144 mm. in diameter,while “vatus is-2%
mim. in diameter.
trilix Bush, Rept. U. S. Fish Com., for 1883, p. 584, 1885, (as
SCLC), lie ConmexAcad. | (Sei) 10% paaa7pli22 tT. 6. °1o,
10a, 12, a-g, pl. 23, f. 10, 15, 1899-1900.
madeonous supyanczaus Wall, Bull. Mii Cay Ze n8. p.278)0 89°
BUS NW TSO. lanai Rema cra oan bala
part). Not supranitidus Wood, Cat. 1842, Crag. Moll., p.
LA 7 MPL LS tsa sa-De kos.
Distribution.—Massachussetts to Hatteras and to Cedar
Keys, West Fla. 15-25 fms. Pleistocene, Knapp’s No. 2
well, Terrebonne Parish, La., at 1519-1632 feet.
Genus LISSGSPIRA Bush
proxima Tryon, Manual Conch., 10, p. 98, pl. 33, f. 4, 1888,
(Gydostrema); Bush, Tr. Conn. Acad. Sei:, 10, p. 130, f. 4,
P22.) 20 T899-1G00,
rugulosum Verrill, Tr. Conn. Acad.,5, p. 533,’82,(Cyclostrema).
Not of G. O. Sars, 1878.
131 BULLETIN 38 16X
ayine Nermill) ir: Conn Acad @nGuipy moo. pluisen 1. es. e4e (as
Cyclostrema). Not of Jeffreys, 1883.
trochoides Dall, Bull. M. C. Z., 18, p. 393, 1889; Dall, Bull. 37,
U.S. N. M., p. 166, ’89, (as Cyclostrema). Not trochoides
(Jeffreys) Sars which is a variety of pettersen7 Friele.
Distribution.—Lat. 41° N., Lon. 65° W. to Lat. 35° N.,
Lon. 74° W. (Bush); West Florida (Dall).
Genus CHORISTELLA Bush
pompholyx Dall, Bull. M. C. Z., 18, p. 394, pl 28, f. 9, ’89, (as
Gydlosizenca) Balla Wem Sui NeeVieap at On plat 2 Saute mon
’805 Bush, ir. Conn: Acad!) Sci-. 10; pp.) 99; | 1401) 1890-
1900.
Distribution.—Blake Station 2, Gulf of Mexico, type locali-
ty. Also Fernandina, Florida, to Cuba. 294-805 fms.
Note.—This species is referred tentatively to Choristella as
Miss Bush suggested that it might prove referable to this
genus. ’
Genus EPICYN!A Moerch
multicarinata Stimpson, MS., Dall, Bull. M. C. Z., 18, pp. 273,
Z02,'/807) Bull27, WS) Nea pr 166)786.) (as, ze7z7ella)\.
Bush, Tr. Conn. Acad., Io, p, 112, ’99-1900.
Distribution.— Hatteras to West Florida. Pliocene, Caloosa-
hatchie beds,” Fla. Upper Miocene, Duplin Co., N. Car.
Perhaps identical with the Avrchitectonica gemma of Holmes
listed by Hilgard from the Lake Borgne borings, La.
Genus ETHALIA A. Adams
The following species are of doubtful generic position. ‘They
were excluded from /¢halia by Pilsbry (Man. Conch., 1,
Pp. 457) when he restricted that genus and gave Ethalia
guamensis Quoy and Gaimard, as the genotype. Dr.
Dall has referred me to Miss Bush’s revision (Trans. Conn.
Acad. Sci., 10, p. 116, 89-1900). Apparently she tentative-
ly retains these three species in Hzhalia.
162 MOLLUSCA OF THE GuLF oF MExIco 132
suppressa Dall, Bull. Mus. Comp. Zool., 18, p. 362, ’89; Bull. 37,
Wee NE Men pu) NOON So)
Distribution.—Goodland Point, West Florida. Collected by
Hemphill.
reclusa Dall, Bull. Mus. Comp. Zool., 18, p. 361, pl. 28, f. 7, 8,
"SOs IEEE Soe OLAS INU [Bae omnadeloy, “Skok
Distribution.—Yucatan Strait, 640 fms. North Carolina
coast, 12-65 fms.
Salida Wall) Bull M.C.Z,, Harv. (Coll 18) plist. 3) 5, 780;
MSU aoe, INE Me pr LOO) pl N 28 uikal ay GanoC)
Distribution.—West Florida and Cuba, 310 fms.
Genus TEINGSTOMA A. Adams
cryptospira Verrill, Trans. Connecticut Acad. Sci. and Arts.,6, p.
PAPE OAL an (as, Acozella)) Wall Bull. 277-9 W4y SayNE Niet. 160;
RSOq EUS lor, Con) Acad: - Sci... 10) pyrrole EN 241 F8Oo-
LooOs Dall Ir Wagner Insts Sei., 33) ps) 4145.92)
Distribution.—Hatteras to Florida. Gulf coast at Crooked
Island, Calhoun Co., Fla. 30-150 fms. Pleistocene of
North Creek, Fla. Miss Bush regards this species as atrue
Teinostoma.
Genus NERITA (Linneeus) Lamarck
pelorenta Linnzeus, Syst. Nat., 10, p. 778, 1758; Reeve, Conch.
NCOs Oy ls Be way Sow dang Miyoyo Wiewowe sO, jos. Bala noyll
Ai 75-7 7cooo Mall Bull=37.) Us Sa Nt Me Koos 1£8So:
Dall andy Simpsons Bull U2 (Si hishe) Commie. 445)
IQOL.
Distribution.—Florida, St. Vincent and Texas region of the
Gulf of Mexico.
tessellata Gmelin, Syst. Nat., p. 3685, 1792; Reeve, Conch,
icon oy pl. ON. 7435) 494,) 6855-) dinyon wane moO py 24, ipl:
133 BULLETIN 38 163
Aer! Seen Dali ull eal SiN INO Goneon Dalle
and Simpson, Bull. U. S. Fish Com., 1, p. 446, 1901,
Distribution.—East Florida to Colon. Gulf coast, West
Florida and Texas, shallow water.
versicolor Lamarck, Anim. s. Vert., 6, pt. 2, p. 193, 1822; Reeve,
Conch: Icons) o. ole 12) fase. a dakOs Somali Etiam.
Uiisu NN. Mes) p: 166.) 780: Dall andi Simpson yBulliue S:
ish) Com: (i) p..446.) 190r
Distribution.—Florida Keys and West Florida to Colon, the
Antilles and Bermuda.
Genus NERITiNA Lamarck
reclivata Say, Jour. Acad. Nat. Sci. Phila., 2, p. 257, 1822, (as
Npeodoxus)\: Reeve. Conchiainon ey 34) a-b)). 55> Wallpean
WSN SUMS. pul2sonpl iim tase nS se iyon Mane srounne
39, pl. 12, f. 25-27, ’88; Singley, Fourth Ann. Rept. Texas
Geol. Surv., p. 342, 1892; Mitchell, List. Texas Shells,
Pp. 7-
Distribution.—St. Augustine to Martinique. Gulf coast:
Mouth of Manatee River, Tampa, Cedar Keys and St.Mark’s,
Fla., Point au Fer and Chandeleurs, La.; Port Lavaca and
Rockport, Tex. Pleistocene, Knapp’s No. 3 well, Terre-
bonne Parish, La. from 1-700 feet.
Note.—The variety palmae Dall is found at Palma Sola,
Texas.
virginea Linnzeus, Syst. Nat.. ed. X, p. 778, 1758, (as Nerita);
Reeve) Conch icon o.ny plea 22a Ge ot S55 uaa
OAS Ne Mays pil 25oM/8i5:) myonme\Vlan..) TO.) Dp: 2Qunplagia.
Hagt-25. 37-42 Dall Maulana 7a eos ON NIG.) py OSes SO.
Singley, Fourth Ann, Rept. Geol. Surv. Texas, p. 342,
1892.
Distribution.—Florida to Brazil. Gulf coast: Various local-
ities of West Florida; Corpus Christi and Rockport, Tex.
x64 MoLLuscA OF THE GULF OF MEXICO 134
pupa Linnzeus, Syst. Nat., ed. X, p. 378, 1758; Tryon, Man., Io,
Os ABS Ole WAL A ore ate ODN, JEENULS Gi OWS: SS) INi2 ME yo),
168, ’89.
Distribution.—Charlotte Harbor, West Florida to Jamaica.
Shallow water.
viridis Linnzeus, Syst. Nat., ed. X, p. 778, 1758; Tryon, Man.,
MOMS ts plu 18) £88) 1888; Dall) Bully Wu Su Ni Vi.
Dylos. 89.
Distribution.—Florida Keys to Barbados, W. 1. Also Texas
region of the Gulf of Mexico.
showalteri Lea, Pr. Acad. NatSci. Phila., p. 55, 1861; Tryon,
(BOS TKOs FOI Get ONAN Aben Snes iacKs en IOBUN IS ybIUG) ceioie Ola Sip
ING Me ps) K68;) .6O:
Distribution.—Alabama and West Florida, Fluviatile.
sparsilineata Dall, Proc, U.S. N. M., 46, p. 236, 1914.
WNenintan spsiindet,, Aldrich, Nantilis) 24))/pi nin ps 131,
IN@s. Jo UGUIE BO folie seme MayM toner),
Distribution.—Upper Miocene or Pliocene. Well near Al-
exandria, La., at 49 feet; Pine Prairie La., well at 1540 ft.
and well % mile S. E. of Pine Praire depot, in dump. Also
Satilla River, Ga., and Burkeville, Texas. One of the most
widespread and characteristic species of the brackish water
formation, extending from Georgia through Louisiana to
Texas.
Genus PLEUROTOMARIA Sowerby
(Petrotrochus) quoyana Fischer and Bernardi, Journ. de Conch.,
Eee HOS. Dla Sut. 1856: Dall Balin Man Z eons 7.o)
gon Crosse journ., de Conch), 22." pa ian s2snMall Bull:
NICE ZEaers) De O7h pli 2ou ta Te Wu tMn Tal Me CH le nae an
Poon mullet 27 UL SuiNG Me ps 168) samenpls wand) fosi
1889.
Distribution.— Yucatan, near Arrowsmith Bank, at 130 fms.
135 BULLETIN 38 265
Also off Barbados, 73 fms., and Marie-Galante Island, near
Guadeloupe.
Genus FISSURELLA Bruguiere
(Cremides) barbadensis Gmelin, Syst. Nat., p. 3729, 1792, (as
iRacewa)i Dall Bull 27. Ui Sy iNWuM wpe 7O.1.8o) nbd story.
Man., 12, p. 164, pl. 37, f. 40-49, ’90; Dall and Simpson,
Ball U2 5) Fish) Come (ie sp) Aa su oom
Genus LUCAPINA Gray
suffusa Reeve, Conch, Icon., in errata, 1851, (as Fissurelia); Dall,
icrans,) Wiaener Insti Set 2) py Azdpleizan ie (22)402k
hondurasensis Reeve, Conch. Icon., f. 70, 1851.
cancellata Sowerby, in part, Thesaurus Conch., p. 200, pl. 8,
(248) f187 only. 362.) Wall Mabini Wan S a4 oN MEN iMG Os
189); bilsbiy, Manin ta. De 2OOMrDlao static pla Mstemsor
’90; Dall and Simpson, Bull. U.S. Fish Com., 1, p. 449,
Igol.
Distribution.—Marco, West Florida to Vera Cruz, Mexico.
Also Antilles and South America to the Island of Fernando
Noronha, Brazil.
Note.—As there has been some question whether the name
suffusa Reeve or cancellata Sowerby should be applied to
this species, Dr. Dall’s opinion was asked. He replied, June,
1921, ‘‘I find nothing to change inthe synonymy of ZL.
suffusa as given in the Wagner memoir.’’
adspersa Philippi, Abbild. u. Beschr., 2, p. 34, pl. 1, f. 3, 1845;
(as) -Fissuvella) Rilsbiny, iMate fe 2h apie el OO). ls O2 eno) wie7s
’°90; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 449,
IgOl.
fasciata Pfeiffer. Fissurellidea fasciata (Pfeiffer) Dall, Bull.
VRS SME p72 iSO),
Distribution.—West Florida region of the Gulf of Mexico to
Cubavandist + Crom Vie a
166 MOLLUSCA OF THE GULF OF MEXICO 136
Genus FISSURIDEA Swainson
alternata Say, Jour. Acad. Nat. Sci. Phila., 2. p. 281, 1822, (as
iassuneiia) Pilsbry,) lane. 12) ps 211) pleg7.\f50-53, pl: On,
Penis) oo7) Dall and | Simpson, Bully Ues:) kish, Com,” 1;
Pp. 450, Igor.
Distribution.—Chesapeake Bay through the Antilles to
Trinidad Isl. and Fernando Noronha, Brazil. Gulf coast:
Cedar Keys, Fla.; Galveston, Corpus Christi, Texas; Vera
Cruz and Progreso, Mexico. Also south of Nicaragua. I-50
fms. Pliocene of South Carolina.
eayenensis Lamarck, Anim. s. Vert., 6, pt. 2, p. 12, 1822; Reeve,
Conch olcon? 7h) 232) Dall, Bull} a7 Uaioe NEM | p12 70)
Ow bilsbry.) Man re) ps 212, plzz 1. above 60, 1890.
Distribution.—Cedar Keys, Fla. to St. Lucia, W. I. Placed
by Pilsbry in synonymy of a/fernata, but is more elongated
and with straighter sides than that species. Perhaps to be
classed as a variety.
Genus PUNCTURELLA Lowe
iriolumMe Wall Bulle MC) Z.\ 9, p.176, Si; 18ip-w4og, pls 26, f.
8, 8b, ’89; Bull. 37, U. S. M. N., p. 168, pl. 26, f. 8, 8b,
1889.
Distribution. —Yucatan Strait, 640 fms.
watsoni Dall, Bull. M. C. Z., 18, p. 403, ’89; Bull. 37, U. S.
ING Me p68.) 780:
Distribution.—Off Yucatan, at 200 fms. Also off Cuba and
Barbados.
cincularis;) Dall) Bull, Mi: (@. Z:> 9, pi75; *8n;/ 18, p. 403, pl. 26,
Pee SO.) Bulls 7a Mey Sa Nee Mesmp ea stOSnapl 26), fai.
7b, ’89.
Distribution.—Florida Strait and the West Indies.
1127) BULLETIN 38 167
(Fissurisepta) triangulata Dall, Bull. M. C. Z., 18, p. 404, ’89;
Bolle) S2 ING ais ipscw7onn So:
rostrata Watson, Chall.Gastr., p. 48, pl, 4, f. 10, 7&5. Not of
Seguenza.
Distribution.—Yucatan coast, 200 fms. Also off Fernan-
dina, Fla., and Culebra Isl., W. I.
(Cranopsis) asturiana Fischer, Journ. de Conch., 30, p, 51, 1882,
(as Rimula); Watson, Chall. Gastr., p. 45, pl. 4, f. 4, ’85;
Dall Bulls IM Cai Zein S) (psi One Son
Distribution.—Yucatan Strait, 640 fms. Also off Havana,
Martinique, and Cape Florida, and in the Gulf of Gascony
at 2018 fms.
Genu EMARGINULA Lamarck
(Rimula) frenulata Dall, Bull. M. C. Z., 18, p. 406, pl. 28, f. 4,
OOM BU e 7 OES INE NED ene Ou 2 Shake Aloe
Distribution.—West Florida, Florida Keys and Hatteras, 6-
52 fms.
(Emarginula) compressa Cantraine, Bull. Acad. Roy. Bruxelles,
9, p. 2. 1835; Jeffreys, Pr. Zool. Soc., p. 679, ’83;Dall, Bull.
Me CuOZ TS pv AOAt esos
Distribution.—Yucatan Strait, 640 fms. Also off Havana,
Barbados, and coast of Portugal. ‘Tertiary of Sicily.
x68 MOLLUSCA OF THE GULF OF MEXICO 138
GNSS) NINA ed ool NI Le RR
ORDER POLYPLACOPHORA
Genus LEPIDOPLEURUS Risso
pergranatus Dall, Bull. M.C. Z., 18, p. 414, ’89, (as Leptochi-
ton): Bull. 37, U:'S. N. M.,. p. 172, 89; Dall and Simpson,
Balle US. hish) Come: 19.451, ply yssitavi aac. Loon:
Distribution.—Gulf of Mexico, western Florida region,
Porto Rico and Dominica, 114-1181 fms.
Genus CHAETGPLEURA Shuttleworth
apiculata Say, Amer. Conch., app., pt. 7, 1830, (as Chzton);
Dallyaaly 27s se NEM per 72. aienih. TO. T8So%
Pilsbryg) Mian’)! 14, p. 35, pli’ 13, tf) 75-7on 1892;) Johnson,
Occmeanibost.. soc. Ni HH: (7, 1p. 8) Toms:
Distribution.—Massachussetts to Haiti, Gulf coast, various
localities of northwestern Florida and Chandeleurs, La. 0-30
fms.
Genus ISCHNOCHITON Gray
limaciformis Sowerby, Proc. Zool. Soc., p. 26, ’32, (as Chiton);
DalkeBully Mey CayZe 18. p. 45.) So svedlsbrypyMans ogy p:
57, pl. 16, f. 9-16, ’92; Dall and Simpson, Bull. U. S. Fish
Com p4 4.52.) LOOT
Distribution.—Gulf of Mexico, southeastern part, Lat. 24°
N., Lon. 83° W., at 37 fms. Also Key West and the An-
tilles, Central America and Peru. Under stones at low
water.
purpurascens C. B. Adams, Proc. Bost. Soc. Nat. Hist., p. 9,
u845. (as) Chzzon))) Dall) Bull..27. Wo Se Ne MeL) pi 172.) So:
Edlsbiny. Manian pa SSa plait. 2oN 2annnSQ 2 ial arid
Simpson, Bull. U.S. Fish Com., 1, p. 452, 1got.
Distribution.—Texas region of the Gulf of Mexico, Florida
Keys, Jamaica and Porto Rico.
139 BULLETIN 38 ; 16g
papillosus C. B. Adams, Proc. Bost. Soc. Nat. Hist., p. 9, 1845;
Dall yBall 27.00 SeiNeMMelnm. 7172) 7 Sonia bia eNleian yaeAy
Pp. 114) pl. 21-8, 40) 401892. Dall andi simpson, Bulli Weise
Fish) Com),/1, ip: 452, 190:
Distribution.—Tampa, West Florida, to St. Thomas,
Wevele
Genus CHITON Linnzeus
fuberculatus Linnzeus, Syst. Nat., ed. X, p. 667, 1758; Pilsbry,
Man., 14, p. 153, pl. 33, f. 58-60, 1892; Dall and Simpson,
Bull. U.S. Fish Com., 1, p. 453, Igor.
Distribution.—Texas and Florida to Trinidad and Ber-
muda.
marmoratus Gmelin, Syst. Nat., p. 3205, 1792; Dall, Bull. 37,
WOE Se ING Mae) ips 2h So nba sbiave Mamie y 4c 0.1 15S, plana
f. 72-76, 1892; Dall and Simpson, Bull. U. S. Fish Com., 1,
Pp. 454, 190t.
Distribution.—Texas, and the Antilles to Barbados. South
to Cartagena, Colombia.
Genus ACANTHOPLEURA Guilding
granulata Gmelin, Syst. Nat., p, 3205, 1792, (as Chzton); Pilsbry,
Man., 14, p. 227, pl. 50, 1892; Dall and Simpson, Bull. U.
S. Fish Com., I, p. 454, 1901.
pica (Gmeling Dall) YBull 27 sOm Sa ONG Ne. Pe) lat isos
Distribution.—Charlotte Harbor, West Fla., to Trinidad,
Bermuda and Bahamas.
Genus ACANTHOCHITES Risso
spiculosus Reeve, Conch. Icon., pl. 9, f. 47, 1847; Dall, Bull. 37,
We SING ME pn 740) OO Med Sbiny NAT NS. | Dah2 nolan
f. 60-62, 1893; Dall and Simpson, Bull. U.S. Fish Com.,
Ae ON USES Wa Keyos (3
170 MOLLUSCA OF THE GULF OF MEXICO 140
Distribution.—Cedar Keys, West Florida, to Barbados.
pygmaeus Pilsbry, Tryon’s Manual Conchology, 15, p. 23, pl 13,
PF OK 159s SOs.
Distribution.—Cedar Keys and Key West, Fla.
I4I BULLETIN 38 E71
GHASS CE RHA @R@ WA
ORDER DIBRANCHIATA
Genus ARGONAUTA Linnceus
argo var. americana Dall, Bull. 37, U. S. N. M., pp, £74, 200,
plas) t. 1 a-by pleat Tazbiplv on Mit 2188.) [ounsen,
@Occei}Pa\ Bost.’ Soc: Nat (Eista7hipai2235)LOls:
Distribution.— Massachussetts north to 43° N. Lat., and
south to the Antilles and doubtfully to Brazil. Gulf of
Mexico, West Florida region.
Note.—The typical Argonauta argo Linnzeus, Syst. Nat.,
ed. X, p. 708, 1758, is Mediterannean. It has two or three
times as many radial folds and carinal nodules as the varie-
ty americana.
Genus SPIRULA Lamarck
australis Lamarck, Anim.s. Vert. 7, p. 600, 1822; Encycl. Me’th.,
pliA6s5, tS, a,b: Walls Science) mi sins, spp. 243-2454 nego.
Dall/and Simpson, (Bull WS; bush Come) 1p.) 359, ilies,
LAs OOT:
spirula Linneeus, in part.
perone (lamarck) Dall, Bul a7 UES INY MCn loin nels
CSA TSO:
Distribution.—Pelagic. West Indies and the Gulf of Mex-
ico, shells being washed ashore along West Florida coast
and at Cameron, La., Galveston and Corpus Christi, Texas.
A specimen with all the soft parts was also taken from the
172
MOLLUSCA OF THE GULF OF MExICO 142
mouth of a fish trawled by the Albatross between the Mis-
sissippi delta and Cedar Keys, at 324 fms. The Spzvula had
just been seized alive by the fish and the soft parts, pre-
viously imperfectly known, were complete. See Science,
13896.
ivnere are! several (tors) of |) Spezza | Siiispzrula. cot
Linnzeus was a composite, according to Dr. Dall, and since
the species cannot be differentiated by the shells alone no
one can tell exactly which species Linneeus had although he
gave America as the habitat. Huxley and Pelseneer (Chal-
lenger Rept.) fixed the name australis Lamarck on the An-
tillean species.
Note.—Arvgonauta and Spirula are the only shell-bearing
Cephalopods reported from the Gulf of Mexico. Of other
forms, the Brazilian, Loligo brevis Blainville is cited from
Cameron and elsewhere on the Louisiana coast, and Z.
hemtptera Wowell from Florida and the Gulf of Mexico.
Ma ee og Seer
“A Bis te %
me
be
ss
pera
BULLETINS
OF
AMERICAN PALEONTOLOGY
Vol. 9
The Miocene of Northern Costa Rica
WitTH Novtss On Irs GENERAL STRATIGRAPHIC RELATIONS
BY
A. A. OLSSON
Harris Co.
Cornell University, Ithaca, N. Y.
UA Savane
OEDICAT =D
WOSF A=
MUSEU IM! Cm GOSsra RICA
y
hg
ay
me =n V1 OS En INE @) a) IN @) eal el ES ESN
G@QSWA RiGA
WITH NOTES ON ITS GENERAL STRATIGRAPHIC
ION.
JW
RELATIONS
BY
A. A. OLSSON
PRELIMINARY REMARKS
GENERAL STRATIGRAPHY OF NORTHERN COSTA RICA
THE CosTA RICAN MIOCENE
A. THE USCARI STAGE
B. THE GATUN STAGE
C. THE USCARI - GATUN UNCONFORMITY
CORRELATION
A. SANTO DOMINGO MIOCENE
B. BOWDEN BEDS, JAMAICA
C. MEXICAN MIOCENE
D. THE East Coast MIOCENE
Tue Costa RicA GEOSYNCLINE
is i,
Bi
aii
t ¥
179 Costa R1cA Mr1ocENE—OLSSON 7
PRELIMINARY REMARKS
The fossil collections and field information, on which this
monograph is based, resulted from over two years of field work
by the writer in Panama and Costa Rica. The writer’s thanks
are due to the Costa Rica Oil Corporation for permission to pub-
lish; and he is most grateful for assistance, and encouragement
received in the course of this work from Drs. A. C. Veatch and
D. F. MacDonald of the Sinclair Exploration Company, to the
Officials of the National Museum of Costa Rica and Professor
J. Fidel Tristan of San Jose’.
The collections were studied in the Paleontogic Laboratory
of Cornell University, the facilities of which, Professor G. D.
Harris, most generously placed at my disposal.
Cornell University,
January, 1922. A. A. OLsSson
181 Costa Rica MrocENE—OLSSON 9
GENERAL STRATIGRAPHY OF NORTHERN CosTA RICA
The northern or Atlantic slope of Costa Rica, the so-called
‘Tierra Caliente’’ is largely composed of Tertiary sedimentary
rocks, which except for some folding, dip away from the slopes
of the Cordillera Central towards the sea. ‘To the east, this
sedimentary belt may be traced into the Province of Bocas del
Toro of western Panama, where its upper and highly fossilifer-
ous beds are well developed and exposed on the islands of the
Chiriqui Lagoon. ‘This Tertiary sedimentary belt ends some 35
to 40 miles east of the city of Bocas del Toro. ‘The Miocene
sedimentary rocks of Batun and of the northern Canal Zone
form a small independent unit, which begins about 30 miles
west of Colon and ends close to the eastern limits of the Canal
Zone.
In a general way, the stratigraphic succession in northern
Costa Rica is as follows:
Pleistocene Slightly elevated coastalswamps, with deposits of cross-bedded
sands, clay-marls ete.
Pliocene Certain reef-limestones composed of coral and hydrozoa re-
mains; also marls, blue lignitiferous clays, blue coarse sand-
stones, and a predominating hard conglomerate, composed of
large boulders of igneous rocks (the so-called boulder-clays of
several writers on Costa Rican Geology).
Miocene The Gatun Stage or Formation. Coral-reef limestones, (near
the coast), marls, more generally a blue or blue-black sand-
stone, dark colored shales, sandy and impure limestones, ligni-
tes and conglomerates. Generally very fossiliferous.
The Uscari Stage or Formation. Usually dark-colored shales
but with some sandstones, limestones and conglomerates. The
smaller types of foraminfera are usually quite abundant, and
are associated with fish-scales, and small! echinoid spines, with
mollusks generally rare.
Oligocene Sandstones, shales and a complex of hard, often semicrystal-
line limestones which pass through all possible gradations to
coarse arkosic sandstones. The most abundant and characteris-
tic fossils are the Orbitoidal foraminifera (Lepidocyclina). The
IO BULLETIN 39 182
age of these beds probably ranges from Lower to Upper
Oligocene.
THE CostTA RICAN MIOCENE
An important part of the stratigraphic succession of north-
ern Costa Rica is composed of rocks of Miocene age, whose
thickness varies according to locality and completeness of sec-
tion, from a few hundred to several thousands of feet. They are
divisible into two stages or formations by an unconformity or
disconformity of greater or less degree. In their natural order
of sequence, these divisions of the Costa Rican Miocene are:
Gatun Stage or Formation: Middle and Upper Miocene
Uscari Stage or Formation: Lower Miocene
THE USCARI STAGE
The Uscari stage or formation derives its name from Uscari
creek in the Talamanca valley of eastern Costa Rica. It con-
sists principally of soft, dark-colored shales and because of their
slight resistance to denundation, their outcrop frequently forms
wide valleys and interior basins. These shales were deposited in
quiet and moderately deep water, and their fauna is largely
composed of the smaller types of foraminifera, such as Globzg-
evina, Textularia, Lagena, Nodosaria and Miliola, but the
large Orbitoidal Lepzdocycline, abundant in the underlying Oli-
gocene, are absent. Mollusks are very uncommon in this forma-
tion. Thesmall fauna described by Gabb from Sapote, Costa
Rica, belongs to the lower part of the Uscari formation.
THE GATUN STAGE
The upper division of the Costa Rican Miocene is equivalent
in part to the Gatun formation of the Canal Zone. The typical
Gatun, so named from its well-known exposures at Gatun and in
which a part of the Panama Canal and the locks at Gatun were
excavated, lies in an independent sedimentary basin across the
1323 Costa Rico MrocENE—OLSSON II
northern end of the Canal Zone and in the adjacent parts of
the Province of Colon. There the Gatun formation consists
of blue sandstones, dark-colored shales, fuller’s earth beds etc.,
generally less than 500 feet thick. Many of its beds are very
fossiliferous. ‘The mollusks of the Gatun stage of Panama have
been studied by Dall, Toula, Brown, Pilsbry and Cossmann.
In Costa Rica, the Gatun is very much thicker than in
the Canal Zone and represents a longer depositional period.
Paleontologically, the Gatun of the Canal Zone, seems to repre-
sent only the lower part of the formation as developed in Costa
Rica. It is largely of sandy character, more resistant to denu-
dation than the Uscari shales and gives rise to a more rugged
and hilly topography. Although typically sandy, it also contains
locally shales, lignites, conglomerates and coralline limestones.
In some localities the limy members are of major importance.
The formation is characterized by marked lateral variation in
which respect it differs. strongly from the much more uniform
shales of the Uscari stage. ‘This feature of lateral variation
gives rise to two important phases: that of the more typical
Gatun, in which the beds are largely sandy in character, with
some lignitic and coarsely conglomeritic members, and carrying
the usual near shore Gatun fauna; and the coralline phase, in
which the beds are more marine, with coralline limestone and
richly fossiliferous marls perdominating, and with no lignitic
beds.
Along the inner margin of the Miocene belt, the Gatun be-
longs to the first and more typical phase; while the coralline
type is more common along the coastal zone, as at Port Limon
and Bocas del Toro. Coralline rocks may occur at any horizon
in the Gatun succession or it may locally be composed of mas-
sive coral-reef limestones. Such is the case in the Limon Penin-
sula, and such coralline limestones, by most observers, have been
regarded as of Pliocene or even of more recent age. At Limon,
£2 BULLETIN 39 784
this confusion has partly arisen from the fact that these Mio-
cene limestones are overlain by the second series, of Pliocene
age, which partly yielded the large fauna described by Gabb,
published shortly after his death in 1881. Dall later contribut-
ed to the discussion of this fauna. The truly Miocene limestones
of Limon may beseen in good exposures along the sea-front
west of the city. ‘There they are massive in character, hard and
recrystallized to the extent that all traces of organic remains
have usually been obliterated or rendered uncertain of determin-
ation. However, some sandy or marly beds are occasionally
found intercalated in these coralline limestones, and from these
layers we have collected a large fauna of unquestionable Mio-
cene affinities, correlating closely in age with the Bowden beds
of Jamaica.
The fauna of the Costa Rican Gatun is largely molluscan.
The following paleontological study records 334 species. Future
collecting will doubtless greatly add to this list, as several species
common it the Canal Zone, still remains undiscovered in Costa
Rica. ‘The smaller types of foraminifera abound in the more
shaly phases of the formation, as well as several species of echin-
oids, belonging to the genera Clypeaster and Encope. Corals of
simple and compound types occur principally in the coralline
phase.
THE USCARI - GATUN UNCONFORMITY
At the close of Uscari time the Miocene sea was with-
draw from the greater part of Costa Rica, but during the
Gatun, it returned. These conditions are indicated by a
stratigraphic break between the two stages, a complete change in
their lithologic composition, the occurrence of Gatun beds in
areas which had received no deposition during Uscari times and
a change in their respective faunas. Although the full strati-
graphic details of this important subject cannot be presented at
the present time, attention is called to it, as the outstanding fea-
ture of the Miocene stratigraphy of Panama and Costa Rica; and
because of its bearing on correlation.
185 Costa Rica MIocEN# —OLSSON 13
In many parts of northern Costa Rica, the Gatun stage
carries at its base a thick and persistent conglomerate, formed of
coarse cobbles of igneous rocks, or of finer pebbles with alternat-
ing layers of sandstone. This conglomerate generally rests dis-
conformably on the Uscari, but in some cases is unconformable
on these or older rocks. ‘This basal conglomerate is generally
firmly cemented and because of its superior hardness, may be-
come an important ridge former. It marks the site of high
water-falls along mauy streams. Fossils of common Gatun
types are abundant at many localitiesin this basal conglome-
rate.
Gatun rocks are frequently found in areas which contain no
Uscari beds. In such cases, the Gatun is found resting, with
strong unconformity, on older rocks. This condition is seen in
the Canal Zone and along the borders of the sedimentary belt of
the western Panama. It indicates, either a complete erosion of
the Uscari shales during the Uscari-Gatun interval or, what is
more likely, a greater transgression of the Gatun sea. This
uncomformable relation of the Gatun upon older sedimentary
beds may be seen in the Chagres Spillway in the Canal Zone,
where the underlying rocks are sandstones, probably equivalent
to the Mt. Lirio of Lower Oligocene age. Inthe Chiriqui La-
goon country, of western Panama, the base of the Gatun is
frequently a coral-reef limestone, which rests directly upon the
older and more disturbed rocks.
%
>
al sk
The present systematic study enumerates 334 species of
molluska of which the majority belong to the Gatun stage. A
few of these species have not been found in Costa Rica or west-
ern Panama, but are common in the Canal Zone, and are includ-
ed here for the sake of comparison with Costa Rican species.
To these we may add the 18 species described by Gabb from
Sapote, Costa Rica, but not included in our collections and the
following species listed below from the Canal Zone. This brings
14 BULLETIN 39 186
the total known Miocene fauna of Panama and Costa Rica to
396 species.
Volvulella micratracta Brown and Pilsbry
Ringicula hypograpta Brown and Pilsbry
Conus haytensis Sowerby Corbula gatunensis Toula
Conus domingensis Sowerby Corbula sphenis Dall
Cruns consobrinus Sowerby Corbula sericea Dall
Conus emulator Brown and Pilsbry
Drillia vaningent Brown and Pilsbry
Drillia gatunensis 'Toula Solecurtus strigillatus Linné
Drillia fusinus Brown and Pilsbry
Drillia Zooki Brown and Pilsbry
Drillia enneacyma Brown and Pilsbry
Cythara heptagona Gagg
Marginella gatunensis Brown and Pilsbry
Anachis fugax Brown and Pilsbry
Murex polynematicus Brown and Pilsbry
Murex gatunsts Brown and Pilsbry
Typhis Gabbi Brown and Pilsbry
Malea camura Guppy
Cyprea Henekint Sowerby, var.
Littium nugatorium Brown and Pilsbry
Natica bolus Brown and Pilsbry
Natica canalizonalis Brown and Pilsbry
Sigaretus gatunensis 'Toula
Sigaretus (Eunaticina) Gabbé Brown and Pilsbry
Cheilea princetonia Brown and Pilsbry
Nucula (Acila) isthmica Brown and Pilsbry
Pecten effosus Brown and Pilsbry
Pecten operculariformis ‘Toula
Pecten reliquus Brown and Pilsbry
Amusium Toule Prown and Pilsbry
Ostrea gatunensis Brown and Pilsbry
Cardium durum Brown and Pilsbry
Cardium gatunensis Dall
Tellina equiterminata Brown and Pilsbry
187 Costa RicA MrockNE—OLSSON 5
Chione tegulum Brown and Pilsbry
Pitaria cora Brown and Pilsbry
Dosinia delicatissima Brown and Pilsbry
Petricola millestriata Brown and Pilsbry
Cyclinella gatunensis Dall
Thracia gatunensis Toula
The Uscari fauna is still poorly known, but it includes sev-
eral very distinctive species of which the following are most
important:
Amusinm Lyonit Gabb
Arca dariensis Brown and Pilsbry also Lower Gatun
Clementia dariena Conrad also Lower Gatun
Canccllaria (Aphera) islacolonis Maury
Neverita neretdis Maury
Sconsia cocleana Olsson
Scaphella costaricana Olsson
Ptychosalpinx ? dentalts Olsson
Dentalium uscarianum Olsson
The Gatun is the great fossil bearing formation of northern
Costa Rica and Panama, and is characterlzed by a rich and high-
ly diversified fauna. This fauna at the present numbers nearly
400 species, but additional collecting will doubtless add consid-
erably to this number. As we have already noted the Gatun
beds present two important facies, the sandy and highly clastic
phase of the typical Gatun, and the deeper water type, illustrat-
ed by the coralline limestones and their associated marls. These
lithological types have their own more or less peculiar and dis-
tinctive faunas.
The typical Gatun is the more common phase and is charac-
terized by a molluscan fauna of heavy, thick-shelled species of
littoral forms. In Costa Rica, the following are the most com-
mon and characteristic:
Arca MacDonaldi Dail Marginella MacDonaldi Dall
Arca Pittiert Dall Voluta Alfarot Dall
Pecten levicostatus 'Toula Strombus pugiloidis Guppy
16 BULLETIN 39 188
Cardium stirtatum Brown, Pilsbry Stormbus gatunensis Toula
Chione Rowelli Olsson Natica Guppyana ‘Toula
Chione mactropsis Conrad Turritella altiliva Courad and its
Dosinia acetabulum Conrad varieties
Conus molis Brown and Pilsbry
The coralline phase is developed at several localities along
the coast, most important of which are Port Limon and Bocas
del Toro. Here the coralline limestones and associated marls
form the greater part of the Gatun section, but coralline rocks
may occur, interbedded with typical Gatun rocks, at any strati-
graphical level.
These coralline limestones were formed, mainly as barrier
reefs some little distance from the old Gatun shore-line. Natur-
ally they offered very different habitat conditions from those of
the typical near-shore and often lacustrine Gatun. Hence we
find a fauna of deeper-water affinities with Pleurotomids and
thin-shelled Cones, as the most important and characteristic
element. In localities where these coralline rocks are exclusive-
ly developed, this fauna bears very little relation to the typical
Gatun fauna, although the two are contemporaneous. The
following are species of the coralline phase:
Pecten coralliphila Olsson Mitra Alamagrensts vat.
Antigona Flarristana Olsson coralliphila Olsson
Gafrarium limonensis Olsson Mitra poas Olsson
Large number of Pleurotomids J/arginella avena Valennencies
Conus planilivatus Sowerby Olivella limonensis Olsson
Conus limonensis Olsson Phos elegans, variety limonensts
Conus stenostomus Sowerby Olsson
Turritella exoleta Linné Stliquaria modesta Dall,
var. limonensis Olsson limonensis var. Olsson
About 13 per cent. of the Gatun fauna is identical or closely
related to the recent species, the majority of which are found
living along the present Caribbean coast, certain others only on
the Pacific side in the Panama Province and a few common to
both. ‘This large percentage of recent forms corresponds closely
189 Costa Rica MioceNE—OLSSON 17
with that characterizing the Chesapeake Miocene fauna of the
eastern United States and shows their close agreement in age.
The recent species and their closely related forms in the Gatun
fauna are as follows:
Arca occidentalis Phil. West Indian
Arca umbonata Lamarck West Indian
Arca candida Gmelin West Indian
Arca auriculata Lamarck West Indlan
Ostrea megodon Hanley Pacific
Anomia simplex d’Orb. West Indian
Phacotdes vadians Conrad var. medioantericanus
Olsson West Indian
Phacoides trisulcatus Conrad West Indian
Glycymeris castaneus Lamarck West Indian
Glycymerts decussatus Linné West Indian
Divaricella guadrisulcata d’Orb. West Indian
Cardium medium Linné West Indian
Cardium spinosum var. Turtont Dall East Atlantic
Cardium serratum Linné West Indian
Pitaria circinata Born and its var. alternata Broderip
West Indian and Pacific
Antigona multicostata Sowerby Pacific
Antigona rugosa Gmelin West Indian and Pacific
Macrocallista maculata Linné West Indian
Chione grus Holmes West Indian
Tellina crystallina Chem. West Indian and Pacific
Strigilla pistformis Linné West Indian
Mactra exoleta Gray Pacific
Mactra alata Spengler West Indian
Labiosa lineata Say West Indian
Corbula Dietziana C. B. Adams West Indian
Tellidora cristata Recluz West Indian
Conus proteus Hwass West Indian
Conus floridanus Gabb var. costaricensis Olsson West Indian
Terebra benthalis Dall var. bocasensts Olsson West Indian
18 BULLETIN 39 190
Turris albida Perry West Indian
Glyphostoma dentifera Gabb West Indian
Acteocina bullata Kiener var. costaricana Olsson West Indian
Acteocina recta d’Orb. West Indian
Volvula cylindrica Gabb West Indian
Anctstrosyrinx elegans Dall variety West Indian
Marginella avena Valen West Indian
Murex messorius Sowerby West Indian
Bursa crassa Dillwyn West Indian
Natica canrena inne West Indian
Architectonica granulata Lamarck West Indian nnd Pacific
Astralium brevispinum Lamarck, variety dasalis
Olsson West Indian
Astralium caclatum Gmelin West Indian
Sastrum nodulosum C. B. Adams West Indian
Latirus tufundibulum Gmelin West Indian
Northia northie variety mtocenica Olsson Pacific
Crepidula plana Say West Indian
Xenophora conchyliophora Born West Indian
Oliva testicea Lamarck var. costaricensts Olsson
It is but natural to expect that the Miocene beds along the
northern or Caribbean coast of Central America, should contain
a fauna largely ancestral to that of the present Caribbean, but
along with these strictly Caribbean types, we also find several
species, identical or closely related to forms now resticted to the
Pacific side. ‘Their occurrence in Caribbean deposits points to a
connection between the Atlantic and Pacific oceans during Mio-
cend time and allows for a certain intermingling of their faunas.
Geological observations indicate that there were at least two
such straits crossing Central America during the Miocene, the
main one across Costa Rica, and a second through the Darien of
eastern Panama.
This Pacific element of the Gatun fauna comprises many of
the most common and characteristic species. Certain of these
species had a wide distribution in the West Indies during Mio-
QI Costa Rico Mi1ocENE—OLSSON 19
cene time and have been noted by several writers. They
comprise the following species from Panama and Costa Rica,
listed together with the Pacific analogues:
Panama or Costa Rica Pacific Analogue
PAGO RAL L OVO SOM CTD paakeiscies stisesaiceire Arca grandis Sowerby
Arca (Noetia) MacDonaldi Dali.....drca reversa Sowerby
Ostrea megodon Hanley
Mactira exoleta Gray
Mactra estrellana O\SSON..........1.000+ + Mactra (Harvella) elegans Sowerby
Semele laevis Sowerby var.
costaricensts Olsson........semele laevis Sowerby
Macoma panamensis Dall variety
canalis Olsson...J7acoma panamensis Dall
Macoma gatunensis Toula............... Macoma elongata Hanley
Tellina crystallina Chemnitz............ also West Indian
Antigona multicostata Sowerby
Antigona rugosa Gmielin................. also West Indian
Pitaria circinata Born variety
alternata Broderip
Cyclinella subqguadrata Hanley variety
guitana Olfson...... Cyclinelia subguadrata Hanley
Cyclinella beteyensis O\SSON.........2+- Cyclinella Kroyeri Philippi
Conus vecognitus GUPPy...--.-..+2e0eeee Conus pyriformis Reeve
Turricula lavinoides Olsson............ Turricula lavina Dall
Cancellaria islacolonis Maury.......... Cancellaria tesselata Sowerby
Cancellaria toroensis Olsson..........+. Concellaria tuberculosa Sowerby
Cancellaria Pluimmert Olsson.......... Cancellaria bullata Sowerby
Mitra Swainsont Brod. variety
limonensis Olsson...... Mitra Swainsont Broderip
UNIEVEMILONMETELATS, MIAULY, (21.004 .5> 23.060 Neverita glauca Humbolt
Northia northiae Gray variety
miocenica Olsson.....- Northia narthiae Gray
INAIECONCOMVUTA (GUPPYs..5.22-64-de006e++-0- Malea ringens Swainson
Oliva testacea Lam. variety
costaricensis Olsson...... Oliva testacea Tam
With the closing of the Miocene straits, this Pacific element
gradually had to give way before the development of the West
Indian fauna, but it is still to be seen in such species as, 7e/lina
crystallina Chem. Architectonica granulata Lamarck, which
have a distribution along both coasts. Moreover the Pliocene
20 BULLETIN 39 192
and Pleistocene beds of Central America contain a well-marked
Pacific element in such species as Worthia northie Gray Oliva
testacea Lamarck and Pecien ventricosus Sowerby.
CORRELATION
Miocene rocks are now known from a great many localities
in the West Indies and the general Caribbean area of Central and
South America. In these areas they comprise a very important
series of sedimentary formations. Beds equivalent to the Gatun,
appear to be the most widespread and generally are highly fossil-
iferous.
The following chart showing the equivalence of the Miocene
horizons in the West Indies and elsewhere to those of Central
America, does not differ materially from the more recent correl-
ations proposed by Maury, Cooke or Vaughan, except in the ad-
dition of the Uscari to the Lower Miocene of Panama and Costa
Rica. The Gatun formation is here recognized as being of Mid-
dle Miocene age with possibly some of its upper beds having
Upper Miocene affinities. The Gatun, at its type locality in the
Central Zone, is not more than 500 to 600 feet thick. In Costa
Rica it is very much thicker, and its higher beds may range into
the Upper Miocene. It is the exact equivalent of the Chesapeake
Miocene of the eastern United States. In this correlation, I have
been guided not only by the paleontologic evidence, but also by
the field relations of the formations, studied over a wide rage of
country in Costa Rica, and in Panama as far east as the Colombia
frontier. .
SANTO DOMINGO MIOCENE
In Santo Domingo and in Haiti, Miocene rocks are exposed
in several areas, but attain their best development along the val-
ley of the Rio Yayue del Norte in the northern part of the is-
land. ‘These beds are highly fossiliferous and because of the fine
perservation of their fossils, have repeatedly attracted the atten-
tion of paleontologists.
The earlier writers dealing with the paleontology of the Do-
minican Miocene, believed thatit represented but a single
X93 Costa RIcA MIocENE—OLSSON 21
stratigraphic unit, and Gabb insisted that no division was possi-
ble. Dr. Maury however, as a result of paleontologic studies
from collections carefully made in the field, was able to divide
the Dominican Miocene into two formations, which she called
the Gurabo and the Cercado. ‘To these Vaughan and his as-
sociates, in more recent stratigraphic work on the island, have
added 4 other subdivisions. The present recognized subdivisions
of the Dominican Miocene are:
Mao clay
MIDDLE Mao Adentro limestone
Gurabo formation
(Sconsza laevigata zone)
Cercado formation
(Aphera islacolonis zone)
Baitoa formation and Bulla
conglomerate
MIOCENE
LOWER
The relation of the Costa Rican and Panama Miocene to
that of Santo Domingo is extremely close; there being about
go species in common. Of this number about 21 are recent,
about 26 species appear to be restricted to the Gurabo, some 27
species are common to both the Gurabo and Cercado formations,
and only 11 species are confined to the Cercado. The list of
of Costa Rican and Panama species in the Dominican Miocene,
is as follows:
Nucula tenutsculpta Gabb Cercado and Gurabo
*Arvca occidentalis Phil Cercado and Gurabo
*Aryca umbonata Lamarck Cercado
Arca Maurye Olsson Gurabo
Arca FHlenekent Maury Cercado
Arca goltoyaguensis Maury Cercado and Gurabo
Arca chiriquiensits Gabb Gurabo
*Arca auriculata Lamarck Gurabo
% Recent
i)
N
BULLETIN 39 194
Glycymerts jamatcensts Dall Cercado
Lime@a solida Dall Gurabo
*Ostrea megodon Hanley Gurabo
Spondylus gumanemocon Brown and Pilsbry, Gurabo
*Anomia simplex d’Orb Gurabo
Placunanomia lithobleta Dall Gurabo
Pteria inornata Gabb Cercado and Gurabo
*Cardium serratum Linné Cercado and Gurabo
Cardium dominicensis Gabb
*Cardium medium Linné Gurabo
Protocardia gurabica Maury Cercado
*Wactra alata Spengler Cercado
Antigona tarquina Dall? Cercado
Antigona Blandiana Guppy Gurabo
*Pitarta circinata Born Cercado
* Tellina crystallina Chemnitz Cercado
Tellina cercadica Maury Cercado
*Strigilla pistformis Linné Cercado
Semele Claytont Maury Cercado
Venericardia scabricostata Guppy Cercado and Gurabo
Echinochama yagquensis Maury Gurabo
Corbula vieta Guppy Cercado and Gurabo
Corbula viminea Guppy Cercado
** Acteocina recta d’Orb Cercado
Acteocina triticum-tritonts Maury Cercado and Gurabo
* Volvula cylindrica Gabb Cercado
Bullaria paupercula Sowerby Cercado and Gurabo
Ringicula dominicana Maury Cercado
Terebra spirtfera Dall Cercado
Terebra bipartita Sowerby Gurabo
Tercbra haitensts Dall Gurabo
Terebra Berlinere Maury Gurabo
Conus haytensis Sowerby Cercado and Gurabo
>? ~~ Sewalli Maury Gurabo
“Recent
CostA RicA MrocENE—OLSSON
Conus stenostomus Sowerby Gurabo
Conus proteus HAwass Cercado and Gurabo
Conus recognitus Guppy Cercado and Gurabo
Conus multiliratus Bose Gurabo
Conus gracilissimus Guppy Cercado and Gurabo
Conus marginatus Sowerby Gurado
Conus consobrinus Sowerby Cercado and Gurabo
Conus domingensis Sowerby Gurabo
* Turris albida Perry Cercado and Gurabo
Drillia venusta Sowerby Gurabo
Drillia consors Gurabo
Cythara heptagona Gabb Gurabo
*Glyphostoma dentifera Gabb Gurabo
Cancellaria Maurye Olsson Cercado and Gurabo
Cancellaria Rowelli Dall
Cancellaria epistomifera Guppy Cercado and Gurabo
Cancellaria tslacolonis Maury Cercado
Oliva brevispira Gabb Cercado and Gurabo
Oliva cylindrica Sowerby Cercado and Gurabo
Olivella muticoides Gabb Cercado and Gurabo
Mitra rudis Gabb Cercado and Gurabo
Mitra longa Gabb Cercado and Gurabo
*Latirus infundibulum Gmelin Gurabo
Melongena consors Sowerby Cercado and Gurabo
Alectrion losquemadica Maury Gurabo
Metula cancellata Gabb Gurabo
Murex cornurectus Guppy Cercado and Gurabo
*Vurex messorius Sowerby Gurabo
Typhis alatus Sowerby Gurabo
Distortio simillima Sowerby Gurabo
*Bursa crassa Dillwyn Gurabo
Cassis sulcifera Sowerby Cercado and Gurabo
Sconsia levigata Sowerby Gurabo
*Recent
i)
Ww
24 BULLETIN 39 196
Dolium camura Guppy Gurabo
Dolium elliptica Pilsbry and Johnson Gurabo
Cypraca Henekent Sowerby Cercado and Gurabo
Cypraca dominicensts Gabb Gurabo
Strombus pugiloides Guppy Gurabo
Serpulorbis papulosa Guppy Cercado and Gurabo
Petaloconchus sculpturatus H.C.lea Cercado and Gurabo
*Architectonica granulata Lamarck Cercado
*Xenophora conchyliophora Born Gurabo
*“Natica canrvena Linné Cercado and Gurabo
Natica Youngi Maury Cercado and Gurabo
Polinices subclausa Sowerby Cercado and Gurabo
Polinices Stanislas-Meuntert Maury Gurabo
Neritina viridemaris Maury Coreado and Gurabo
LNeverita neretdis Maury Cercado
Most of the above listed species belong to the Gatun forma-
tion. Cancellaria (Aphera) islacolonis Maury, Neverita neretdis
Maury and Corbula viminea Guppy occur in the Upper Uscari
and in the Lower Gatun or transitional beds of eastern Costa
Rica. This leaves but the following 8 Cercado species in the
Gatun fauna, or a proportion of about 3.25 to 1.
Arca Henekent Maury Winter Cay
Glycymeris jamaicensts Dall Winter Cay
Protocardia gurabica Maury Water Cay and Gatun
Antigona tarquinia Dall? Saury
Tellina cercadica Maury Saury
Semele Claytont Maury Hotel Creek
Corbula viminea Guppy Middle Creek
Ringicula dominicana Maury Middle Creek, Red Cliff
Creek and Estrella River
*Recent.
197 Costa RicA MioceEnm—OLSSON 25
GENERAL CORRELATION CHART NO. I.
Costa Rica and | Canal Santo Jama Porto
western Panama Zone Domingo tea | Rico
SSI SN Ese Set SC (Se Se itera SEL ee TA SRNL cI USEA EINE INT Ie Dick) TOSI ne
|
v .
5 ms Limon. Monkey Las Matas
aan Point, Bocas Stage
m=
Be Conglomerates
(boulder clays)
SRB BR MRE RB aT anes lhe tl Seah meted llinshl tome) \fecriph ea | fost. suse y Nike | %
ed h | |
a | Cerros de | |
BY 2? Toro | Sal Stage hoe
P Gatun Stage Stage Ui @ucbras
\Bowden| dillas
Stage act
A Mao Clay
4A Gatun
jaa 5 Stage |Mao Adeotro
o &. Gatun Stage | limestone °
on Gurabo Stage
>
s! !
FA € ie iS * % Ae Bees Vlies Sn bce c < < oe me | S asl) Tas v
fa Cercado Stage |
= -Uscari Stage |
e) (Sapote) |
aN | Baiton Stage
Reisen Reise
Seles | ena Seed oe Ni aap fet <r Te ro KITS ach ha ‘ |
o) | | |
5 ‘ Emperado l’s | |
5 Undifieren- | Culebra Stage | Cevicos l’s |
ae tiated | Bohiocongl.| Tabera Stage |
©
26 BULLETIN 39 198
GENERAL CORRELATION CHART NO. il.
Colombia and Mexico Eastern
Venezuela United Staies
* x a * * * me * * ¥ Dr x %* % * * * * * * + x% +
Caloosahatchie
| Waccamaw
|
o st x x x ¥ xX s ft * TAO TAR eM Ph Ge) Te hes oe aS
Txuetepec, State} & Yorktown and Duplin Stage
3 of Oaxaca a &
is ‘<> Murfreesboro Stage
5 Santa Rosa and |
Cartagena and |& Barranca de v
9
Cumana a Santa |
5 2,
\2 Maria Tatetla, |s'. St. Mary’s Stage
he] Nw
© State of Vera aie
° oq Cruz a) é Choptank Stage
|
| Calvert Stage
eae ac Viyisen sen Wese UAC SORA R NE NESE na Cai rat igiay a ean | atl bea Niet eg |S aa)
i a)
Peano aay @ Shoal River marls
ae eet om eeyan| Oak Grove marls
Bre Hoe Bay Chipola marls
Soa
\<q
x x aa w ea ileal eri wah lee CY cogil aay 2 [Py EA apy es Veo te iiMlezas Weel ea ea Se
Tampa Stage
Vickburgs Stage
199 ~ Costa Rico MrocENE—OLSSON 27
The Arca Henekent, Glycymerts jamaicensis and Protocardia
gurabica occur with a typical Gatun fauna at Water Cay, west-
ern Panama, associated with Sconsza, and other Gurabo shells.
With the exception of Ringicula dominicana,the other species are
generally rare. ‘The evidence is therfore strongly in favor of
the equivalence of the Gatun beds with the Gurabo of Santo
Domingo.
BOWDEN BEDS, JAMAICA
Next to Santo Domingo, the highly fossiliferous beds at
Bowden, Jamaica, are the best known. From this locality, Dall
has listed about 435 species, of which 12 per cent. appear to be
identical with recent species.
Recent correlation by Cooke places the Bowden beds as
equivalent to the Gurabo formation of Santo Domingo,and there-
fore to the Gatun formation; a correlation which is borne out by
our own studies. About 38 Gatun species, not including recent
forms, are here considered equivalent to Bowden shells. In gen-
eral, the Costa Rican localities which show the closest agree-
ment with Bowden, are the upper coralline horizons, so that the
Bowden beds may be a little higher than the main Gurabo and
they are so considered here. The following Bowden shells oc-
cur in the Costa Rican Gatun.
*Acteocina bullata Kiener
*Acteocina recta d’Orb Petaloconchus sculpturatus Wea
*Volvula cylindrica Gabb Alaba turrita Guppy
Lullaria paupercula Sowerby *Crepiduia plana Say
Terebra bipartita Sowerby * Nrtica near canrena Linne
Conus planiliratus Sowerby Glycymeris jamaicensis Dall
Conus stenostomus Sowerby *Arca occidentalis Philippi
Conus granozonatus Guppy *Arca auriculata Wamarck
Conus recognitus Guppy *Ostera megodon Hanley
Conus interstinctus Guppy Lime@a solida Dall
Conus gracilissimus Guppy Placunanomia lithobleta Dall
*Turris albida Perry Echinochama antiguata Dall
*Recent
28 BULLETIN 39 200
Drillia consors Sowerby Venericardia scabricostata Guppy
Drillia venusta Sowerby Myrtea limoniana Dall
Cancellaria Barretii Guppy Phacoides actinus Dall
Oliva cylindrica Sowerby *Levicardium serratum Vinne
*Latirus infundibulum Gmelin Cytherea Blandiana Guppy
Melongena consors Sowerby Tellina lepidota Dall
Metula cancellata Gabb *Strigilla pistformis TLinne
Typhis alatus Sowerby Abra triangulata Dall
Distortrix simillima Sowerby Corbula viminea Guppy
“Bursa crassa Dillwyn Gastrochena rotunda Dali
Cassis sulcifera Sowerby Phalium moniliferum Guppy
Sconsia sublevigata Guppy Malea camura Guppy
Strombus pugiloides Guppy Serpulorbis papulosa Guppy
MEXICAN MIOCENE
From Mexico, Boseft has described several small faunas
from the Isthmus of Tehuantepec, and from the States of
Oaxaca and Vera Cruz. These faunas, he considered as Upper
Miocene and Pliocene.
The most interesting of these localities, in the present con-
nection, are those at Tuxtepec in the State of Oaxaca, and
Santa Rosa and the Barranca de Santa Maria Tatetla in the State
of Vera Cruz. From these localities, we may note together
with their Costa Rican and other affinities, the following
species:
Fecten santarosanus Bose Fecien levicostatus Youla Gatun
Amustum Mortont Ravenel Amusium luna Brown and Pilsbry
Levicardium sublineatum Conrad Gatun Chesapeake Miocene
Anomia simplex d’Orb Gatun-Chesapeake to recent
Venus Ebergenvit Bose Atone ulocyma Dall Upper Chesapeake
Solarium Villarelloi Bose Architectonica granulata Lamarck
Lower Miocene or recent
Vermetus pulcher Bose Serpulorbis sculpturatus H.C. Lea
Gatun and Upper Chesapeake Miocene
Strombus pugilis Linne Strombus pugtloidis Guppy Gatun
Sconsia sublevigata Guppy Bowden
Phos mexicanus Bose Gatun of Water Cay
Melongena Mengeana Dali Solenosteira Vaughani Dall variety
medioamericana Olsson,Gatun of
Water Cay
* Recent.
+ Boletin del Instituto Geologico de Mexico, No. 22, 1919, pp. 11 - 88
204 Costa RicA Mriockn® —OLSSON 20
fleurotoma alesidota Dall var.
MAGMA BOSC......000-. Drillia consors Sowerby, Gatun
Conus Agassizi Dall variety
multilivatus BOSe........... Conus multiliratus Bose, Gatun
COMUSMEUYGRRATLD BOSC aN soe seces. Gatun
The above species are from Tuxtepec and Santa Rosa. To
these we may add; from Barranca de Santa Maria Tatetla, the
following forms:
Ostrea sculpturata Conrad.............. Ostrea costaricensis Olsson, Gatun
Lucina quadrisulcata Vinné........... Divaricella quadrisulcata d’Orb
Gatun, Upper Chesapeake Miocene to
recent
Levicardium serratum Vinné........ Gatun, Upper Chesapeake to recent
Dosinia acetabulum Conrad..........-. Gatun - Chesapeake Miocene
The above list is but a partial one of the entire fauna from
these localities. However,’ it shows the close relationship be-
tween these Mexican species and those of the Gatun and the
Bowden, such as Sconsia sublevigata, Phos mexicaius, Conus
multilivatus and others, witha few typical Chesapeake species
such as Chione ulocyma Dall, Dosinta acetabulum Conrad and
Cardium sublineatum Conrad. ‘This illustrates the interming-
ling of the Gatun-Bowden-Gurabo fauna with that of the Chesa-
peake, and proves their equivalence in age.
THE East Coast MIOCENE
A fine development of Miocene rocks is found along the
Atlantic coastal plain and they extend more or less continuously
from New Jersey south into Florida. They are continued around
the borders of the Gulf of Mexico into Texas, but in these re-
gions, their marine phases are deeply covered by younger forma-
tions, and they are known only from deep wells. These Mio-
cene beds are divisible into two series, which early were classifi-
ed as (a) the older or sub-tropical Miocene and (b) the younger
and cold-water Miocene. These terms are descriptive of their
main faunal differences. The upper division consisting of sever-
30 BULLETIN 39 202
al formations or stages is frequently known as the Chesapeake
Miocene or Chesapeake series, and the lower consisting solely of
the Alum Bluff formation, may be designated for sake of discus-
sion, the Alum Bluff series. These Miocene formations or stages
are the following:
| Yorktown, Duplin Stages
Upper Murfreesboro Stage
ml, * * * * | Chesapeake Series/* * * * * * * *
a | |St. Mary’s Stage
ea Middle | Choptank Stage
Oi Calvert Stage
oO | |
4 | Mie a Meira aa egos.) Re | BEC RCT RA aaa a ies |e
= | Alum Shoal River marl
| _ Bluff
| |Alum Bluff Series} Stage Oak Grove marl
| Lower Chipola marl
|
This two-fold division is a very natural one. Each series
represents a distinct trangression of the Miocene sea over parts of
the Atlantic coastal plain, that of the Chesapeake being the
more extensive. In Florida, where the Chesapeake Miocene is
sometimes found resting upon the Alum Blufi beds, as at Alum
and Jackson Bluffs, the contact is seen to be an erosional uncon-
formity. At these localities, the Chesapeake Miocene is equiva-
lent to the Murfreesboro stage or formation of Virginia and
North Carolina, and hence the time interval represented by this
break is equivalent to the Lower Chesapeake or the Marylandian
Miocene. Dall,* who has paid special attention to the relations
of the Alum Bluff and the Chesapeake series, both stratigraphi-
cally, and paleontologically, characterizes this break as ‘‘the
most sudden, emphatic, distinct in the whole post-Cretaceous his-
* Trans, Wagner Free Inst., 1903, p. 1594
203 Costa Rica Mi1ocENE—OLSSON 31
tory of our southeastern Tertiary, and indicates physical changes
in the surrounding region, if not in Florida itself, sufficient to
alter the course of ocean currents and wholly change the temper-
ature of the waters of our southern coasts.”’
The fauna of the Lower Miocene or Aluin Bluff series is of
warm-water character and largely composed of Antillean types.
In the Chesapeake series, the fauna is of cold or temperate-wat-
er character and it finds its relations with the present Atlantic
fauna north of Cape Hatteras. These faunal differences must be
take into account in any correlation between the Miocene of the
United States with that of the West Indies and Central
America.
In the Correlation Charts,I have placed the Gatun formation
as equivalent to the Chesapeake series, the Uscari formation
with the Alum Bluff. The evidence upon which this correla-
tion is based, may be summarized as follows:
1. The two-fold division of the Costa Rican and Panama
Miocene, by an unconformity at the base of the Gatun forma-
tion.
2. Wherever the Gatun beds have been found overlain by
more recent formations, the age of these overlying beds is Plio-
cene or younger.
3. The high percentage of recent species in the Gatun
fauna, which agrees with the average as shown for the Chea-
peake Miocene.
4. The Gatun fauna, contains a few species which appear
identical with strictly Chesapeake species, such as the follow-
ing:
Dosonia acetabulum Conrad
Petaloconchus sculpturatus WH. C. Lea
Chama congregata Conrad
Plicatula marginata Say?
32 BULLETIN 39 404,
The Dostnia acetabulum-Conrad is one of the most abundant
and characteristic species of the Chesapeake Miocene and in its
course through the Chesapeake formations, several varieties are
developed, of great use in correlation. The typical acetabulum
is however confined to the Upper Chesapeake formations and it
was figured by Conradin his Medial Tertiary, from a fine large
example from Smithfield, Virginia. It is characterized by its
large size, broad outlines and even, concentric banding. This
Upper Chesapeake or typical acetabulum corresponds very close-
ly in size, form and sculpture to the Costa Rican examples. It
is very common at certain localities in Costa Rica, especially in
the lower part of the Gatun formation. Bose records it from
Mexico. The VPetaloconchus sculpturatus is a widely distributed
species in the West Indies and Caribbean Miocene, from which
it is more generally recorded as P. dominigensts, a name given to
the Santo Domingian shell by Sowerby. Close comparative
study of a large collection of typical seulpturatus from Virginia
and North Carolina, failed to show any real difference, either in
details or sculpture or in nature of coiling. The Petaloconchus
sculpturatus occurs in the Bowden beds of Jamaica; the Gurabo
formation of Santo Domingo; the Quebradillas limestone of
Porto Rico; the Springvale beds of Trinidad; and the Gatun of
Panama and Costa Rica. It is also known from Cumana, Vene-
zuela: Cartagena, Colombia; and Santa Rosa, Mexico. In the
Chesapeake Miocene, it is restricted to the Murfreesboro, York-
town and Duplin formations, that is, to the Upper Chesapeake.
It does not occur in the overlying Pliocene so far as is known.
5. Possibly more significant because of their greater num-
ber are the following species, which appear for the first time in
the Atlantic coast Miocene in the Chesapeake formations. They
also continue into the Pliocene and generally into the recent
fauna.
205 Costa Rica MrockENE—OLSSON aa
Anomia simplex d’Orb St. Mary’s to recent
Cardium spinosum var. Tur-
tont Dall Murfreesboro to recent
Cardium serratum Linné © Murfreesboro to recent
Cardium medium Linné St. Mary’s to recent
Labiosa lineata Say Duplin to recent
Tellina umbra Dall Duplin to recent
Chione grus Holmes Duplin to recent
Phacotdes radians Conrad Duplin to recent
Phacotdes trisulcatus Conrad Chesapeake to recent
Divaricella quadrisiulcata d’Orb Murfreesboro to recent
Oliva sayana var.immortua
Pilsbry and Brown typical shell, upper
Chesapeake to recent
Fasctolaria tulipa Linné Pliocene
Tur Costa RICAN GEOSYNCLINE
The wide distribution of Miocene beds around the borders
of the Gulf of Mexico and Caribbean sea, indicates a condition
of general deposition during Miocene time. Costa Rica was
largely under water, at least during the Lower Miocene, and
was the locus of a strait or geosyncline that then connected the
Atlantic and Pacific Oceans; while Panama, parts of Nicaragua,
and other central American areas were mainly above sea level.
This condition is shown in the general igneous character of the
backbone of Panama, while in Costa Rica, sedimentary rocks of
Miocene age occur even in the high interior and at elevations of
several thousands of feet. This geosyncline covered the greater
part of Costa Rica, a portion of western Panama, and western
Nicaragua. It should be noted that this area, at the present
time, is the locus of a large number of active volcanoes.
Certain changes of importance took place at the close of
Uscari time, resulting in a brief withdrawal of the Miocene sea.
The effects of these changes were widespread. These great land
movements, which probably reached a culmination in the Plio-
34 BULLETIN 39 206
cene, had far reaching consequences and to them we must ascribe
even the sudden and marked change between the fauna of the
Alum Bluff and Chesapeake series of the eastern United States.
It probably resulted in a partial closing of the Atlantic-Pacific
straits and a disarrangement of the direction of oceanic currents.
The strongly clastic character of the Gatun formation as compar-
ed with the more shaly Uscari, is also a result of the elevation of _
the land.
The Pliocene was a period of high elevation and, in Costa
Rica, of intense volcanic activity. The coarse conglomerates,
or the so-called boulder clays, which have perplexed most geo-
logists who have visited Costa Rica, were formed from the
erosion of these Pliocene lava flows and deposited as flood-plain
material. In a few localities where these conglomerates reached
the sea, they contain intercalated shale beds carrying marine
fossils of late Pliocene age.
MU 7 1909
MONE Sax
GWASS| GAS RROPODA
ORDER OPISTHOBRANCHIATA
SUBORDER TECTIBRANCHIATA
Genus ACTAEON, Montfort
Actzon costaricensis, n. sp.
Shell moderately large, ovate-cylindric; body-whorl large
and more than 3 of the total length of the shell; spire pointed,
conic; whorls about 6, convex and with deep sutures; spiral
sculpture of fine, nearly regular, smooth threads separated by
deep and somewhat wider interspaces; there are about 9 spiral
threads on the spire-whorls and about 34 on the last whorl; the
interspaces are crossed by fine, elevated, distant threads; aper-
ture linear-lanceolate, rounded below; columella with a single
large fold.
Height 19.5 mm, diameter Io mm, aperture 14. 5 mm.
This isan unusually large and cylindrical species with a
relatively high spire. Suggestion of the original coloration is
indicated by a white band which encircles the middle of the
last whorl commencing at the upper end of the aperture,— the
general ground-color of the surface being a faint purple or red-
dish brown. ‘The interspaces between the spirals are crossed by
fine, elevated threads, but they appear merely punctate on the
early whorls.
Gatun Stage: Hill No. 3, Banana River.
Middle Creek.
Genus ACTEOCINA, Gray
Acteocina bullata Kiener, var. costaricana, n. var. Plate 15 figures 20, 21
cf. Tornatina bullata Kiener, Icon. Coq. Viv., p. 5, pl. 1, fig. 4.
206a a 34a
ct
34b BULLETIN 39 206)
Shell rather large, moderately solid; outlines subcylindrical
but a little wider about the anterior half of the shell; the spire
is short, composed of 4 or more whorls, and a small nucleus of
the usual type, asin canaliculata,; sutures channelled; the sur-
face appears smooth, until slightly magnified when it is seen to
be covered with very fine spiral lines; columella carries a single
strong fold. ;
Height 10, diameter 3, aperture 9 mm.
The Costa Rican fossils are related to the recent A. bullata
Kiener, of the West Indies but differ in their shorter spire and
are somewhat more expanded about their anterior half. It is the
largest species in the Costa Rican Miocene, often reaching a
length of 1o mm. Its surface is covered with fine, spiral lines
and the columella carries a single, strong plication.
Gatun Stage: Middle Creek.
Coll. 4, East Grape Point Creek.
Acteocina recta d’Orbigny
' Bulla vecta d’Orbigny, 1845, Delas Sagra, Hist. Pol. y Nat. Isla de
Cuba, vol. 5, p. 67, no. 55; Atlas 8, pl. 4, dzs, figs. 16-20.
Tornatina recta Gabb, Trans. Phil. Soc., vol. 15, p. 246.
Tornatina coix-lacryma Guppy, 1876, Quart. Journ. Geol. Soc. Lon-
don, vol. 22, p. 518. Probably not of Guppy, 1867.
Acteocina recta Maury, 1917, Bull. Amer. Pal., vol. 5, p. 178, pl. 3,
sete ey
A small species, typically with a short exserted spire and a
broadly cylindrical form. The Costa Rican examples are all
less than 3mm in height. There is considerable variation in
the height of the spire, which may be sunken in some cases as
illustrated by the larger cozx-lacryma Guppy from the Miocene
of Venezuela and Martinique, to others with high spires and
strongly descending whorls as in the recent canaliculata Say.
However they all appear to belong to the same species. From
the young of bullata var. costaricana, they are at once distinguish-
ed by their smooth, plain surface.
Gatun Stage: Coll. 7, Estrella River.
Coll. 5, Red Cliff Creek.
206c Costa Rica M1ockENE—OLSSON 34c
Subgenus CYLICHNELLA, Gabb
Acteocina triticum-tritonis Maury
Cylichnella bidentata Gabb, 1873, (in part) Trans, Amer. Phil. Soc.,
vol. 15, p. 273. Not of d’Orbigny, 1845.
Cylichna bidentata Guppy, 1876, Quart. Journ. Geol. Soc. London, vol.
32, p. 518.
Acteocina (Cylichnella) triticum-tritonis Maury, 1917, Bull. Amer.
ale VOUS Py L7O. Pla siutie., Ae
This shell was separated by Dr. Maury from the recent 67-
dentata d’Orbigny, with which Gabb, Guppy and others had
identified it. In ¢vzticum-tritonts the anterior of its plications is
very small, the posterior one large, which is the reverse of the
conditions as seen in the living d¢dentata. ‘The base of the shell
is finely grooved with incised, spiral lines. ‘The specimens from
Costa Rica agree very closely with typical specimens from the
Miocene of Santo Domingo.
Gatun Stage: Middle Creek.
Genus VOLVULA, A. Adams
Volvula cylindrica Gabb
Volvula cylindrica Gabb, 1873, Trans. Amer. Phil. Soc.. vol. 15, p.
246.
ie cylindrica Guppy, 1876, Quart. Journ. Geol. Soc. London,
vol. 32, p. 518.
cf. Bulla (Volvula) cf. oxytata Toula, 1908, Jahrb. der K-K. Geol.
Reichsanstalt, Wien, vol. 58. p. 709, pl, 28, fig. 4.
Volvula cylindrica Maury, 1917, Bull. Amer. Pal., vol. 5, p. 180, pl. 3,
fig. 5.
This, is a small species of no unusual characters. It is nar-
rowly cylindrical in form, shortly produced and pointed at its
posterior extremity. It was described by Gabb from the Mio-
cene of Santo Domingo, but Dall united with it, the oxytata
Bush, in which case, it is also found in the recent fauna. Gabb
records acuta d’Orb. from the Pliocene of the Limon Peninsula.
Uscart Stage: Middle Creek.
Genus BULLARIA, Rafinesque
Bullaria paupercula Sowerby Plate 4, figure 14
34d BULLETIN 39 206d
Bulla paupercula Sowerby, 1849, Quart. Journ. Geol. Soc. London,
vol. 6, p. 52.
Bulla paupercula Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
246.
Bulla paupercula Guppy, 1874, Geol. Mag. London, p. 437.
Bulla paupercula Guppy, 1876, Journ. Geol. Soc. London, vol. 32, p.
518.
Bulla paupercula Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 3,
pt. 1, p. 18.
Bulla striata Dall, 1903, Trans. Wagner Free Inst., Sci., vol. 3, pt. 6,
. 1583.
ie ee Maury, 1917, Bull. Amer. Pal., vol. 5, p. 182, pl.
Balen Se
Bullaria paupercula Maury, 1920, N. Y. Academy of Science, Tertiary
Mollusca of Porto Rico., vol. 3, pt. I, p. 75.
This species is rare in Costa Rica, and all our specimens are
from the Lower Gatun rocks of Middle Creek. It is possibly
identical with the recent Aullaria amygdala Dillwyn, abundant
on the beaches along the northern coast of Panama and Costa
Rica.
Gatun Stage: Middle Creek.
Genus RINGICULA, Deshayes
Ringicula dominicana Maury
Ringicula semistriata ? Gabb, 1875, Trans. Amer. Phil. Soc., vol. 15,
p. 225. Not #. semistriata d’Orbigny.
Ringicula dominicana Maury, 1917, Bull. Amer. Pal., vol. 5, p. 185,
Oi ain semegy Nae,
This is the most abundant of the Opisthobranch mollusks in
the Miocene beds of Costa Rica. The whorls are in general more
strongly sculptured with incised lines than typical dominicana
from the Miocene of Santo Domingo, and the spiral lines extent
over the greater part of the shell,except for a wide, smooth band
about the suture.
The R. hypograpta Brown and Pilsbry,from Gatun is a larg-
er shell with a heavier outer lip and more crowded spiral lines
about the base. ‘The Costa Rican shells measure about 1.5 mm
or less in height.
Gatun Stage: Middle Creek. Coll. 5, Red Cliff Creek.
Coll. 7, Estrella River.
Ww
qn
207 Costa Rico M1ocENE—OLSSON
ORDER CTENOBRANCHIATA
(A) SUPER-FAMILY TOXOGLOSSA
Genus TEREBRA ApDANSON
Terekra subsulcifera Brown and Pilsbry
Terebra subsulcifera Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 339, pl. 22, fig. 7.
Terebra (Myurella) subsulcifera Cossmann, 1913, Jour. de Conchylio-
logie, vols 61, ps 14) ple tr, e.)25:
The single, fragmentary specimen comes from the Gatun
beds of Mt. Hope in the Canal Zone. From ézpartita, which it
resembles in its sculpture, this species and haztenszs Dall, differ
in having only a single columellar fold.
Gatun Stage: Mt. Hope, Canal Zone.
Terebracf. haitensis Dall Plate 1, figure 3
Terebra haitensis Dall, 1895, Proc. U. S. Nat. Mus., vol. 18, p. 35.
Terebra haitensis Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt.
- 6, p. 1632, pl. 59, fig. 30.
Terebra haitensis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 194. pl. 4,
185) Bie
The collection contains a single, imperfect shell from Sousci
Creek, which should probably be referred to this species. From
the swbsulcifera, it differs in its more numerous and oblique ribs,
and wider, less pronounced spiral bands. Like subsulcifera, the
columella carries a single, sharp plication. Our specimen con-
sisting of only 5 whorls measures 37 by 13 mm.
Gatun Stage: Sousct Creek, a branch of Fone Creek, C. R.
Terebra bipartita Sowerby Platemmmehourent, 2.
Terebra bipartita Sowerby, 1849, Quart. Jour. Geol. Soc. London, vol.
5; P- 47.
36 BULLETIN 39 208
Terebra bipartita Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
225.
Terebra (Acus) bipartita Dall, 1895, Proc. U.S. Nat. Mus., vol. 18,
p-. 38.
Terebra bipartita Maury, 1917, Bull. Amer. Pal., vol. 5, p. 187, pl. 3
fig. 14.
This species, possessing much the same type of sculpture as
sulcifera, subsulcifera, haitensis etc., is recognized by having two,
instead of one columellar plication. Full-grown specimens
have a bipartite sculpture while in sz/cifera and the others it is
tripartite.
The Costa Rican collection contains three specimens from
Saury creek, near Cahuita, the largest of Io incomplete whorls
mearures 77 by I9 mm.
Gatun Stage: Zone E. Saury Creek
Terebra gatunensis Toula Plate 1, figures 4-6
Terebra gatunensis Toula, 1909, Jahrb. der K-K Geol. Reichsanstalt,
vol. 58, p. 705, pl. 25, fig. 14.
Terebra gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phils.) volii63)) p.339pls.22sihige 2s
Terebra wolfgangi Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phil., vol. 63, p. 340, pl. 22, figs. 1, 3-6, but not of Toula, 1908.
Terebra (Myurella) gatunensis Cossman,1913,Jour. de Conchyliologie,
vol, 61, p. 13, pl. 1, figs. 26-29.
In the Canal Zone, the Gatun beds contain two common
species of Terebra, which were described by Toula as gatun-
ensis and wolfgangi. ‘These species have been more or less con-
fused by Brown and Pilsbry, so that their figures and description
of wolfgang?, really represent gatunensis. Their figure 2 isa
large and less common variety of gatunensis. ‘Toula’s figure, al-
though of a young imperfect shell, is clear and shows the main
209 Costa RicA MiockENE—OLSSON
Ww
“I
characters of the species, the spirally striated, faint sutural sul-
cus and fine spiral scu)lpturing of the whorls. He however in
his notes, allies this species with the recent 7. dislocada of Say,
which is misleading. Toula’s figure can be duplicated by scores
of specimens in our collection.
Terebra gatunensts is characterized by its large size, (length
50 mm or more), its faint sutural sulcus which may be smooth
or with 2 or 3 faint spirals, showing best on young shells. The
spiral cords of the main body of the whorl are variable in num-
ber, spacing and strength, as well as the longitudinal ribs. A
large coarse variety, with persistent sulcus, and heavier sculp-
ture occurs rarely and is shown by Brown and Pilsbry as their
figure 2. The pillar is nearly smooth, carrying a very broad
fold above, and a small one below.
Gatun Stage: Gatun and Mt. Hope, C. Z.
Terebra wolfgangi Toula Plate 1, figure 11, 12
Terebra wolfgangi Toula t909, Jahrb. der K-K Geol. Reichsandstalt,
vol. 58, p. 705, pl. 28, fig. 7.
Not Zerebra wolfgangi Brown and Pils. which is 7. gatwnensis Toula.
Although 7. gatunensis and wol/gangz have not been collect-
ed by us except in the Canal Zone, where they are abundant,
they are included here for the sake of completeness. It is also
not unlikely that future collecting may still reveal their presence
in the synchronous beds in Costa Rica and adjacent parts of
Panama.
T. wolfgangi differs from the common 7. gatunensis by its
long, tapering form, deep, persistent sulcus anda sutural band
generally free from spirals. It is also a much smaller species,
the largest specitnen in our collection measuring about 38 mm in
length.
Gatun Stage; Mt. Hlope, C. Z.
38 ; BULLETIN 39 220
Terebra gausapata Brown and Pilsbry Plate 1, figures 8, 10
Lerebra gausapata Brown and Pilsbry, April 1911, Proc. Acad. Nat.
Sci) Phila.) iveli163,)p./ 340) (pls 22) fies. (8. 9:
Tevebra acuaria Toula, Dec. 1911, Jahnb. der K-K Geol. Reichanstalt,
vol. 61, p. 19, pl. 2, fig. ro.
Tevebra (Myurella) acuaria Cossm., 1913, Journ. de Conchyliologie,
vol. 61, p. 12, pl. 1, figs. 19-24.
The 7. gausapata, was described by Brown and Pilsbry,
from young specimens, measuring only 19 mm in length. Later
in the same year appeared Toula’s 7. acuaria, also based on
young shells.
The important characters of this species, are its nearly
straight ribs, deep sulcus, a sutural band with faint spirals
and even sculpture. Figures 23 and 24 of Cossmann I believe
represents wolfgangt. T. wolfgang? as will be noted from the
accompanying figures is a much more slender species and usually
with a smooth sutural band. The pillar carries two broad folds,
which are better developed than in wol/gangi or gatunensts.
Length 33, breadth 8.5 mm.
Gaiun Stage; Mb. Tope (Cay
Water Cay.
Terebra spirifera Dall Plate 1, figure 13
Terebra dislocata Gabb, 1873, Trans. Amer. Phil. Sci., vol. 15, p. 225,
Not of Say.
Terebra (Acus) bipartita Sowerby, variety spirifera Dall, 1895, Proc.
U. S. Nat. Mus., vol. 18, p. 38.
Terebra (Oxymeris) bipartita Sowerby, variety spirifera Dall, 1903,
Trans. Wagner Free Inst. Sci., vol. 3, pt. 6, p. 1632, pl. 59, fig. 13.
Terebra spirifera Maury, 1917, Bull, Amer. Pal., vol. 5, p. 188, pl. 3
figs. 15, 16.
In 7. Gatunensis and wolfgang? the pillar is nearly smooth
Axi Costa R1icA M1rockNE—OLSSON 39
or with two board, sub-obsolete folds. In spzrifera, the two folds
are strong and recall those seen in the recent 7: dislocata. The
sutural band is strongly sculptured by the end of the ribs which
are entirely dislocated by the deep sulcus. Typical specimens
have three or four, even spirals, which are stronger than the
longitudinal ribs. Variation occurs in the number, and regu-
larity or these spirals,
Length 43, breadth 9 mm.
Gatun Stage: Banana River, Hill No. 1.
Water Cay.
Terebra spirifera Dall, variety midiensis, n. var. Plate 1 figure 7
This variety, represented only by a single specimen from
Middle Creek, C. R., differs from typical spzrifera, in having on-
ly three spirals, which are broad and strap-like in form.
Gatun Stage: Middle Creek. C. R.
Terebra Berlinerae Maury Plate 1, figures 17, 18
Terebra Berlinere Maury, 1917, Bull. Amer. Pal., vol. 5, p. 198, pl.
Als SONA, 9/3) le
This beautiful species was described by Dr. Maury, from the
Cercado (Miocene) formation of Santo Domingo. ‘The whorls
are slightly convex, and its sculpture-is closer, finer and more
delicate than is seen in any of the associated Costa Rican Tere-
bras. Our specimens, all imperfect, were collected from the
hard Gatun sandstones of Boucary Creek. The most complete
specimen measures 21 by 6.5 mm.
Gatun Stage: Boucary Creek.
40 BULLETIN 39 | 21zZ
Terebra estreliana, n. sp. Plate 1, figures 9, 16
Shell small; nucleus of three smooth tapering whorls; post-
nuclear whorls about 9, sculptured with a faint sutural sulcus,
which is lacking or nearly so from the early whoris; axial sculp-
ture of about eleven, straight or slightly oblique ribs which pass
from suture to suture; but are nearly lacking from the base;
spirals lacking or very faint on the spire-whorls, so that they ap-
pear smooth and solid; the base of the last whorl is strongly con-
tracted and shows four or five, very faint spirals; pillar straight
and smooth; beak twisted.
Length 7.5, diameter 2.5 mm.
This is a small species of rather solid appearance, strong
ribs, but otherwise nearly smooth whorls. The spirals are very
faint and are seen best only when viewed at right angles to the
source of light. Our specimen came from the Estrella River.
Gatun Stage: Zone 7, Estrella River, C. R.
Terebra costaricensis, n. sp. Plate 1, figure 19
Shell small and slender, with about three, long, tapering
smooth, nuclear whorls; post-nuclear whorls eleven; sutural
band rather large and prominent, strongly tuberculated; the
spire-whorls are slightly concave and have at their base or just
above the suture, a revolving series of tubercles, which are the
enlarged bases of the longitudinal ribs; longitudinal ribs on the
last whorl number about twelve, are nearly straight and large
and strong on each side of the suture, low and weak in the mid-
dle of the whorls; the surface of the whorls are otherwise smooth
or in some shells feebly spiralled; base slightly rounded with
eight or more spirals which become finer as they approach the
canal; the longitudinal ribs are continued across these spirals.
Length 13, diameter 3 mm.
The above description is based mainly on two specimens
214 Costa Rica M1ockNE—OLSSON Al
from Hill No. 3, of the Banana River. Our collection contains
several smaller shells from Zone 7, of the Hstrella River in
which the base is somewhat more heavily sculptured and with
very fine spirals over the main surface of the whorls.
Terebra benthalis Dall var. bocasensis, n. var. Plate 1, figures 25, 26
Cf. Terebra (Acus) benthalis Dall, 1889, Bulli. Mus. Comp. Zool., vol.
18, p. 65, pl. 29, fig. 6.
This species, fairly abundant in the shale beds, just north
of the city of Bocas del Toro, Panama, agrees closely with Dall’s
figure in the Blake Report, except that the revolving series of
tuberclesin the fossil shells seem to be lower and broader.
The whole surface on slight magnification is seen to be finely
spirally striated.
Terebra benthalis was described by Dall from specimens
dredged from Ioo to 400 fathoms of water off the Morro Light,
Havana, Cuba. The species belongs to the section Fusoferebra
Sacco.
Length 27, diameter 6.5 mm.
Gatun Stage: Bocas del Toro, Panama.
Terebra pumbriensis, n, sp. Plate 1, figure 27
Shell of medium size, composed of 9 whorls (type with a
small portion of the apex lost); the early post-nuclear whorls
(the 1st 3 or 4) with two revolving rows of tubercles, formed by
the intersection of fine, oblique riblets and two spiral bands; on
the 4th whorl of our specimen, a median spiral makes its ap-
pearance and rapidly increases in strength so that the later
whorls have three rows of tubercles, the upper or most posterior
one being somewhat the strongest; the longitudinal ribs are nar-
row and oblique, and on the last whorl number about 20; the
space between the ribs have three or four fine, incised growth-
lines; periphery of the base has an additional tuberculated spiral,
42 BULLETIN 39 214
below which is a band composed of four smooth, close spirals ; be-
tween these spirals and the keel is a deep, concave zone; pillar
nearly straight and smooth, with a single sharp keel on the
back.
Length 20, diameter 4.75 mm.
Of this interesting and distinctive species, we have only a
single, imperfect specimen from the Gatun beds of Pumbri
Creek. The three rows of tubercles are evenly spaced and there
is no sutural band.
Gatun Stage: Pumbri Creek, C. R.
Genus CONUS Linné
Conus molis Brown and Pilsbry Plate 2, figures 1, 2
Conus molis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila.,
VOl6gh pi 242 wplyi2guntomis
Conus molis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 200.
This is the common Cone in the Gatun beds of Panama and
Costa Rica and large specimens may reach a length of nearly 150
mim. ‘The upper surface of the whorls carry 5 or more fine
spirals and perfect examples of young shells have a few of the
early post-nuclear whorls finely coronated. Described by Brown
and Pilsbry from the Canal Zone, the species has also been col-
lected by us at several stations along the Panama coast between
Colon and Bocas, at Water Cay, and along the Banana River in
Costa Rica. A large specimen from Water Cay measures:
length 144 mm, width 81 mm.
Gatun Stage: Gatun, Water Cay, Banana River, ete.
Conus stenostomus Sowerby Plate 3, figure 2
Conus stenostoma Sowerby, 1849, Quart. Jour. Geol. Soc. London,
vol. 5 p. 44.
Conus stenostoma Guppy, 1866, Zd., vol. 22, p. 287, pl. 16, fig. 2.
Conus stenostoma Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
230.
i
ae
CH
Costa Rico MioceNE—OLSSON 43
Conus catenatus Guppy, 1876, Quart. Jour. Geol. Soc. London, vol. 32,
Pp. 527, in part b
Conus stenostomus Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3,
pt. 6, p. 1583.
Canus stenostomus Maury, 1917, Bull. Amer. Pal., vol. 5. p. 203, pl. 6,
fig. 4.
A species characterized by its sharp angulated shoulder,
concave upper surface of its whorls and attenuated anterior can-
al. Well-preserved examples showing a highly polished sur-
face and traces of yellow blotches of color markings were col-
lected quite abundantly in the coralline of Port Limon. The
species also occurs in Jamaica and Santo Domingo.
Gatun Stage: Port Limon.
Conus concavitectum Brown and Pilsbry
Conus concavitectum Brown and Pilsbry, 1911. Proc. Acad. Nat. Sci.
Philal vol. 62. p. 341, pl. 23. figs 54 6.
Conus (Lichoconus) concavitectum Cossmann, 1813, Jour. de Conchy-
liologie, vol. 61, p. 43, pl. 4, figs. 3, 4.
In adult shells, the spire whorls become concave. The
species is also recognized by its sharp shoulder and low spire.
It may be confused with varieties of proteus but the sharp should-
er angle and spiralled spire whorls will separate it. Our speci-
mens come from Water Cay, and measure 48 by 27 mm.
Gatun Stage: Canal Zone and Water Cay.
Conus proteus Hwass Plate 2, figures 3, 4
Conus proteus Wwass, 1789, Enc. Meth. vers, 1 pt. 2, p. 682.
Conus leoninus WHwass 1789, Enc. Meth. vers, 1 pt. 2, p. 683.
Conus proteus ? Gabb, 1873, Trans, Amer. Phil. Soc., vol. 15, p. 232.
? Conus Berghausit ? Gabb, dem, p. 232. Not of Hoernes, Foss. Wien.
Becki ly ii hige 38
Conus leoninus Gabb, 1881, Jour. Acad. Nat. Sci. Phila., vol. 8, 2nd
series, p. 359.
44 BULLETIN 39 216
Conus proteus Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 3, pt. 1,
p. 26.
Conus proteus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 206, pl. 6,
sifegy 10h
This isa common species in the Miocene and Pliocene beds of
Costa Rica and Panama. It is generally possible to distinguish
two forms, a smaller one which agrees with typical examples of
proteus in form and coloration (which is frequently preserved, )
and a larger and heavier type, like the so-called /eontnzus. ‘This
second variety may show several dark spiral bands in addition to
the square yellow blotches. Such markings we have not observ-
ed on recent examples of /eonznus.
Gatun Stage: Banana River, Port Limon, Water Cay. Plio-
cene: Monkey Point; Bocas.
Conus Veatchi, n. sp. Plate 2, figures 5, §
Shell rather large, heavy and broad, with a low, nearly flat
spire, but pointed, slightly projecting apex; spire-whorls about
Io in number, witha slightly raised border near the suture and
marked with fine spirals which on the last whorl number ro or
11; shoulder of the last whorl rounded or beveled; last whorl,
large and broad, nearly smooth above, except for the very faint
spirals, but with numerous, fine, irregular spirals on the canal;
pillar straight; aperture straight and narrow, slightly wider
below.
Length 43, breadth 28 mm.
The present collection contains two specimens, the larger of
which from Water Cay, serves as the holotype. The species is
very unlike any of the associated Gatun species in its low, near-
ly flat spire. The Conus demiurgus Dall, of the Chipola Mio-
cene of Florida, is a related species.
It is named for Dr. A. C. Veatch, Director of the Hxplora-
tion Work of the Sinclair Consolidated Oil Corporation, for his
217 Costa RicA MIocENE —OLSSON 45
interest and co-operation in obtaining the extensive collections
which form the basis of the present work.
Gatun Stage: Water Cay, Panama.
GOO On VE
Conus floridanus, var. costaricensis, n. var. Plate 3, figures 3, 9
Shell conic, witha moderately high spire of about eleven
whorls; profile of the spire flat or slightly concave to the long
projecting apex seen in well-preserved specimens; shoulder angle
rather sharp with the whorls slightly concave above between the
sutures; spire-whorls generally show 3 or more faint, raised
spirals and strongly arcuated growth lines; the last whorl below
the shoulder angle is usually nearly ‘smooth above (sometimes
with faint spirals) but with heavy regular spiral groovings to
the number of about fifteen on the canal; the canal is long, rap-
idly attenuated and usually somewhat flexed below.
Height 44, breadth 21 im.
Type from the Gatun beds of the Banana River.
This fossil, quite comimon in Costa Rica, should probably be
classed as a variety of the recent Conus floridanus Gabb, differing
only in always being somewhat more slender. They also agree
closely with Dall’s figure of Conus chifolanus Dall from the
Chipola Miocene of Florida.
Care should be taken in distinguishing this from C.
zmitator of the Canal Zone. In that species, the full-grown shell
is larger, with nearly straight sides and finer irregular spirals on
the canal.
Gatun Stage: Banana River; East Grape Point Creek.
Conus imitator Brown and Pilsbry Plate 2, figure 6
Conus tmitator Brown and Pilsbry, April, Ig11, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 342, pl. 23, fig. 4.
Conus Dallt Toula, Dec., 1911, Jahrb. der K-K Geol. Reichsanstalt,
Wien, vol. 61, p. 508, pl. 31, figs. 23 a-d.
46 BULLETIN 39 218
Conus (Lithoconus) Dalli Cossmann, 1913, Jour. de Conchyliologie,
VOM GI pA pls Baines 2 OniaT ama MAN TO SEIZE Sy
? Conus, Pall Maury, 1917, Bull! Amer Pale vol. 5) p.) 212), pl. gy
fig. 15.
The Conus Dalli Toula is doubtless the young of zmztator
described earlier in the same year by Brown and Pilsbry. Fig-
ure 6 is of a specimen from Mt. Hope which measures 51 by 2:
mim. From Costa Rica, the collection contains a series of per-
plexing Cones, which probably should be referred to this speces.
From costaricensis they differ in their nearly straight sides, more
irregular spiral groovings on the anterior canal, and they usually
show faint spirals over most of the body-whorl, which I have
not observed on costaricensis. A large shell from Zone C of
Comadre Creek near Cahuita, measures 65 by 29 min.
Gatun Stage: Mt. Hope and Gatun C. Z.
Water Cay, Panama.
Banana River, East Grape Point Creek,
Comadre, Middle Creek, C. R.
Conus recognitus Guppy Plate 2, figure 9
Conus solidus Sowerby, 1849, Quart. Jour. Geol. Soc. London, vol. 6,
p- 45, Not C. solidus Sowerby, 1841.
Conus solidus Guppy. 1866, Quart. Jour. Geol. Soc. London, vol. 22,
P. 287.0 pl. (26) fig.) 1.
Conus recognitus Guppy, 1867, Proc. Sci. Assoc. Trinided, p. 171, Har-
ris Reprint, Bull. Amer. Pal., vol. 8, p. 198.
Conus pyriformis Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 229,
Not of Reeve, 1843.
Conus vecognitus Guppy, 1876, Quart. Jour. Geol. Soc, London, vol.
32, Pp. 527.
Conus vecognitus Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt.
(CN OBInansgo rey
Conus recognitus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 209, pl. 7,
fig. 9.
This species is easily recognized by its well-rounded should-
er and appressed sutures. It is closely related to the recent C.
BLO Cosra Rica M1ockENE—OLSSON 47
pyriformis Reeve of the west coast of Central America but the
fossil shells have generally a lower and flatter spire. It is also
known from the Miocene of Santo Domingo, Jamaica and
Trinidad.
Gatun Stage: Banana River, Hill No. 3.
Conus limonensis, n. sp. Plate 3, figures 7, 8
Shell rather large, but delicate and rather slender; spire low,
concave to the slightly projecting apex; spire whorls Io or more,
nearly flat between the sutures or slightly concave due toa
slightly raised border; spire-whorls with 3 or 4 low spirals which
become faint on the later whorls so that they are nearly smooth;
sutures deep; shoulders of the last whorl angled or somewhat
raised; body-whorl long and slender, polished and smooth, ex-
cept fora few faint spirals near the tip of the canal; anterior
canal long and straight, evenly tapering.
Length 41, breadth 24 mm.
This species is based on 2 specimens from the coral lime-
stones of Port Limon, where it is associated with Conus stenost-
omus and planiliratus. Froim stenostomus, its straight canal, low
spire and nearly smooth, polished surface is most noticeable.
Gatun Stage.) Port Lemon, \C. R:
Conus musaensis, n. sp. Plate 1, figures 22, 24
Shell small and solid; spire short, conic, composed of about
2 pellucid, globular and projecting whorls and 6 post-nuclear
whorls; the spire-whorls are flat or slightly cancave and project
slightly over the anterior sutures; body-whorl with the upper
half smooth, but with 12 deep grooves on the anterior canal
which produce wide, spiral bands: these grooves are delicately
incised by raised longitudinal lines; color markings are some-
times preserved, which appear as 8 narrow, gray bands on the
“upper half of the whorl.
Ny
Length 19, breadth 9 mm
This small species, the Gatun analogue of the recent West
43 BULLETIN 39 220
Indian Conus Agassizt Dall, is abundant along the Banana
River and elsewhere in Costa Rica.
Gatun Stage: Banana River, Zone 6 Red Cliff Creek, C. R.
Conus cf. interstinctus Guppy Plate 3, figure 12
Conus interstinctus Guppy, 1866, Quart. Jour. Geol. Soc. London, vol.
22) DA 2oS ime MO. sh oa ah
A single specimen from Hill 1, of the Banana River, agrees
closely with Guppy’s figure of this Jamaican species. The shell
measures 39 by zo mm. ‘The spire is moderate in height, with
a slightly concave profile. The spire-whorls are smooth between
the sutures but also show a slight coronation. ‘The last whorl
carries about 15 narrow, widely spaced, spiral cords which an-
teriorly are slightly beaded.
Dr. Maury unites zzdersttuctus, with Sowerby’s catenatus,
but that species is based on a very young and scarcely distin-
guishable specimen. We therefore prefer to compare the Costa
Rican sheil with Guppy’s larger and well-figured zzterstinctus.
Gaiun Stage: Banana River, Hill No. 1.
Conus cf. Sewalli Maury
Conus Sewalli Maury, 1917, Bull. Amer. Pal., vol. 5, . 201, pl. 5, fig. 3,
pl. 6, fig 3.
The collection contains 2 young shells which may belong to
this Dominican species. They differ from the preceding zter-
stinctus in being shorter and broader, and the spire-whorls in
place of being smooth are marked with 5 or 6 strong spirals. The
last whorl is sculptured with 15 or more widely spaced beaded
spirals. The dimensions of the largest shell are: Length or
height 30, breadth 19 mm:
Gatun Stage: Banana River.
Conus toroensis n. sp. Plate 2, figure 7
Shell of medium size, with a conic spire a little more than
¥ the length of the aperture; the whorls of the spire numbering
2ar Costa RicA MI1ocRENE—OLSSON 49
12 plus are flat or slightly concave and bordered below by a low
carina projectly slightly above the suture; this carina on all ex-
cept the last whorl bears low nodes, about 20 to the later whorls;
the spire-whorls are otherwise smooth except for the arcuate
growth lines; the last whorl is a tapering cone, nearly smooth,
except for the faint spirals about its lower one-third; the aper-
ture is narrow, straight, with a thin arcuate outer lip.
Height 45, diameter 21, aperture 33 mm.
This shell belong to the Conus consobrinus group, agreeing
in its form and strongly nodulated spire whorls. The main
difference is that éovoensis is nearly smooth, the spirals showing
only on the lower one-third, while cozsobyinus has most of the
last whorl covered with beaded spirls.
Gatun Stage: Toro Cays.
Conus (Chelyconus) tortuosostriatus Toula Plate 1, figure 15
Conus (Chelyconus) tortuosostriatus Toula, 1911, Jahrb. der K-K.
Geol. Riechsandstalt, Wien, vol. 61, p. 508, pl. 31, fig. 22.
Conus (Hemiconus) tortuosostriatus Cossmann, 1913, Jour. de Conchy-
liologie, vol, 61, p. 40, pl. 3, figs. 28, 29.
Conus tortuosostriatus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 205, pi.
6, fig. 9
This shell should probably be considered as a short variety
of gracilissimus Guppy, differing chiefly in its broader form and
shorter spire.
The body-whorl is sculptured with about 24 raised,even, spiral
cords, separated by deep, even interspaces. These interspaces
are crossed by neat, incised lines, corresponding to the lines of
growth. ‘The nucleus consists of about 3, long, tapering smooth
whorls, after which follow several turns of carinated and finely
coronated post-nuclear whorls.
Length 22, breadth 9 mm.
Gatun Stage: Bocas del Toro.
50 BULLETIN 39 222
Conus granozonatus Guppy Plate 3, fig2 15
Conus granozonatus Guppy, 1866, Jour. Geol. Soc. London, vol. 22, p.
237) MplyTOw ioe
This is a rare species in Costa Rica, our collection containing
only a single specimen from the Gatun beds of the Rio Blanco.
It is characterized by a rather high, conic spire, with smooth or
only slightly coronated whorls. ‘The last whorl has about 18,
wide, spiral bands, which areheavily granulated above. The
specimen agrees well with Guppy’s figure of the Jamaican
shell.
Length 25, breadth rz mm.
Gatun Stage: Rio Blanco, C. R.
Conus planitiratus Sowerby Plate 3, figures 10, 13
Conus planiliratus Sowerby, 1849, Quart. Jour. Geol. Soc. London, vol.
6, p. 44.
Conus planiliratus Guppy, 1866, Sdem. vol. 22, p. 287, pl. 16, fig-
Tbe) P77
Conus planiliratus Gabb, 1873, Trans. Amer. Phil. Soc., p. 230, in
part.
Conus planiliratus Guppy, 1876, Quart. Jour. Geol. Soc. London, vol.
Aiea! ON) ENaLeNe
Conus planiliratus Dall, Trans. Wagner Free Inst. Sci., vol. 3, pt. 6,
Pp. 1583.
Couus planiliratus Cossmann, Jour. de Conchyliologie, vol. 61, p. 48,
PIs wmSs25h2onbore
Conus regularis Gabb, Jour. Acad. Nat. Sci. Phila., vol. 8, 2nd series,
p- 359, pl. 46, figs. 45-48. Not of Sowerby.
Conus planiliratus Maury, Bull. Amer. Pal., vol. 5, p. 209, pl. 7,
fig. 10.
The occurrence of this species in Costa Rica is limited to
the coral limestones of Limon, Gabb had specimens from here
which he identified with the recent West Coast Conus regularis
of Sowerby. The Limon shells agree very well with Guppy’s
figure in the Quarterly Journal and with Cossmann’s figures of
Bowden and Mindi specimens. The species appears to be closely
223 Costa Rico MirocENE—OLSSON 51
related to the recent Conus Stimpsont Dall from the Florida
Seats CbTOe NUE, Sah Nabi Mids.) voli 24m pa503,pluizon tion!) 7!
1903).
Surface of whorls are heavily sculptured with 25, 30 or
more subequal spiral cords. Upper surface of whorls, flat or
slightly concave and with 3 or more spirals. ‘The shoulder is
sharp and bounds a rather high, conic spire.
Length 31, breadth 16 mm.
Gatun Stage: Port Limon.
Conus planiliratus, var. hocasensis, n. var. Plate 3, fig. 14, 16
Shell of medium size, solid witha broad, stubby outline;
spire low, conic with nearly straight sides and composed of seven
or more whorls; the rst 4 or 5 post-nuclear whorls are coronated;
spire-whorls with 3 faint spirals and curved growth-lines; body-
whorl large and wide, with 25 strong but somewhat irregular
simple spiral cords; their interspaces are wide and with crowded,
raised lines of growth and sometimes intercalated spirals; anter-
ior canal nearly oe
Length 32, breadth 17 mm.
This Cones which should probably be considered as distinct,
is fairly common in the shale beds, belonging to the Gatun
Stage on Bocas Island. From typical planiliratus it GTS in its
broad stubby form and coarse sculpturing.
Gatun Stage: Bocas del Toro, Panama.
Conus marginatus Sowerby Plate 1, fig. 20
Conus marginatus Sowerby, 1849, Quart. Jour. Geol. Soc. London,
vol. 6, p. 44.
Canus marginatus Gabb, 1873, Trans. Amer. Phil. Soc., vol. 16, p.
230.
Conus marginatus Guppy, 1876, Quart. Jour. Geol. Soc. London, vol.
32, p. 528, pl. 20, fig. 5.
Conus marginatus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 210, pl. 7,
shoes
52 BULLETIN 39 224,
On the basis of Guppy’s figure in the Quarterly Journal,
this cone is seen to belong to the type exemplified by the recent
C. verrucosus, with its sulcated sides and appressed or beveled
shoulder angle. Our collection contains a single specimen from
Zone 5 of Red Cliff Creek which is very close to Guppy’s figure.
Cossmann’s specimen of marginatus from Martinique should
probably be referred to some other species.
Gatun (stages: Zone Gi ean Glas Graces GLiree
Conus gracilissimus Guppy Plate 1, figure 14
Conus gracilissimus Guppy, 1866, Quart, Jour. Geol. Soc. London,
vol. 22, p. 288, pl. 16, fig. 4.
Conus Orbignyi Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 230,
Not C. Ordtenyt Audonin, 1830.
Conus gracilissimus Gappy, 1876, Quart. Jour. Geol. Soc. London, vol.
32, P- 527+
Conus gracilissimus Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3,
DEVON spe SOs:
Conus gracilissimus Cossmann, 1913, Jour. de Conchyliologie, vol. 61,
Plea mess
Conus gracilissimus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 204, pl.
6, figs. 8.
A small, but long and slender cone, sulcated with 25 or
more, strong, regular spiral cords. The spire is long and slend-
er with slightly coronated whorls and the area between the su-
tures is delicately decussated by 3 or more spirals crossed by the
raised and arcuated lines of growth.
Our collection contains specimens from Bocas and Limon,
the largest measuring 26 by 9 mm.
Gatun Stage: Bocas del Toro, Pan., Port Limon, C. R.
Conus Burckhardti Bose Plate 3, figures 4, 5
Conus Burckhardti Bose, 1¥906, Bull. de Inst. Geol. de Mexico,
numero 22, p. 50, pl. 5, figs. 39, 40.
Shell of moderate size, witha rather high slightly concave
spire and long, acute apex; sutural whorls smooth, except for the
YI
W
225 Costa Rica MIocENE —OLSSON
faint arcuate lines of growth and very indistinct spirals; the
earlier whorls have a carinated suture but are only very faintly
coronated; last whorl heavily sculptured with about 20, wide,
even spiral bands, separated by grooves of about 1-3 of their
width; these grooves are faintly incised by the longitudinal
lines of growth; the spiral bands are slightly wider above
and smooth, becoming granulated on the canal; this granulation
appears on the posterior side of the bands only; aperture long and
narrow.
Length 36, diameter 14.5 mm.
33 (apex broken) diameter 15 mm.
The original specimen of Conus Burckhardti was described
from the Miocene of Mexico, State of Oaxaca. Our specimens
from Panama and Costa Rica area little larger, but show the
characteristic high, smooth, spire and strong, flat, spiral bands of
the body-whorl.
Conus tortuosopunctatus "Toula is a similiar but smaller
species. The difference in sculpture of the two species may be
seen in the accompanying figures, the spiral bands of fortuoso-
punctatus being double or incised, while in Burckhardt, they are
broad and simple.
Gatun Stage: Water Cay, Panama.
Zone 2, East Grape Point Creek.
Conus Harrisi, n. sp. Plate 3, fig. I
Shell of moderate size, subcylindric with a high, even, conic
spire; spire-whorls about 6 plus, flat, with close sutures, except
for the upper whorls which are slightly carinate; the surface of
the spire-whorls is nearly smooth, except for a few faint spirals;
the last or body whorl is strongly sculptured with widely spaced
and sub-regular grooves; these grooves form about 14 wide,
spiral bands about the lower 3-4th of the last whorl; the upper
ath below the shoulder angle forms a wide, smooth band; below,
the spiral bands are generally smooth except those about the lower
54 BULLETIN 39 226
half of the shell, which may become, mesially divided and bear
afew small granulations or beads about their upper half; aper-
ture sublinear.
Height 51, diameter 23 mm.
A single specimen of this elegant species was collected from
the Gatun of the Banana River. It is a species of the Conus
Burckhardt type but much larger and when perfect reaching a
length of nearly 60 mm. The sculpture is of wide bands formed
by deep, and regularly spaced grooves about the lower 3-4ths of
the shell.
Gatun Stage: Hill No. 3, Banana River.
Conus multistriatus Bose Plate 1, figures 21, 23
Conus Agasstzt Dall, variey multistriatus Bose, 1906, Bull. de Inst.
Geol. de Mexico, numero 22, p. 49, pl. 5, figs. 34-38.
Conus gaza Johnson and Pilsbry, 1g91t, Proc. Acad. Nat. Sci. Phila.,
vol. 63, pl. 23, figs. 2, 13.
Conus gaza | Maury,.)1917, Bulle) Aimer) kale, vol) '5;)) sp. ) 210.) play
Aigehen2)
This is the Conzws gaza of Johnson and Pilsbry described
from the Miocene of Santo Domingo and the Isthmus. It isa
small, pretty species, recognized by its nearly biconic outlines
and neat sculpture. The spire is high; with nearly smooth, spire-
whorls and carinated sutures. The last whorl is rather short and
heavily sulcated with about 18, regular, spiral cords, separated
by intervals of their own width.
All our specimens come from Water Cay, where the species
is fairly abundant. The largest specimen measures 20 by II
mm. Bose’s original specimens are from the Miocene of the
State of Oaxaca, Mexico. They are somewhat larger, measuring
Zieh) OMA), a's) panbaol yi
Gatun Stage: Water Cay.
Conus tortuosopunctatus Toula Plate 3, figures 6, 11
Conus tortuosopunctatus Toula, 1911, Jahrb. der K-K. Geol. Reichsan-
stalt, Wien, vol. 61, p. 507, pl. 31, fig. 21.
On
Ui
227 Costa Rica MrocknE—OLSSON
Shell small, with a high smooth, conic spire of about 12
whorls the earlier post-nuclear whorls are finely nodulated, the
succeeding have the sutures carinated and then simple; body-
whorl with 20 or more strong, beaded, spiral cords; three spiral
cords are usually double or incised with a median line, especially
on the anterior canal; the beading of the spiral cords is develop-
ed best on their posterior portions; interspaces of the cords near-
ly as wide as the cords themselves, smooth or with faint, longi-
tudinal lines and frequently a small median spiral.
Length 30, breadth 13 mm.
Gatun Stage: Mt. Flope.
FAMILY TURRITIDA
Genus TURRICULA Schumacher
SECTION SuRCcULA H. anp A. ADAMS
Turricula lavinoides, n. sp. Plate 4, figure 6
Shell rather large and moderately solid; whorls about 12 in
number, heavily sculptured with ribs and spiral cords; nucleus
of two smooth, slightly bulbous whorls; post-nuclear whorls in-
creasing uniformily in size; concave above, widest below the per-
iphery; suture strongly appressed with a spiral cord in front of
it; anal fasciole flat or concave, appearing nearly smooth except
for about three, small, widely spaced spirals; remainder of
whorls strongly sculptured with 8 or 12 axial ribs which are
lacking from the anal fasciole and from the base of the last
whorl; spiral sculpture of 3 or 4 strong cords on the spire-whorls,
becoming 10 or more on the last whorl in addition to those of the
anterior canal; finer, intercalated threads are present between
the main spiral cords on the later whorls; the interval between
the spirals is finely, longitudinally striated by long growth lines;
anterior canal long and nearly straight, slightly calloused along
56 BULLETIN 39 228
the inner lip; outer lip thin, with a shallow anal sinus, lying in
the sutural fasciole.
Length 54, diameter 20, last whorl 38 mm.
This elegant species may be compared with the 7. jaguensis
Sowerby from the Santo Dominican Miocene and the recent 7.
lavinia Dall from the West Coast of Mexico. The Dominican
species lacks the sutural cord, as well as differing in details of its
sculpture. 7. /avinia, appears to be more closely related, differ-
ing in its finer sculpture and proportions.
Gatun Stage: Hill ra, Banana River.
Turricula favinoides var. limonensis, n. var. Plate 4, figure 12
Shell of medium size and with about 10 whorls; nuclear
whorls about 2 in number, at first smooth, becoming finely rib-
bed on the Jast half; post-nuclear whorls spirally and axially
sculptured; anal fasciole concave, smooth, with a strong spiral
cord, just in front of the appressed suture; axial sculpture of
about 8 sharp or pointed ribs on the periphery of each whorl,
lacking entirely from the anal fasciole and from the base of the
last whorl; spiral sculpture of 3 cords on the early spire-whorls
becoming more numerous on the later, due to the intercalation
of additional spirals; on the last whorl, the spirals number about
22,including those on the canal; anterior canal long and straight;
anal sinus shallow; outer lip thin.
Length 47, diameter 18, last whorl, 32 mm.
67 27 47
This shell when full-grown appears distinct from the /avz-
noides but young specimens cannot be easily separated. This
variety differs from the typical form, by its sharp, narrow, and
widely spaced axial ribs, which produce a sharp, angled and
not rounded shoulder to the periphery of each whorl. The anai
fasciole is also wider and more smooth.
Gatun Stage: Toro Cays. Port Liman, coralline limestones.
229 Costa Rica M1ockNE—OLSSON 57
Turricula taurina, n. sp. Plate 5, figure 3
Shell small, slender and fusiform; mucleus of nearly two
smooth, convex whorls; post-nuclear whorls about seven; su-
tures appressed and bordered by a spiral cord; fasciole about
one-half the width of the spire-whorls, smooth; the sculpture of
the whorls is spiral, axial ribs lacking; on the spire-whorls there
are four raised spiral cords with the flattened interspaces about
three times the width of the spirals themselves; on the last whorl
and anterior canal, there are 18 or 19 spirals; anterior canal,
long, slender and strongly twisted or curved; outer lip simple;
anal sinus as indicated by the growth lines is wide, moderate-
ly deep and occupies the middle of the fasciole.
Length 25, diameter 8, last whorl 16 mm.
This isa small and delicate species, characterized by its
slender, fusiform outlines and lack of an axial sculpture. The
spirals are narrow, raised cords, rather widely spaced. The
single specimen was collected at Bocas del Toro.
Gatun Stage: Bocas del Toro, Panama.
Turricula Terryi, n. sp. Plate 5, figure 4
Shell small, slender and fusiform, with a long, evenly taper-
ing spire and slender, twisted anterior canal; nucleus of two,
smooth, small whorls, followed by 8 sculptured post-nuclear; su-
ture appressed, bordered by a low, but wide, spiral cord; upper
half or more of each spire-whorl, flat or slightly concave and
forms the fasciole; fasciole nearly smooth or sculptured with
few, low, faint spirals; early spire-whorls strongly sculptured
with 12 or 13 axial ribs which are most heavy on the periphery
of each whorl; these ribs become progressively weaker and on the
last whorl are only feebly developed on the shoulder or periph-
ery; spiral sculpture persistant but weak, consisting on the spire-
whorls of 4 cords on the lower half, alternating with much weak-
er spirals: the fasciole as already noted is nearly smooth, or very
weakly spiralled; on the last whorl, there are 20 spirals, includ-
58 BULLETIN 39 230
ing those on the anterior canal; aperture elliptical with a thin
outer lip and long twisted anterior canal.
Length 23.5, diameter 7, last whorl 15 mm.
The small exposure of Gatun shales, on the east shore of
Columbus Island, about one mile north of the city of Bocas del
Toro, has yielded a large number of small interesting gastropods.
More than a dozen species of Pleurotomids were collected at this
place by Mr. R. A. Terry and the writer during a short period
of collecting.
The 7. faurina, also from this locality, is somewhat similiar
but differs by its more convex whorls, deeper fasciole and prati-
cally lacks all axial sculpturing.
Gatun Stage: Bocas del Toro, Panama.
Genus TURRIS Bolton
Turris albida Perry Plate 4, figres I, 2
FPleurotoma albida Perry, 1811, Conch. Expl., pl. 32, fig. 4.
Fleurotoma virgo Wamarck, 1822, An. s. Vert., vol. 7, p. 94.
Pleurotoma haitensis Sowerby, 1849, Quart. Jour. Geol. Soc. London,
vol. 6, p. 50.
Fleurotoma virgo Moore, 1853, Quart. Jour. Geol. Soc. Lond., vol. 9,
Paso:
Fleurotoma barretti Guppy, 1866, Quart. Jour. Geol. Soc. London, vol.
22M ZOOMIPLe) V7 Aad Sa Os
Pleurotoma antillarum Crosse, 1865, Journal de Conchyliologie, vol.
13, p. 34, pl. I, fig. 8. Not of d’Orbigny.
Turris (Surcula) virgo Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15,
Pp. 206, 207.
Pleurotoma albida Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 3,
pt. 1, p. 28, pl. 4, figs.da.
Pleurotoma albida Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 343.
Pleurotoma haitensis Cossmann, 1913, Jour. de Conchyliologie, vol.
61, p. 16, pl. 2, figs. 1-4.
Pleurotoma cf. antillarum Cossmann, 1913, Jour. de Conchyliologie,
VOL OLN panei ple 2 wes s5 ho!
Turris albida Dall, 1915, Bull. go, U. S. Nat. Museum, p. 38, pl. 5, fig.
Tu DPV CA NATO NT)
223i Costa Rico M1iocENE—OLSSON 59
Turris albida Maury, 1917, Bull. Amer. Pal., vol. 5, p. 214, pl. 8, figs.
4-8.
This is a common species in the Gatun beds of Central Amer-
ica. ‘The usual form is a large, strongly sculptured and should-
er shell, which corresponds best to the variety described by
Guppy from Jamaica as darreééz. Our largest specimen from
Toro Cays, Panama measures as follows: Length 86, diameter
27, last whorl 56 mm. 7. a/dida, is alsoa common fossil in the
Miocene of Santo Domingo, Jamaica, Trinidad and Venezuela.
Its geologlcal range is from the Oligocene to the recent.
Gatun Stage: Gatun,C. Z.
Toro Cay, Water Cay, Panama.
Genus LEUCOSYRINX Dall
Leucosyrinx chloris, n. sp. Plate 5, fignre 19
Shell subfusiform, thin, with an acute spire and a long,
straight, anterior canal; nucleus small, globular of about 2,
smooth, convex whorls; post-nuclear whorls about 7; the spire-
whorls are strongly angled in the middle, leaving a wide, con-
cave or sloping zone about the upper suture; the sculpture con-
sists of small, more or less confluent tubercles on the periphery
of the whorls and number on the last whorl about 14; in addi-
tion, the whole surface is strongly sculptured with raised, sub-
equal, spiral threads; base strongly contracted tothe long,
straight, anterior canal; the outer lip is broken; anal sinus, as
indicated by the growth lines, lies in the concave zone, close to
the upper suture.
Length 14.50, diameter 5.00, aperture 7.25 mm.
This appears to be a true Leucosyrinx, characterized by its
thin shell, long, anterior canal and acute spire. It occurs in the
Bocas shales with Dyrillia lithocolletotdes, cocostna, Tevrebra benth-
alts var. bocasensts and others, shells which are all closely related
to recent deep-water species.
Gatun Stage: Bocas del Toro.
60 BULLETIN 39 232
Genus ANCISTROSYRINX Dall
Ancistrosyrinx elegans Dall, variety Plate 4, figure 17
Ancistrosyrinx elegans Dall, 1881, Bull. Museum Comp. Zool., vol.
9, P. 54.
Axcistrosyrinx elegans Dall, 1889, Bull. Museum Comp. Zool., vol-
LSM PU 2hp A, Ous ple soul See
The discovery of this elegant, recent species in the Gatun
beds of Costa Rica is of more than usual interest, the record
being based on a single, small but otherwise quite typical speci-
men from Hill 1a of the Banana River. Comparison with Dall’s
figure in the Blake Report, shows no important difference, ex-
cept that the sculpture of the fossil shell is somewhat finer.
A. elegans, is recorded by Dall from the Florida Reefs and
from 805 fathoms of water off Havana, Cuba.
The dimensions of our shell is as follows: Length 15, diam-
eter 5.75, last whorl 11 mm.
Gatun Stage: Hill ra, Banana River.
Ancistrosyrinx Dalli, n. sp. Plate 4, figure 16
Shell larger than the preceding and very nearly smooth; nu-
cleus very small and smooth; post-nuclear whorls seven or more
with a large and prominent, dentate or spinous keel; between
the keel and the suture, there is a wide, nearly flat area, which
carries a strong, smooth, median, spiral cord, behind which lies
the small concave anal fasciole; the rest of the shell is smooth
and without spirais;on the last whorl, the peripheral keel carries
about 18 short, broad, spiny teeth; the canal is long, slender and
straight; outer lip thin and fragile.
Length 14, diameter 7, last whorl 9.5 mm.
20 10, 15 mm.
Two specimens are represented in our collection, and the
smaller but more perfect specimen is selected as the Holotype.
It differs from the described species of Axncistrosyrinx by its
233 Costa Rica MioceENE—OLSSON 61
nearly total lack of spiral sculpturing. The species is named
for Dr. W. H. Dall of the National Museum, the author of the
genus Azncistrosyrinx.
Gatun Stage: Toro Cay, Water Cay, Panama.
Genus DRILLIA Gray
Drillia venusta, Sowerby Plate 4, figure 9
Fleurotoma venusta Sowerby, 1849, Quart. Jour. Geol. Soc. London,
VOl--65)p. 50; pl: Lo, fg. 7.
LPleurotoma venustum, Guppy, 1866, Quart. Jour. Geol. Soc. London,
VOls 22)" ps 280)
Turris (Drillia) venusta, Gabb, 1873, Trans. Amer. Phil. Soc., vol.
15, p. 208.
fleurotoma venusta, Guppy, 1876, Quart. Jour. Geol. Soe. London, vol.
Behe D 527):
Fleurotoma venusta, Guppy and Dall, 1896, Proc. U. S. Nat. Museum,
vol. 19, p. 305.
Drillia venusta Maury, 1917. Bull. Amer. Pal., vol. 5, p. 216, pl. 8,
BUSRS) AS TEA,
The Drillia venusta and consors of Sowerby, are the two
commonest Pleurotomids in the Gatun beds of Costa Rica.
Both are characterized by a neat, regular and reticulate sculp-
ture of spirals and axial ribs. In vwenusta, the anal fasciole is
narrow, and appears asa deep constricted zone, encircling the
upper part of each whorl. The spirals are narrow bands, pro-
duced between deep, incised lines and in typical specimens num-
ber on the last whorl about 18. Thecanal is rather long and
usually more or less twisted.
Drillia venusta, occurs also in the Miocene of Jamaica,
Trinidad and in Santo Domingo.
Gatun Stage: Gatun, C. Z.
Banana River, Port Limon.
Drillia consors, Sowerby Plate 4, figures 8, Io, 13
Pleurotoma consors, Sowerby, 1849, Quart. Jour. Geol. Soc. Londen,
WO, ©, {Os GOs
62 BULLETIN 39 434
FPleurotoma consora, Guppy, 1866, Quart. Jour. Geol. Soc. London,
vol. 22, p. 280.
Turris (Drillia) militaris, Gabb, 1873, Trans. Amer. Phil. Soc., vol.
15, p. 207. Not of Hinds, 1843.
fleurotoma consors, Guppy, 1876, Quart. Jour. Geol. Soe. London, vol.
32, Pp. 527-
Drithia militaris, Gabb, 1881, Jour. Acad. Nat. Sci. Phila., vol. 8, and
series, p. 337.
FPleurotoma alesidota, Dall, var. magina Bose, 1908, Bol. Inst. Geol de
Mexico, No. 22, p. 47, pl. 5, figs. 30, 31, 33, 45,
Pleurotama sp. aff. Pl. alesidota (Dall) var. mactlenta Toula, tort,
Jahrb. der K-K Geol. Reichsanstalt Wiea,vol. 61, p. 505, pl, 30, fig.
11. Not alesidota or macilenta Dall, 1889.
Drillia consors Brown and Pilsbry, 1911, Proc. Acad Nat. Sci. Phila.,
vol. 63, p. 345.
Drillia consors, Cossmann, 1913, Jour. de Conchyliologie, vol. 61, p.
20, pl. 2, figs. 8-14.
Drillia consors, Maury, 1917, Bull. Amer. Pal., vol. 5, p. 216, pl. 8,
figs. 15, 16.
This species is easily distinguished from the vezusta, by its
more slender form, wide but shallow anal fasciole, sharper sculp-
ture and straight, longer, anterior canal. ‘The most common
variety, illustrated by figure 10, averages in length about 35
mm. The axial ribs are numerous (about 28), straight or
slightly oblique and only slightly heavier than the raised spiral
cords. In addition the surface is overrun by fine and almost mi-
croscopic spirals. Figure 8 of a large variety from Grape Point,
has about 9 whorls and measures 60 mm in length. The early
spire-whorls have the usual sculpture as seen on typical cozsors.
On about the seventh whorl, the ribe increase suddenly to near-
ly twice their normal number. They gradually become obsolete,
so that on the last whorl, pratically only the spirals are left.
The fine, microscopic spirals seen on typical consors are lack-
ing.
The Drillia alesidota and its variety macilenta Dall of the
recent West Indian fauna, are closely related to consors.
235 ~ Costa RrcA MrockenE—OLSSON 63
Gatun Stage: Mt. Hope, C. Z.
Banana River, Port Limon.
Boras.
Brillia cf. moensis, Gabb
Drillia moensis, Gabb, 1881, Jour. Acad. Nat. Sci. Phila., vol. 8, 2nd
SCHIES, py 3514 Pl. 46.) fe. 33.
A single small speimen of a Drzllia was collected from the
coral limestones near Limon, which agrees partly with Gabb’s
figure and description. Our specimen has 7 instead of 8 ribs,
and lacks the sutura! spiral mentioned by Gabb. The dimen-
sions of this shell are: Length 12.5, diameter 4.5, last whorl
7 mm.
Gatun Stage: Port Limon.
Drillia papaya, n. sp. Plate 4, figure 5
Shell of medium size, slender and subfusiform; whorls 10
(mucleus eroded); anal fasciole, a deep concave zone, occupying
about 1-3 of a whorl interval and bordered above by a small su-
taral cord; sculpture of heavy, knob-like ribs, crossed by spiral
threads of different strengths; axial ribs about 6 on the last
whorl, lacking from the fasciole and from the base and anterior
canal; spiral sculpture of primary secondary and tertiary threads,
there being about four primary threads on each spire-whorl,
on the last whorl there are 8 or more primary threads, with an
intermediate secondary and finer tertiary threads on each side;
the anterior canal has numerous threads of varying strength;
aperture long and subelliptical; anterior canal long and slightly
bent backwards; anal sinus asindicated by the growth lines is
deep and wide.
Length 32, diameter ro, last whorl 19 mm.
A single specinien was collected from certain sandstones of
Lower Gatun age from a small tributary stream of the Rio
Cocles near Old Harbor, C. R. The recent Drillia haliostrephis
Dall from the Gulf of Mexico appears closely related, but the fos-
64 BULLETIN 39 236
sil shell has a more slender spire and slightly longer anterior
canal.
Gatun Stage: Rio Cocles, C. R.
Drillia cocees, n. sp. Plate 5, figure 7
Shell small, slender and fusiform; whorls Io or more, nu-
cleus eroded; fasciole deep and concave, about % of the width of
a spire-whorl, bordered above by a strong sutural cord, other-
wise smooth; sculpture of strong axial ribs, which are lacking
from the fasciole and from the base of the last whorl; these ribs
number on the last whorl about 10; the ribs are crossed by
strong, raised cord-like spirals, there being at first 2 on the earl-
iest spire-whorls and on the later 3; the last whorl shows 8
spirals and the wide interspaces are occupied by 3 or more small-
er ones; additional and more crowded spirals occur on the long
slender canal; anterior canal long and slender, slightly bent.
Length 21, diameter 6, last whorl 12 mm.
A small, very slender and fusoid species with strong, persis-
tent spiral and axial sculpture. A single specimen was collect-
ed at Coco Plum, a small cocoanut plantation about 40 miles east
of Bocas del Toro.
Gatun Stuge: Coco Plum.
Drillia chiriquiensis, n. sp. Plate 5, figure 2
Shell solid of medium size; whorls 7 (apex lost), of slightly
convex profile; anal fasciole small but quite deep and nearly
smooth; suture strongly appressed and bordered just below by a
strong, elevated spiral cord; sculpture of numerous, somewhat
oblique riblets and numerous weak spirals; the ribs commence
at the lower edge of the fasciole, pass slightly obliquely across
the whorls to the suture or on the last whorl across the base to
the canal; the ribs on the last whorl number 15 or 16; spirals of
numerous weak threads which are generally alernating in charac-
ter; the spirals are very small on the fasciole, quite large and
237 Costa RicA M1IocENE—OLSSON 65
coarse on the anterior canal; the outer lip is heavily thickened
by a large rib; anterior canal short and bent.
Length 25, diameter 8, last whorl 15 mm.
A single shell from Bocas, serves as type for this species. It
recalls the D. consors Sowerby, but differs in its much finer
sculpture.
Gatun Sage: Bocas del Toro, Panama.
Drillia citria, n. sp. Plate 5, fiyures-12, 13
Shell small; whoris about 8, of which the first 2 belong to
the small, obtuse, smooth nucleus; the post-nuclear whorls are
strongly sculptured with ribe and spirals; the upper 1-3 of the
spire-whorls carries a concave fasciole, bordered posteriorly by a
strong sutural cord to the rst post-nuclear whorl; the fasciole is
smooth or only faintly sculptured with fine spirals; the sculpture
consists on the last whorl of about 9 ribs which fade out rapidly
on the base and on the fasciole; the ribs are crossed on the early
spire-whorls by 2 spiral cords, becoming 3 on the later; on the
last whorl there are 12 more of which 6 are on the anterior ca-
nal; the intervals between the spirals, are 3o0r more times as
wide as the spirals themselves and are sculptured with fine
spirals threads; aperture sub-elliptical, with a short and slightly
twisted anterior canal; anal sinus moderate.
Height t1, diameter 4 mm.
A small species, common in the coralline phase of the Gatun
at Port Limon.
Gatun Stage: Port Limon.
Drillia aquanica, n. sp. Plate 5, figures 16, 17
Shell small, slender with a long spire and a shorter last
whorl; whorls about 11, the nucleus composed of 2 small, smooth
convex whorls and about 9 post-nuclear; whorls strongly sculp-
tured with ribs and spiral cords; fasciole nearly 1-3 the width of
the spire-whorls, marked with fine spirals and bordered postert-
66 BULLETIN 39 238
orly by a strong sutural cord; the spiral sculpture consists on
the early whorls of 2, later 3 and finely 4 cords on the penulti-
mate whorls; the last whorl has 8 spiral cords and 10 others on
the canal; the spiral cords are separated by interspaces of slight-
ly greater width; the interspaces are finely striated with small
spiral threads; the axial sculpture consists of about 9 knot-like
ribs over which pass the enlarged spiral cords; the ribs pass over
the base of the last whorl to the canal and slightly over the fac-
ciole; aperture elliptical with a short, straight canal.
Height 18, diameter 6, last whorl 9 mm.
This is a diminutive of Drillia fusiformis of the Miocene of
Santo Domingo, the two species having practically the same
sculpture. Drillia fusiformis of 11 whorls measures 35 mm in
length, while agzanica of the same number of whorls hasa
length of only 19 mm. The Maury collection contains a small
but quite typical specimen of aguanica from the Rio Gurabo,
Santo Domingo.
Gatun Stage: Water Cay.
Drillia limonica, n. sp. Plate 4, figure 15
In general form like Drillia consors Sowerby, with a long
spire and but slightly shorter anterior canal; whorls about 8% of
which 1% belongs to the small, smooth, blunt nucleus; the
post-nuclear whorls are sculptured with ribs and weaker spirals ;
on the 1st three whorls, the anal fasciole is but slightly differ-
entiated; it increases slowly in strength until on the later whorls
it has become a concave zone asin comsors; a sutural cord is in-
troduced on the 2nd post-nuclear whorl and is present on all the
succeeding whorls; on the spire-whorls there are about 11, wide-
ly spaced ribs; these ribs are lacking from the anal fasciole, ex-
cept on the very earlest whorls, where they are faintly continued
towards the upper sutures; on the last whorls, the ribs are small-
er and number about 13; between these main ribe on the last
whorl there appear occasionally smaller and shorter ribs which
do not reach to the edge of the fasciole; there are 5 faint spirals
239 Costa Rico MrockNE—OLSSON 67
crossing the ribs on the spire-whorls and very much finer ones in
the anal fasciole; the last whorl shows about ro spirals together
with others on the canal; aperture elongate with the anterior
canal rather long and straight.
Length 19, diameter 6.5 mm.
Like the Drillia consors in form, this species is distinguish-
ed by its fewer ribs and shallower anal faciole. The ribs on the
earlier whorls are oblique, but become very nearly straight and
vertical on the later.
Gatun Stage: Port Limon.
Drillia bocatoroensis, n. sp. Plate 5, figure 6
Shell of medium size, solid and strongly sculptured with
ribs and spirals; whorls about 11, the nucleus eroded in the type
specimen; fasciole occupying about 1-3 of the width of the spire-
whorl, with a large sutural cord above which the edge of the su-
ture is finely and regularly beaded or granulated; the surface of
the fasciole is slightly undulated by the obsolete ends of the ribs,
otherwise smooth; axial ribs about 8 on the last whorl, straight
and nearly in line across the face of the spire-whorls to the apex,
but interrupted and lacking from each fasciole; spiral sculpture
of the few raised cords with wider interspaces; there are 3
spirals on the spire-whorls, about 6 on the last, with 9 more on
the anterior canal; the interspaces in addition carry fine and sub-
microscopic spiral lines; the base of the last whorl is somewhat
constricted with a short anterior canal.
Length 23, diameter 8, last whorl 13.5 mm.
From the several species of fusoid Pleurotomids in the Gatun
beds of Panama and Costa Rica, this species differs in its heavy,
persistent ribs and sharp spiral sculpturing. The fasciole is
nearly smooth, bordered above by a heavy sutural cord.
Gatun Stage: Bocas del Toro.
68 BULLETIN 39 240
Driffia theobroma, u. sp. Plate 5, figure 11
Shell subfusiform; whorl 7 (apex broken); fasciole a narrow,
concave zone, not strongly differentiated from the rest of the
whorl, spirally sculptured and bordered above on the early spire-
whorls by a sutural cord; sculpture consists on the spire-whorl
of about ro low ribs, becoming more numerous and subobsolete
on the last; the spiral sculpture consists of raised, alternating,
revolving cords; there are 2 or 3 spiral cords on the early spire-
whorl, becoming 5 or more on the later; the last whori has about
10 spirals in addition to those on the anterior canal; between
each pair of the main or primary spirals there is a smaller, sec-
ondary spiral; base rounded or contracted tothe slender anterior
canal.
Length 21, diameter 7.5, last whorl 13 mm.
It is possible that this species, belongs to the genus Glypho-
stoma, as the type specimen seems to show a slightly thickened
and inflated outer lip. The aperture is however so completely
filled with a rocky matrix, that this observation cannot be fully
verified. "The axial sculpture is practically lacking from the
last whorl, leaving a surface simply marked with the rough
spirals.
Gatun Stage: Bocas del Toro.
Drillia aurantia, n. sp. Plate 5, figure 15
Shel! small, solid; whorls about 9; the nucleus consists of
about 114 smooth, convex whorls which through the gradual in-
troduction of the ribs, followed by the spirals blend in with the
post-nuclear whorls; the anal fasciole is rather wide (about 1-3
of the spire-whorl) and shallow; it uppears very early and car-
ries from the start a strong, sutural cord; the sculpture consists
of numerous closely spaced, linear ribs which are absent from
the anal fasciole; these ribs number on the last whorl about 21;
the spirals are uneven threads which are strongest below and on
the base: the fasciole also carries one or more fine threads except
24t Costa RicA MirocENE —OLSSON 69
on the base of the last whorl; the spirals do not cross the ribs so
that the summit of the ribs are left nearly smooth; aperture
small with a short, slightly twisted anterior canal.
Length 11, diameter 4 min.
A small species sculptured like the D. elegans Conrad of
the Chesapeake Miocene of Virginia, but with more numerous
and more closely spaced ribs.
Gatun Stage: Port Linon,
Drillia musacina, n. sp. Plate 5, figures 27, 28
Shell small, slender; the nucleus of about 2 whorls, the rst
blunt, convex and smooth, followed by the znd which is strongly
carinate; there are 7 post-nuclear whorls of slightly convex pro-
file; the fasciole is a narrow band on which the ends of the ribs
become strongly deflected to the left; the axial sculpture of about
16, narrow ribs on the last whorl; the ribs are continuous across
the spire-whorls to the upper sutures but become bent and
strongly deflected on the fasciole; on the last whorl, the ribs are
continued to the base of the canal; the spirals vary somewhat
in strength but are usually subregular spiral bands of which there
are 7 or 8 on the spire whorls and about 14 on the last whorl and
the canal; the back of the last whorl is strongly humped; the
outer lip was probably thin (broken in all our specimens); canal
short, straight.
Length 9, diameter 2.75, last whorl 5.00 mm.
A small species abundant in the Gatun beds of the Banana
River.
Gatun Stage: Hill ra, Banana River.
Hill 3, Banana River.
Drillia estrellana, n. sp. Plate Io, figures 31, 32
Shell very small, solid, porcellaneous; slender with a long
spire and a short body-whorl; nucleus small of 2 smooth, convex
70 BULLETIN 39 242
whorls; post-nuclear whorls about 7; sculpture with strong ribs
and impressed spiral lines in their interspaces; the anal fasciole
is a constricted band about the upper 1-4 of the whorl and which
passing over the ends of the ribs dislocates them asin Zerebra;
there are 11 ribs on the last whorl which commencing at the up-
per suture pass across the sutural fasciole but somewhat dimin-
ished in strength, continue across the face of the whorl ina
slightly oblique direction to the columellar region; the summits
of the ribs are smooth but the interspaces are sculptured with
impressed lines produced by spiral bands of regular width; there
are 4 such bands on the spiral-whorls and about 12 0n the last
whorl, including a few on the canal; aperture small, subellipti-
cal and a short twisted canal.
Height 7.5, diameter 1.75 mm.
A small species of Terebroid appearance, the ends of the ribs
being strongly dislocated as they pass across the small anal
fasciole.
Gatun Stage: Coll. 7, Estrella River.
Drillia limonetta, n. sp. Plate 5, figure 10
Shell small, solid, porcellaneous; whorls about 8, coarsely
ribbed and polished; fasciole absent; the sculpture consists of
moderately strong ribs which on the spire-whorls pass suture to
suture but are only slightly flexed near the upper suture; on the
last whorl, the ribs number about 11; the last rib is much en-
larged and forms the outer lip; the ribs are but shortly continu-
ed upon the base and do not reach the canal; in addition, the
surface is covered with fine, irregular threads which are nearly
obsolete on the upper portions of the whorls but are strong on
the base and on the canal; aperture subelliptical with a short
canal, somewhat bent or excavated ventrally; anal sinus simply
as a vertically directed notch at the junction of the outer lip with
the body-whorl and is bordered by a tooth-like denticle on its in-
ner side.
Length 12.25, diameter 5.00, last whorl 7.25 mm.
243 Costa RicA MrocKkNE—OLSSON al it
A small, solid and porcellaneous species, belonging to Dall’s
Cymatosyrinx. The sutural fasciole being absent, the ribs are
continuous to the upper suture and the surface is finely striated
with small, irregular spiral threads.
Gatun Stage: Port Limon.
Drijlia carruca u. sp. Plate 4, figure I1
Shell of medium. size, solid; whorls 7 plus (apex broken),
rather rapidly tapering; no fasciole; axial sculpture of about 8,
narrow ribs with deep and wider interspaces; the ribs commence
at the top of the canal and pass across the whorls to the sutures;
the ribs are nearly in line across the spire-whorls to the apex;
spiral sculpture of slightly elevated cords, separated by wide, flat
interspaces; there are about 7 spiral cords on the spire-whorl;
with 12 on the last whorl; aperture sub-elliptical, the outer-lip
somewhat thickened and a short canal.
Height 18, diameter 7.25, last whorl 11 mm.
This shell resembles Drz//ia musa in its general sculpture of
fine, widely spaced, spiral cords.
Gatun Stage: Coll.4, East Grate Potnt Creek.
Driiiia limonensis, n. sp. Plate 5, figures 8, 4
Shell of medium size and in form recalling D. venusta So-
werby; nucleus of nearly three, smooth convex whorls; post-nu-
clear whorls 8; anal fasciole scarcely perceptible on the carly
whorls, becoming on the later, simply a narrow, constricted zone,
bordering the suture; sculpture of rather numercus riblets, which
are abruptly bent to the left near the posterior suture on crossing
the constricted anal fasciole; three ribs number on the last whorl
about 15; the spiral sculpture consists of very fine threads; the
space between each pair of adjacent spiral threads is finely and mi-
croscopically decussated by the raised lines of growth, producing
a shagreening of the whole surface; in mature,shells, the last rib
becomes unusually large and heavy, producing a large and strong-
72 BULLETIN 3g 244
ly thickened outer lip; canal short, slightly bent inward or out:
wards; callous on the inner lip.
Length 20, diameter 7.5, last whorl 11 mm.
The general form of thissheill is that of Drillia venusta Sow-
erby, which it also recalls in its constricted anal fasciole. The
whole surface appears minutely shagreened, an effect produced by
the fine, sub-microscopic, spiral threads as well as a minute de-
cussation of their interspace. his species is fairly abundant in
certain sandy layers found intercalated in the hard and gener-
ally barren coral-reef limestone along the sea-front at Port
Limon.
Gatun Stage: Port Limon.
Drillia Cristobali, n. sp. Plate 5, figure r
Shell of the general form of wenusta, rather solid; whorls 6,
(apex broken in the type specimen); anal fasciole a narrow con-
stricted zone, bordering the upper suture; no sutural cord; sculp-
ture of numerous, straight, parallel ribs and fine spirals; ribs on
the early whorls about 12, becoming about I9 on the last; on the
anal fasciole, the ribs are shortly deflected or bent; spirals very
numerous and on slight magnification seen to consist of narrow,
even raised threads with interspaces of nearly twice the width of
the spirals themselves; anterior canal rather short and slightly
bent backwards; aperture sub-ellipitical with a thin outer lip;
anal sinus small and shallow.
Length 20, diameter 8.5, last whorl 11.5 mm.
This species, like the /Zémonensis, has the general form and
constricted fasciole of Drillia venusta Sowerby, but differs by its
smaller size, short canal and quite different spiral and axial
sculpture. JD. limonensis, is more closely related, but that
species differs in its more slender form and peculiar submicro-
scopic sculpture.
The single specimen serving as the type, was collected
from the Gatun shale, exposed about a mile north of the city
of Bocas del Toro, on Columbus or Cristobal Island.
245 Costa Rica MrockNE—OLSSON 73
Gatun Stage: Bocas del Toro, Panama.
Drillia bocasensis, n. sp. Plate 5, figure 5
Shell of medium size, solid; nucleus of about 2 smooth
whorls; post-nuclear whorls about 10; anal fasciole a narrow con-
stricted zone, nearly filled with a large, sutural cord; axial sculp-
ture of large, heavy ribs, about 9 on each whorl; the ribs pass
from suture to suture and on the last whorl acrcss the base to the
anterior canal; a large, hump-like rib is developed on the back
of the last whorl, formed during a resting stage; no spirals; the
growth-lines produce a minute but characteristic sculpture of
heavy, raised threads, which pass obliquely across the surface of
the whorls and ribs, and follow the curve of the anal
sinus across the fasciole; the sutural cord is finely granulat-
ed by the raised growth-lines; canal short but straight; outer lip
large, with a deep anal sinus at its posterior union with the
body-whorl.
Length 22, diameter 8, last whorl 12 mm.
This species should more properly be referred to the genus
Cymatosyrinx Dall, based on the Pleurotoma lunata H. C. Lea,
a Chesapeake Miocene fossil. The form and general sculpturing
of bocasensis is similiar to that of many species of this group but
it may be recognized at once by its very peculiar, submicroscopic
sculpture formed by the close, heavy growth-lines.
Gatun Stage: Bocas del Toro.
Drillia aquaensis, n. sp. Plate 5, figure 25
Shell small and solid; nucleus of 1+ whorls (mostly broken
on type specimen); post-nuclear whorls about 8; anal fasciole
lacking; axial sculpture of about six, large, heavy and straight
ribs, which pass from suture and across the base of the last whorl
onto the anterior canal; each set of ribs is in a straight line
from the canal across the spire-whorls to the apex; spiral sculp-
ture of about 8 impressed lines which produce a series of flat but
74. BULLETIN 39 246
quite wide, spiral bands; on the last whorl there are about 22
spiral bands; much finer near the suture, larger and wider on
the middle of the whorl and on the canal; canal short and slight-
ly bent to the left; inner lip calloused.
Length 15, diameter 5.5, last whorl 8.5 mm.
Characterized by its six axial ribs and finer sculpture of flat,
spiral bands. It belongs to the genus Cymatosyrinx of Dall.
Gatun Stage: Water Cay, Panama.
Drillla musa, n. sp. | Plate 5, figure 26
Shell about the same size as the preceding but witha
more slender spire; nucleus of 2 smooth whorls (broken); post-
nuclear whorls about 8; no fasciole; axial ribs about 9, pass
from suture to suture and across the base to the anterior canal
the ribs are in line across the whole series of spire-whorls; spirai
sculpture quite heavy, consisting of raised threads with wider in-
terspaces; the spirals number on the spire-whorls 8 or 9, and on
the last whorl about 22; aperture large and suboval; anterior
canal short, narrow and heavily calloused to form the inner lip.
Length 17; diameter 6, last whorl 9 mm.
Related to the preceding, but differs by its more slender
form; with g instead of 6 ribs and a much coarser spiral sculp-
ture.
Gatun Stage: Banana River, C. R.
Drillia lithocolletoides, n. sp. Plate 5, figure 18
Shell small, glassy or translucent; nucleus of about 3 smooth,
convex and glassy whorls, followed by 6 post-nuclear whorls; the
whorls are strongly angled in the middle by tubercular-like ribs
but leaving a concave zone about the upper sutures which forms
the anal fasciole; on the last whorl, the ribs or tubercles number
I1; are set obliquely and on the back of the whorl, they are con-
tinued slightly over the base; the surface is smooth and polished;
last whorl slightly constricted above the short, straight canal;
aperture subovate.
24.7 Costa Rico MioceNE—OLSSON
“I
wn
Height 10.75, diameter 3.25 mm.
The Drillia lithocolleta Watson is a_ recent deep-water
species, dredged by the Challenger and the Blake, at several
stations in the West Indiesand off the Florida coast, from depths
of 400 to nearly tooo fathoms. ‘The Bocas shell seems to be
very closely related to the recent species; differing only from
Dall’s figure in the Blake Report (plate 11, fig. 61), in being
somewhat more slender, with heavier tubercles and a longer base.
Gatun Stage: Bocas del Toro.
Drillia cocosina, n, sp. Plate 5, figure 14
Shell small, glassy or translucent in texture; spire twice or
more the length of the last whorl and the canal; whorls 8 plus,
the earlier ones missing; no anal fasciole; the sculpture consists
at first of a lower row of small tubercles, but a second or upper
row soon begins to appear and on the last whorl, this upper set
of tubercles is very nearly equal to the lower; the last whorl has
about 13 of these tubercles, in addition the whorl is finely sculp-
tured with fine, regular, closely spaced spiral threads which cov-
er the entire whorl and the canal, but leaving the tops of the
tubercles smooth; aperture ovate, with a short, twisted canal.
Length 12, diameter 3.5 mm.
This interesting species from the shales near the city of
Bocas del Toro, is related to the recent Dril/ia oleacitna Dall
dredged from rather deep water in the Gulf of Mexico and else-
wherein the West Indies. In differs from that species in being
less slender and with a larger body-whorl.
Gatun Sage: Bocas del Toro.
76 BULLETIN 39 248
Genus GLYPHOSTOMA Gabb
Glpyhostoma dentifera, Gabb.
Glyphostoma dentifera Gabb, 1872, Proc. Acad. Nat. Sci. Phila., vol.
24, p. 270, pl. 11, fig. 4.
Glyphostoma dentifera Gabb, 1873, Trans. Amer. Phil. Soc. vol. 15, p.
210. ;
Glyphostoma dentifera Dall, 1889, Bull. Mus. Comp. Zool., vol. 18,
pt. 2, p. 108.
Glyphostoma dentifera Cossmann, 1913, Jour. de Conchyliologie, vol.
61, p. 31, pl. 2, figs. 15, 16, 17.
Glyphostoma deniifera Maury, 1917, Bull. Amer. Pal., vol. 5, p. 225,
pl. 9, fig. 46.
A single specimen of this species was collected from the cor-
alline phase of the Gatun, near Port Limon. The shell showing
5 whorls (apex lost), measures 18 by 8mm. The pecular sha-
greening, characteristic of the genus is strong and easily seen.
This shell agrees closely with the figure given by Cossmann,
based on a Dominican specimen. It differs in several respects
from the specimen figured by Dr. Maury in her Dominican Fos-
sils from a metatype, sent by Professor Gabb to the Cornell Mu-
seum. Dr. Maury’s shell is larger, has a longer spire, and the
microscopic shagreening is much finer.
Dall in the Blake Report, doubtfully refers to this species,
an imperfect specimen collected in 15 fathoms of water at Sand
Keys.
Gatun Stage: Port Limon.
Glyphostoma moinica, n. sp. Plate 5, figures 29, 30
Shell small, solid with a coarse, tuberculate, subreticulate
sculpture anda thickened, enlarged onter lip; nucleus small,
pointed and tapering of 3 or more, smooth, convex whorls fol-
lowed by about 4 post-nuclear; the sculpture consists of about
'4 narrow ribs continuous from the canal to the upper suture
and crossed by even, spiral cords; their intersection form small
249 Costa RicA MrockNE—OLSSON 77
rounded beds or tubercles; immediately after the nucleus, there
is but one spiral, soon followed by two and on the penultimate
there are three; the last whorl has five spirals and three or four
more on the canal; aperture elliptical with a large expanded out-
er lip, smooth within and a deep, anal sinus at its upper end;
canal short and straight.
Height 5.75, diameter 2.75, last whorl 3.75 mm.
A small species doubtfully referred to Glyphostoma, having
the outer lip smooth within and lacking the peculair, submicro-
scopic structure of typical Glyphostoma.
Its sculpture is coarse, the intersection of the ribs and spirals
forming small beads or tubercles.
Gatun Stage: Port Limon.
Genus CYTHARA Schumacher
Cythara terminufa var. costaricensis, n. var. Plate 5, figures 21, 22
cf. Cythara terminula Dall, 1890, Trans. Wagner Free Inst. Sci., vol.
BODE IGHP srs On Pls 2e tis. 5:
The Cythara terminula Dall, to which the present shell ap-
pears closely related, was described from the Caloosahatchie
beds of Florida, of Pliocene age. The Costa Rican shell with
the same number of whorls is somewhat small (typi-
cal terminula of 7 whorls, 16 mim), (variety costartcensts 7 or 8
whorls, 13 mm), and has 8 instead of 9 ribs. The details of the
spiral sculpturing is very similiar, consisting of flattened bands
separated by sharp channels. The spiral bands carry a central
incised line, which produce the appearance of being in pairs.
C. cercadica Maury from Santo Domingo, is a larger species
with higher spire and simple and not banded spirals.
Length 13, diameter 5, last whorl 10 mm.
Gatun Stage: Fill ra, Banana River.
Cytharelia limata, n. sp. Plate 5, figure 20
Shell small, nearly smooth and porcellaneous; spire slightly
78 BULLETIN 39 250
shorter than the narrow aperture; nucleus of about 3% smooth,
convex whorls, the last 4% turn being very finely and closely rib-
bed; there are 3% post-nuclear whorls; sculpture consists of
narrow, slightly oblique ribs which pass across the spire-whorls
from suture to suture and on the last whorl follow down on the
anterior canal to its tip; there are 8 ribs on the last whorl; the
spaces between the ribs are wide, flat and smooth; the tops and
sides of the ribs themselves are carved or etched with fine, sub-
obsolete spiral lines; aperture linear-elliptical, the outer lip some-
what thickened by the last rib, but smooth within; anterior ca-
nal long straight.
Height 9, diameter 3.50, last whorl 7mm.
Gatun Stage: Port Limon.
Genus BORSONIA Bellardi
Borsonia coceensis, n. sp. Plate 5, figures 23, 24
Shell small, biconic; whorls about 7; nucleus of 2, nearly
smooth whorls, followed by 5 strongly shouldered post-nuclear
whorls, the angle of which on the spire-whorls comes just
above the lower suture; above the shoulder, the whorls
are flat or slightly concave to the upper suture; sculp-
ture with a double or triple beaded cord on the peri-
phery and with alternating beaded spirals both above and below;
on the last whorl, the primary spirals number 15 or 16, above
the shoulder 3 or 4; the suture is bordered in front by a strong
cord; pillar long, slightly twisted in front and with 2 sharp, pli-
cations; the anal sinus lies on the peripheral angle as in typical
Turris.
Length 16, diameter &, last whorl 11 mm.
The present species is based on two specimens fron: Coco
Plum about 30 miles east of the city of Bocas del Toro. The
B. varicosa Sowerby from the Dominican Miocene, is a related
species, differing in its porportionately longer spire.
Gatun Stage: Coco Plum.
25% Costa RicA M10cENE—OLSSON 79
Genus SCOBINELLA Conrad
Scobinella Morierei (Laville) Cossmann Plate 4, figures 3, 4
Euchilodon Morierei Vaville, in Cossmann, 1913, Jour. de Conchylio-
logie, vol. 61, p. 34, pl. 3, figs 6, 7.
Shell of medium size, with spire and last whorl of nearly
equal lengths; sculpture of numerous, close, bead-like spiral
cords, 3 or 4 on the spire-whorls, 18 or more on the last whorl
and canal: the sutural fasciole is rather narrow, shallow and
concave and with 4 much finer beaded cords; the suture is bor-
dered in front by a heavy beaded spiral, which on the early spire-
whorls, nearly fills the entire fasciole ; columella with four sharp
folds, the largest above; canal nearly straight; aperture narrow,
with a lirated outer lip.
Length 49, diameter 15, last whorl 33 mm.
This is one of the most elegantly sculptured Pleurotomoids
in the Gatun beds of Panama and Costa Rica. Described by
Laville and Cossmann from Mindi in the Canal Zone, we have in
addition collected the species from Toro and Water Cay, (where
it is fairly common), Bocas del Toro and from Limon. The pre-
dominating spiral sculpture and columellar folds are very sug-
gestive of Mitra.
Gatun Stage: Mindi, C. Z. (Laville and Cossmann) -
Toro aud Water Cays, Panama.
Bocas del Toro, Panama.
Port Limon, coral limestones.
Genus HALIA Risso
Halia americana, n. sp. Plate 4, figure 7
Shell large, thin, buccinoid in form; spire elevated with
shouldered whorls and deep sutures; whorls 3 plus, the earlier
ones missing; the profile of the whorls is convex with a wide,
shouldered, flattened or even slightly concave band just in front
of the suture; the surface of the spire-whorls is smooth and un-
sculptured but the early ones show very faint spiral bands; the
80 BULLETIN 39 252
upper half of the body-whorlis smooth but below, the whorl car-
ries about 16, low, faint, spiral ribbons like those of J/alea but
much less strong; these spiral ribbons average about 1.25 mm in
width; the growth lines cross the face of the whorl from the tip
of the canal to the shoulder angle in a slightly convex curve, the
convex side of which is towards the aperture; from the shoulder
angle to the suture the growth lines are reversed with the con-
cave side facing the aperture; aperture broadly elliptical; the ca-
nal was probably short and turned inward or ventrally.
Height 66, diameter 39 mm.
This truly remarkable species evidently belongs to the Pleu-
rotomoid genus Hala, of which the 4 known species are Euro-
pean and North African in their distribution. Its type species
flalia Priamwzs Meuschen is living off the coast of Spain and
North West Africa. According to Cossmann, there are three
Tertiary speciesin Europe, . precedens Pant. from the Helve-
tian of Italy, 7. Deshayeseana da Costa in the Tortonian of
Portugal and Italy, and AH. helicotdes Br. in the Plaisancian of
the maritime Alps.
The Huropean species are described as being entirely smooth,
but the Gatun shell is sculptured on its lower half with ribbon-
like spirals bands.
Gatun Stage Mi. Flope OZ
Genus CANCELLARIA Lamarck
Cancellaria dariena Toula Plate 6, figure 8
Cancellaria dariena Toula, 1909, Jahrb. K-K. Geol. Reichsanstalt,
Wien, vol. 58, p. 31, pl. 25, fig. 13, pl. 28, fig. 2.
Cencellaria dariena Brown and Pilsbry, I911, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 345, pl. 24, figs. 3, 4.
Cancellaria darienesis Cossmann, 1913, Jour. de Conchyliologie, vol.
61, p. 51, pl. 4 figs. 9, Io.
var. trachyostraca Brown and Pilsbry.
Cancellaria dariena, var. Toula, 1909, Jahrb. der K-K, Geol. Reich-
sanstalt Wien, vol. 58, p. 37, pl. 28, fig. 11.
253 CosTA RicA M10ocENE—-OLSSON 81
Cancellaria dariena var. trachyostraca Brown and Pilsbry, 191t, Proc.
Acad. Nat. Sci. Phila., vol. 63, p. 345, pl. 23, figs. I, 2.
This species is most abundant in the Canal Zone. It is dis-
tinguished from the Barretti Guppy, which it most closely re-
sembles by its shorter spire, larger body-whorl and a more irreg-
ular spiral sculpture. A large specimen from Mt. Hope, C. Z.,
has the following dimensions: Length 38, diameter 22; last whorl
30 mm. Shorter and more globose shells form the variety ¢ra-
chyostraca Brown and Pilsbry.
Gatun Stage: Gatun and Mt. Hope, C. Z.
Water and Toro Cays, Panama.
Veto Gocles\ Gunes
Cancellaria Barreitti Guppy Plate 6, figure 6
Cancellaria Barretti Guppy, 1866, Quart. Jour. Geol. Soc. London,
vol. 22, p. 289, pl. 17, fig. 11.
Cancellaria reticulata Gabb, 1873, Trans, Amer. Phil. Soc., vol. 15, p.
236. In part.
Cancellaria Barretti Guppy, 1876, Quart. Jour. Geol. Soc. London,
vol. 32, p. 520.
Not C. Barretti Maury, which is C. Maury@, n. sp.
This species is the West Indian Miocene analogue or the re-
cent Cancellaria reticulata Linné. ‘The most important differ-
ence to be noted between thetwo shells is that in Barretiz the
columellar plicee are more anteriorly situated and heavier. It
is a Miocene fossil in the Bowden beds of Jamaica.
Gatun Stage: Banana River
Cancellaria Cossmanni, n. sp. Plate 6, figures 9, II
Shell of medium size, solid; nucleus of 3, small, smooth
whorls; post-nuclear whorls 5, sutures deep; the whorls are
slightly channeled or coronated just below the sutures; sculp-
ture is evenly but coarsely reticulated by numerous, slightly ob-
lique, narrow ribs, crossed by heavy, raised, narrow, spiral cords;
the last whorl show 21 or 22 ribs; the spirals consists of narrow,
82 BULLETIN 39 254
raised cords, separated by wide, deep interspaces; there are 4
spirals on the spire-whorls, with 15 on the last whorl and the
canal; base of last whorl rounded and contracted to the short,
beak-like anterior canal; columella with 2, narrow, but strong
plications, of which the upper is much the strongest; aperture
sub-elliptical with the outer lip carrying 8, narrow, entering
liree.
Length 25, diameter 14.5, last whorl 18 mm.
This shell should probably be considered as a variety of C.
Larretti Guppy, but in the large series in our collection, the shell
is always much smaller and with a much coarser and heaver sculp-
ture. In some specimens, this sculpture becomes quite sharp
and harsh, brought about by the crossing of the narrow spiral
cords and equally narrow axial ribs.
It is an abundant and characteristic species of the Gatun beds
along the Banana River, C. R.
Gatun Stage: Banana River.
Cancellaria Maurya, n. sp. Plate 6, figure 5
Cancellaria Barretti Maury, 1917, Bull. Amer. Pal., vol. 5, p. 226, pl.
10, fig. 1. Not of Guppy, 1866.
Shell large, ovate; an evenly conic spire in height less than
the length of the aperture and a large, evenly convex last whorl;
the nucleus is rather small, of 2 smooth, convex whorls followed
by about 6 post-nuclear whorls; the profile of the spire-whorls is
slightly convex with deep, distinct suture; the sculpture is can-
cellate or reticulate, the spirals and ribs of very nearly equal
strength; of the axial ribs there are 40 or more on the whorl,
crossed by 24 spiral cords; on the spire-whorls there are 5 spiral
cords becoming -7 on the penultimate; the spirals are fairly regu-
lar, separated by interspaces 1% times their width, in which
smaller spirals may occasionally appear; faint indications of rest-
ing marks show as crowding of the ribs or as smooth, slightly
humped spaces; aperture large, ovate; a thin outer lip with 17,
long entering lireze; columella with 2 simple plicee, of which the
255 Costa Rico MrocENE—-OLSSON 83
upper is much larger; a third is probably developed on the an-
terior border of the canal; a well marked ridge about the base
and a strong siphonal fasciole.
Height 36, diameter 23, last whorl 29, aperture 24 mm.
This shell was figured by Dr. Maury in her Dominican fos-
silsas C. Barretti of Guppy, but it differs from that species in
much shorter spire, more globose shell, larger aperture and by
its columellar plicze. In the species closely related to the veéz-
culata, such as Barreiti, Cossmanni and others, the columellar
plicze are heavy and the posterior one is usually more or less
bifid. In this species, the posterior plication is large, wide and
thin, especially in the interior of the shell.
The Costa Rican collection is limited to a single, imperfect
specimen, so we have used as the type of this species, the origi-
nal figured specimen of Dr. Maury from the Miocene of Santo
Domingo.
Gatun Stage: Water Cay.
Cancellaria epistomifera Guppy
Cancellaria Moorei Gabb. 1872, Trans. Amer. Phil. Soc., vol. 15, p.
236. Not C. Moorei Guppy, 1866.
Cancellaria epistomifera Guppy, 1876, Quart. Journ. Geol. Soc. Lon-
don, vol. 32, p. 520, pl. 28, fig. 9.
Cancellaria epistomifera Cossmann, 1913, Journ. de Conchyliologie,
vol. 61, p. 53, pl. 4, figs. 5, 6.
Cancellaria epistomifera Maury, 1917, Bull. Amer. Pal., vol. 5, p. 227,
plo ess 35) 47 (Se
For figures of this species, the reader is referred to Dr.
Maury’s Dominican Fossils. We have collected a few, small
aud imperfect specimens at Gatun in the Canal Zone. Small
shells are easily confused with the common C. darzena, but in gen-
eral the spire is shorter, and the spiral sculpture is heavier and
more regular. At maturity, the outer lip develops below a
peculiar and characteristic sulcus or pout, which is not seen on
any of the associated species of Cancellaria.
Gatun Stage: Gatun, C. Z.
84 BULLETIN 39 256
Cancellaria Rowelli Dall Plate 6, figure 7
Cancellaria Rowelli Dall, 1898, Proc. Acad. Nat. Sci. Phila., vol. 19,
Doze, DIO. fetn
Cancellaria Rowelli Maury, 1917, Bull. Amer. Pal., vol. 5, p. 227, pl.
16, fig. 2
Two specimens of a Cancellaria which seem to be this spec-
tes, were collected from the Gatun beds of East Grape Point
Creek. ‘The axial ribs are numerous, oblique and somewhat ir-
regular. The spaces between the ribs are finely spiralled with
deep, incised lines, producing narrow and regular spiral bands.
The columellar plications are large and more or less bifid. Shell
measures:
Length 30, diameter 18, last whorl 21 mm.
The Cornell Museum contains, in the Gabb collection, a
single specimen which is probably this species. It agrees with
the Costa Rican shells. This species was described by Dall from
the Miocene of the Rio Amina, Santo Domingo.
Gatun Stage: Collection 2, East Grape Creek, C. R.
Subgenus TRIGONOSTOMA Blainville
Canceliaria toroensis, n. sp. Plate 6, figure 4
Cancellaria ( Trigonostoma) afi. Cancellavia bullata Sowerby, Toula,
1911, Jahrb. der K-K. Geol. Reichsanstalt, vol. 61, p. 504, pl. 30,
fig. 10.
Shell with 6 large, expanding whorls; the nucleus of 2%
whorls is small, closely coiled and smooth; the succeeding 2
post-nuclear whoris are small, convex and finely reticulated with
6 spirals and fine ribs; the last two whorls are deeply channeled
or excavated along the suture; this excavated band is smooth,
except as crossed by oblique growth lines; the last whorl is
somewhat contracted above and between this contracted zone and
the sutural channel is a large, heavy, more or less bifid spirai
cord. ‘This spiral cord is rendered coarsely and sharply nodulose
by the axial sculpture; the sculpture consists of about 21, narrow,
low, oblique riblets crossed by strong spiral cords; the spirals are
257 Costa Rica MIOCENE —-OLSSON 85
rendered slightly nodulose by the crossing of the longit udinal ribs
the penultimate whorl shows 6 or 7 spirals and 14 or 15 on the
last whorl; the contracted band around the upper part of the whorl
is nearly free from spirals, but is bordered above as already not-
ed by the large, bifid and strongly nodulose spiral cord; base
deeply but narrowly umbilicate, spirally sculptured within; col-
umella with 3, oblique plications; inner lip with a wide callus,
finely pustulated; the outer lip is broken in the type.
Length 29, diameter 20, last whorl 23 mm.
This is the Miocene analogue of the recent West Coast C.
tuberculosa Sowerby. It is probably the species figured by Toula
from the Canal Zone and which he compared with C. dullata
Sowerby. More recently Pilsbry and Johnson have described as
C. tnsularis,a Trigonostoma from the Miocene of Santo Dom-
ingo and which they consider as possibly identical with Toula’s
Isthmian specimen. In their description they mention but 14
ribs while toroenszs has 21 and they relate their species with the
Chesapeake Miocene perspecttva Conrad and the Tampa depressa
Dall, species which belong to an entirely different group.
Gatun Stage:
Toro Cays, Providence of Bocas del Toro, Panama.
Cancellaria Plummeri, n. sp. Plate 6, figures 2, 3
Shell thin and delicate, with large, loosely coiled whorls;
the whorls are prominently shouldered and deeply channeled or
excavated along their upper sutures; nucleus of three small,
smooth whorls; post-nuclear whorls 4; the sculpture consists of
very fine and delicate spiral threads or lines and three rows of
slightly elevated spine-like elevations or small tubercles: the
larger of these rows is found along the shoulder of the whorls
bordering the excavated zone, a smaller on the middle and a
small scarcely noticeable one below; on the penultimate whorl,
only two rows show and on the upper spire-whorls but one; the
umbilicus is deep, but narrow: the interior is concealed in the
matrix.
86 BULLETIN 39 258
Height 34, diameter 26, last whorl 17, aperture 19 mm.
This is a very distinctive species, characterized by its thin,
delicate shell and fine sculpture. It may be compared with the
recent 7. bullata Sowerby from the Pacific coast of Central
America, which differs in its larger, more expansive body-whorl
and coarser sculpture. The single specimen was collected from
the lower part of the Gatun formation on Toro Cays, in the east-
ern part of the Chiriqui Lagoon.
It is named for Mr. Fred Plummer of the Royal Dutch Oil
Company of Hague, Holland.
Gatun Stage: Toro Cays.
Subgenus APHERA H. and A. Adams
Cancellaria islacolonis Maury Plate 6, figure 12
Cancellaria tessellata Gabb, 1873, Trans. Amer. Phil. Soc. vol. 15, p,
236. Not of Sowerby, 1832.
Cancellaria islacolonis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 229.
DI Noy Mies 2 haeaD)
The C. zslacolonis was selected by Dr. Maury, as the guide
fossil for the lower of her Miocene formations, the Cercado. It
is a very rare fossil in Costa Rica and we have collected it only
along Red Cliff Creek, in the eastern part of the Republic.
There it occurs in the upper part of the Uscari and in the lower
part of the Gatun.
It is closly related to the recent West Coast C. ¢esselata
Sowerby.
Uscavi formation, Coll. 6, Red Cliff Creek.
Gatun formation; Coll. 4, Red Cliff Creek.
259 Costa RicA MrocENE—OLSSON 87
(B) SUPER-FAMILY RHACHIGLOSSA
Genus GLIVA Bruguiere
Olives are among the most common fossils in the Miocene
beds of Panama and Costa Rica. ‘The various species, however
show so few distinguishing characters, that their identification
becomes at times, both difficult and uncertain. The following
key has been prepared showing the characters which have been
most relied upon for their separation. ‘This key should be used
in conjunction with the figures.
A. Aperture narrow and of about the same width along its
whole length. Inner lip finely and regularly crenulated
above.
B. Sutures deep or channelled, with the edge of the
whorl projecting slightly above; spire of medium
height, length 35 mm.
O. sayana vat. immortua Pilsbry & Brown
BB. Sutures not channelled and with the upper edge of
the whorl beveled or appressed.
C. Spire generally low, with concave profile and
usually a projecting and globular nucleus.
Length 35 mm. O. drevispira Gabb
CC. Spire high or short, with conic sides; nu-
cleus or protoconch not large or prominent.
Di Spire’ high) jandiconie.) shell) wlansxe!
Length 40 mm or more
O. cylindrica Sowerby
DD. Spire short, shell usually small. Length
rarely over 30 mm.
O. gatunensts Toula
AA. Aperture wide and usually expanded and flaring in
front; inner lip is not crenulated above. (Agaronia)
B. Spire long, about 1-3 of the length of the shell;
88 BULLETIN 39 261
inner lip not strongly calloused. Lenght up to
45 mm. O. mancinella, n. sp.
BB. Spire shorter, about 1-4 of the length of the
Shell; inner lip usually strongly callused.
Length about 40 mm.
O testacea vat. costaricensts, 1. var.
Oliva cylindrica Sowerby Plate 7, figure 1
Oliva cylindrica Sowerby, 1849, Quart Jour. Geol. Soc. Londou, vol.
6, p. 45.
Oliva reticularis Guppy, 1866, Quart. Journ. Geol. Soc. London, vol.
22, p. 288,. Not of Lamarck.
Oliva cylindrica Gabb, 1873, Trans. Amer. Phil. Soe., vol. 15, p.
215.
Oliva cylindrica Guppy, 1876, Quart. Journ. Geol. Soc. London, vol.
BOY Os \ ESPXO)
Oliva cylindrica Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt.
6, p. 1583.
Miva cylindrica Cossmann, 1913, Journ. de Conchyliologie, vol. 61, p.
SP lenS iu tis Seu Dui at
Oliva cylindrica Maury, Bull. Amer. Pal., vol. 5, p. 231, pl. 10, figs.
TAN Aal,
A single, large specimen from Costa Rica, is doubtfully re-
ferred to this Dominican Miocene fossil. It differs from the more
common 6revispira by its larger size and longer, more conic
spire. This shell is also closely related to the recent O. avaneosa
Lamarck of the West Coast.
Length 47, diameter 19 mm.
Gatun Stage: Upper Gatun beds, 1 mile south of shore, alton:
Old Man Sam Creek, C. R.
Oliva brevispira Gabb Plate 7, figures 2, 3, 4
Oliva brevispira Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
215.
Oliva brevispira Maury, 1917, Bull. Amer. Pal., vol. 5 p. 232, pl. 10,
figs. 16, 17.
26k Costa Rica MroceNE—OLSSON 89
variety Giraudi Cossmann
Oliva Giraudi Cossmann, 1913, Journ. de Conchyliologie, vol. 61, p.
56, pl. 5, figs. 4-8.
A fairly common species in Cost Rica. Typically the shell is
short; moderatly convex, a short projecting spire and concave
spire-whorls.
Unusually broad forms as illustrated by figure 4 may be sep-
arated by the name of Girvaudz Cossmann.
A large typical shell measures:
Length 35, diameter 17 mm.
Gatun Stage: Coll.6, Red Cliff Creek, C. R.
fleadwaters of Middle Creek, C. R.
Cols Grape Point CreckanGs ies
Oliva gatunensis Toula Plate 7, figure 5
Oliva gatunensis Toula, 1909, Jahrb. der K-K. Geol. Reicesanstaldt,
vol, 58, p. 702, pl. 25, fig. 12.
Oliva reticularis gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat.
Sci. Phila., vol. 63, p. 348.
Oliva gatunensis Cossmann, 1913, Jour. de Conchyliologie, vol. 61, p.
58, pl. 5, figs. 9-12.
A common species at Gatun. The largest specimen in our
collection measures 28 mm. in length but Brown and Pilsbry men-
tion shells of an length of 35-38 mm. Oltva gatunensis, resem-
bles somewhat the drevirsozra of Gabb, but has a more even con-
ic spire and lacks the projecting nuclear tip. The sutures are
appressed. Brown and Pilsbry, place this shell as a variety of
the larger, recent Oliva reticularis Lamarck.
Gatun Stage: Mt. ope, | C.)Z,
Oliva sayana var. immortua Pilsbry and Brown Plate 7, figs. 6, 7
Oliva sayana var. immortua Pilsbry and Brown, 1917, Proc. Acad.
Naty Set Phtlar on 3a ui ple 15, fganG:
The Oliva sayana Ravenel (Oliva literata of most writers)
go BULLETIN 3g 262
is acommion recent species along the eastern coast of United
States, becoming replaced in the West Indies by the Oliva reiz-
cularis Laniarck. These two species are principally distinguish-
ed from each other, in that the sutures of O. sayana are quite
deeply channelled, above which projects the upper edge of the
whorls while in reticularis the sutures are merely deep with
rounded or beveled shell margins.
O. sayana var. tmmortua was described by Pilsbry and
Brown, from Gatun beds in the vicinity of Cartagena, Columbia.
What appears to be the same shell is common in the exposures
found along the lower part of the Banana River. Like the drev-
estria it is rather darkly colored, but no trace of a pattern is pre-
served. Our specimens averaged in length about 36 mm. The
largest specimen measures 41 by 19.5 mm.
Gatun Stage: Banana River, C. R.
Oliva testacea Lamarck, var. costaricensis, n. var. Plate 7, figs. 12, 13
Shell rather solid, with a moderately projecting spire, about
1-4 or less of the total length of the shell; whorls about 5, the
last, large, convex and widest about the middle; the spire-whorls
appear as if mesially divided by an encircling weak callus about
the lower halt; aperture broadest below, with the inner lip heav-
ily calloused above near its junction with the outer; the inner lip
is non-denticulated above, along its lower part with 3 or more,
irregular oblique, plaits; a broad band, arises from about the mid-
die of the inner lip, passes obliquely downward over the back of
the shell to the lower end of the outer lip.
Length 42, diameter I9 mm.
‘This Oliva is abundant in the Gatun beds of the Banana
River, C. R. It is closely related to the recent O. festacea Lam-
arex but seems to differ in being wider, shorter and with a lower
spire.
The Oliva testacea Lamarck (in part O. hiatula of some au-
thors) is abundant along the Pacific coast of Panama, and in the
Manuals, its range is given as the West Coast. Recently a close-
263 Costa Rrco MiocENE—OLSSON 91
ly related if not identical form, was collected by us from the
Pleistocene at Almirante, Panama, and several broken specimens
from the beaches at Bocas del Toro, Panama, and from Manzan-
illa, C. R. This recent Atlantic form differs only from the typ-
ical West Coast /estacca in being somewhat more slender and ap-
parently differently colored.
Tryon in the Manual of Conchology unités /estacea with
hiatula Gmelin from West Africa.
Gain Stage: Banana River, Coll 520 of, \Redp- Clif,
(Cgaliox (ON IKE
Oliva mancinella, n. sp. Plate 7, figures 8, 9
Shell slender and more delicate than the preceding; spire
long and pointed and about 1-3 of the total length of the shell; 5
whorls; sutures deep; the spire-whorls have their lower half
thickened by a smooth, encircling band of callus; aperture broad-
est about the lower half; inner lip smooth above, with only a
small and weak callus at its junction with the outer lip; the out-
er lip carries below, 3 or 4 heavy and very oblique plaite; the
outer lip is smooth and sharp.
Length 48, diameter 11 mm.
This species differs from O. ¢estacea and its variety costaricen-
sts by its much more slender and delicate shell. The porpor-
tions of the spire are about 1-3 of the total length of the shell.
Gatun Stage: Coll. 4, East Grape Potnt= Creek, C. R.
Genus OLIVELLA Swainson
A. Shelllarge, length 20 mm. or more; spire long and about
¥% the total length of the shell.
Olivella goliath, n. sp.
AA. Sheli smaller; spire long or short, usually less than %
of the total length.
B. Shell short and stubby, spire rather short,
92 BULLETIN 39 2604
bluntly pointed; 1-3 or less the total length.
Olivella limonensts and variety bocas-
énsts, 1. sp. and var.
BB. Shell with a longer spire, oftensharply point-
ed,
C. Shell rather broad, with a heavy cal-
lus about the upper part of the inner
lip and extending partly over the pen-
ultimate whorl. Length 12 mm. plus.
Oltvella muticotdes Gabb.
CC. Shell slender, with a long, pointed
spire; callus about the upper of the
inner lip, slight.
Olivella Boussact Cossmann, var.
Gliveila goliath, n. sp. Plate 7, figures 22, 23
Shell large, with a long, pointed spire of about % of the to-
tal length of the shell; whorls about 6, with plain, nearly straight
profile; sutures linear; last whorl with the greatest convexity
about the middie; aperture subelliptical, broadest in front; in-
ner lip with a thin, flat callus and with 3 small plaits below.
Length 26, diameter 10, last whorl 20, spire 13 mm
23 8.5 18 A roaven
An unusually large species, represented by two specimens
from Red Cliff Creek and Banana River. Its general form is that
of O. indevisa Guppy and O. Boussaci Cossmann, but nearly 4
times as large.
Gatun Stage: Banana River.
Call ia wkied | Olapm@necton C.F.
Olivella muticoides Gabb Plate 7, figures 11, 14, 17, 18
Oliva muticoides Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
Ais
Olivella mutica variety muticoides Dall, 1890, Tarns. Waguer Free,
Inst. Sci., vol. 3, pt. I, p. 45-
265 Costa RicA MioceENE—OLSSON 93
Olivella muticoides Maury, 1917, Bull. Amer. Pal., vol. 5, p. 232, pl.
TATe io Se ATs
A. broad, chubby species with a spire of moderate length. A
large, thick callus is formed about the upper part of the inner
lip, which spreads partly over the back of the penultimate whorl,
giving to it a hump-back appearance. Measurements of our
Costa Rican shell run as follows:
Length 17, diameter 7, last whorl 14, spire 6.5 mm.
13 6.5 II 5-5 mm.
Gatun Stage: Coll. 5, Red Cliff Creek.
Coll. 4, East Grape Point Creek.
Olivetla Boussaci Cossmann, variety Plate 7, figures 15, 16
Olivella Boussact Cossmann, 1913, Journ. de Conchyliologie, vol. 61,
p. 60, pl. 5, figs. 16-19.
The Olivella tndivisa Guppy (Proc. U.S. Nat. Mus., vol.
19, p. 308, pl. 30, fig. 10) from Jamaica and Olivella Boussact
Cossmann from Martinique, belong to a closely related series
characterized by their high and sharply pointed spire. The
main difference seems to be that of size. The zzdivisa hasa
length of 6.5 mm., the Boussaci of 9 mm. The Costa Rican shells
are generally larger and when full-grown, often 13 or more mm.
in length.
Often very common.
Length 13, diameter 5, last whorl 10, spire 6.5 mm (6
whorls)
Gatun Stage: Hill ra, Banana River.
fiill 3, Banana River.
Olivella limonensis, n. sp. Plate 7, figures 19, 20
Shell short and plump, witha small, broad, conic spire;
whorls about 5, separated by deep sutures; last whorl very large,
broadly cylindrical in form and slightly contracted in the middle;
the inner lip has a wide callus, somewhat heavier above, with 5
94 _ BULLETIN 39 266
fine, denticles in the middle and few, more oblique plaits below;
the base is obliquely encircled by a lighter colored band; outer
lip sharp, smooth within.
Length 1o.5, diameter 4.75, last whorl 9, spire 2.25 mm.
This is a common shell in the coralline phase of the Gatun
at Port Limon. The middle of the last whorl is usually dark col-
ored, with the spire and the encircling basal band and callus on
the inner lip white.
Gatun Stage: Port Limon.
Olivelia limonensis var.bocasensis, n. var. Plate 7, figures 24, 25
A much smaller shell than the /monenszs and possibly a dis-
tinct species. The spire is somewhat higher and the shell less
cylindrical and pump. Dimensions as follows:
Length 6.5, diameter 2.75, last whorl 6, spire 1.5 mm.
ah BuDi5e 6.5 1.75 mm.
Gatun Stage: Bocas del Toro, Panama.
Genus ANCILLARIA Lamarck
Ancillaria aquaensis, n. sp. Plate 7, figure Io
Shell of medium size, solid; spire elevated, composed of about
5 whorls, the sutures of which are concealed by a wide band of
callus this band of callus commences on the body-whorl, a
short distance below the suture, and extends to the apex; where
this band covers the sutures, it forms just above, a depressed or
constricted band on the middle of each whorl; the last whorl is
large, with a basal band of callus, which commences near the
upper end of the inner lip, descends obliquely across the back of
the last whorl to the anterior tips of the outer and inner lips; a
short distance above this basal band, the shell carries a single,
incised line; a small umbilical pit is found just behind the mid-
dle part of the inner lip; aperture broadly sub-elliptica] with a
sharp outer lip.
Length 18.5, diameter 7.5, last whorl 12.5, spire Io mm.
267 Costa Rica M1ocENE—OLSSON 95
The Ancillaria pinguis Guppy, from Jamaica is probably a
closely related species. Guppy’s figure in the Geological Maga-
zine, vol. 1, Decade 2, isvery poor and insufficient. Our shell
seems to differ by its much longer spire and more slender shell.
The Ancillaria chipolana Dall, figured on Plate 41, fig. 3, of
the Wagner Institute, is larger and has a longer spire. No um-
bilical pit is shown in the figure.
Gatun Stage: Water Cay, (sla de Aqua).
Genus MARGINELLA Lamarck
Marginella MacDonaldi Dall Plate 6, figures 14, I5, 17, 19
Marginella MacDonaldi Dall, 1912, Smith, Misc. Coll., vol. 59, No.
2aRD STE
This large M/arginella is one of the most common and char-
acteristic species of the Gatun beds of Costa Rica. It is extreme-
ly valiable in size and general from and heaviness of its shell, as
may be seen in the accompanying figures of the more common
varieties.
Typically the shell is oblong-cylindrical with a small spire of
about 4 whorls, usually completely covered in front by a broad
mass of callus, which spreads over the whole base of the shell
and the outer lip, leaving an elevated ridge along each side as
frequently seen in Cyprga, From the back, the spire may be
seen lying in the mass of callus.
The Marginella mindiensis Cossmann is a smaller related
species from the Canal Zone. It differs also in having its outer
lip finely denticulated, while in /acDonaldz the outer lip is us-
ually smooth.
Length 28, diameter of base 15, vertical diameter 10 mm.
24 14 g mim.
25 13 9.5 mm.
ola) II Gjois\ioab ol.
Gatun Stage: Banana River.
96 BULLETIN 39 268
Marginella mindiensis Cossmann Plate 6, figures 16, 23
Marginella mindiensis Cossmann, 1913, Journ. de Conchyliologie, vol.
6f, p. 61, pl. 5, figs. 13-15.
This shell is very closely related to MW. MacDonaldi. It dif-
fers only in being smaller and usually with the outer lip more or
less denticulated. The outer lip of WZ. MacDonaldi is generally
smooth but occasional specimens occur in which the outer lip is
faintly denticulated. It isa species of the Canal Zone.
Length 19.5; basal diameter 11, vertical diameter 8 mm.
16 9 7 mim.
Gatun Stage: Mindi, (Cossmann)
Gatun Gok
Marginella latissima Dall
Marginella latissima Dall, 1896, Prac. U. S. Nat. Mus., vol. 19, p. 308,
pli2o hos
This is a short and very broad species, described by Dall
from Moen Hill, C. R. Dall remarks:‘‘This is perhaps the short-
est and widest American species.’’
Length 11, diameter 8.5 mm.
Gatun Stage: Moen, Costa Rica, (Gabb).
Marginella latissima, var. pilstryi, mn. var. Plate 10, figures 1, 2
Marginella coniformis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sct.
Phila., vol. 63, p. 348, pl. 24, fig. 12. Not of Sowerby, 1849.
Like M7. latissima Dall, but less broad, more pointed anteri-
orly, a slightly higher spire and a less heavy outer lip. In /afzssv-
ma, the two posterior plications are shown as being very oblique,
while in the present form they are nearly transverse as is seen in
the recent cizcta. ‘The outer lip is finely granulated.
Length 14, basal diameter 9.5, vertical diameter 6.75 mm.
Us II 9g mim.
This shell is figured asthe Marginella contformis Sowerby
a common Dominican fossil, by Brown and Pilsbry. The A/arg-
269 Costa Rica’ MiockENE—OLSSON 97
inella coniformis has been figured by Guppy in the Quarterly
Journal and similiar shells were collected in abundance by the
Maury expedition to Santo Domingo. It isa larger and more
cylindrical form.
It isan abundant shell in the quarries west of the Gatun
locks at Gatun. A single, large shell figured as figure 1 was
collected from the Banana River beds.
Gatun Stage: Gatun, C. Z.
Banana River, C. R.
Marginella avena Valenciennes © Plate 7, figures, 21, 28
Marginellaavena Valenciennes, 1814, in Kiener, Coq. Viv. Marginella,
PAL Zip le Oy ao 2Ay
Marginella avena Gabb, 1881, Journ. Acad, Nat. Sci. Phila., vol. 8,
2ud series, p. 355.
A common, recent species along the Caribbean coast of
Panama and Costa Rica. As a fossil, it occurs plentifully in the
coralline phase of the Gatun at Port Limon and Bocas del Toro.
The fossil shells do not differ materially from their recent repre-
sentative.
The shell is elongate-cylindrical in outline, with a low, but
slightly projecting spire. The aperture is norrowly linear,
slightly wider in front. Outer lip thick, smooth within. The
columella is provided with 4, obliquely descending plications.
A series of specimens from Limon and Bocas measure as fol-
lows.
Length 11.75, diameter 4.25, spire 2.
I1.50 4.25 1.15,
12.50 4.50 a,
Gatun Stage: Port Limon, C. R.
Bocas del Toro, Panama.
Coll. 4, Red Cliff Greek.
Marginalla collina, n. sp. Plate 7, figures 26, 27
Shell of the general form and size of JZ. avena, but more
98 BULLETIN 39 270
solid and with a lower and less differentiated spire; whorls 4 or
more with sutures scarcely distinguishable under their glaze or
coat of callus; aperture linear-elongate, widest in front and
slightly contracted in the middle; outer lip thickened, smooth
within; pillar with 4, oblique plaits of nearly equal strength.
Length 11, diameter 4.5, spire 1.5 mm.
TO 4.25 I mm.
Ein 25 5. I mim.
Closely related to the MWarginella avena Val., this species
differs in being more solid, less slender and with a lower spire.
In avena the spire is clearly differentiated, with well marked su-
tures to its spire-whorls. In the present shell, the spire is much
lower, and the sutures are more thickly covered with glaze. The
thick outer lip is carried across the end of the lip and firmly
joined to the spire without any preciptible line of demarcation.
In avena, the outer lip and spire are clearly differentiated from
each other.
Abundant in the Gatun beds of the Banana River.
Gatun Stage: Hill ra, Banana River.
Marginelia leander Brown and Pilsbry Plate 6, figure 22
Marginella leander Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 347, pl. 24, fig. 13.
Not known from Costa Rica, and the figure here given is
from a specimen in our collection from Gatun. It is a broader
and more cylindrical species than colliza, and with a much low-
er and scarcely elevated spire. Our specimen measures 9 by 4
mm.
Gatun Stage: Gatun, C. R.
Marginella musacina, n. sp. Plate 10, figure 25
Shell small and slender, with a high spire of about 1-4 of
the total length of the shell; whorls about 5, with sutures light-
ly concealed under a thin coat of glaze; last whorl narrowly cylin-
271 Costa Rico MiocENE—OLSSON 99
drical, widest about the upper half; aperture sublinear, with a
slightly thickened lip, somewhat contracted in the middle, smooth
within; inner lip smooth, or with only a thin wash of callus and
with 4, very oblique plications at its extreme anterior end.
Length 9.25, diameter 3.25, spire 2.75 mm.
A small, slender species of unusual form. Two specimens
were collected in.Costa Rica, the type from the Banana River,
the other from Old Man Sam Creek, near Manzanilla Point.
Gatux Stage: Banana River,
Along Old Man Sam Creek, one mile south
of the beach, C. R.
Genus VO@LUTA Linnzeus
Voluta alfaroi Dall Plate 8, figure 2
Voluta alfarvoit 1912, Smith, Misc. Coll., vol. 59, No. 2, p. 8.
This fine species described by Dr. Dall, from the Banana
River, isa common and very characteristic fossil or the Gatun
beds in Costa Rica and Western Panama, but it still remains to
be recorded from the Canal Zone. Allied to the recent West
Indian V. musica Linneeus it differs most importantly in its
much smaller nucleus.
The shell is heavy and in its typical form the whorls are
shouldered and carry about 12 ribs, which may be quite sharp
and high on the shoulder angle. The whorls are sometimes
smooth, without ribs and a shoulded angle. Spiral threads usu-
ally occur on the lower one-quarter of the last whorl and on the
canal, but in some cases on the spire-whorls as well. A large
specimen from Water Cay measures;
Height 59 mm. diameter 34 mm.
Gaiun Stage: Water Cay Panama.
Coll. 3, Red Cliff Creek; Old Man Sam Creek,
r mile from shore, Coll. 2, QOuttana Creek;
Comadre Creck, Soust creek; Coll. 5, Estrella
River; Banana River, Rio Blanco, Port Limon.
100 BULLETIN 39 272
Genus SCAPHELLA Swainson
Scaphella costaricana,n. sp. Plate 15, figure 13
Type fragmentary consisting of the nucleus and part of the
three succeeding whorls; the sheil is subfusiform; nucleus large,
smooth and mamumilate at its apex; the three following whorls
are slightly contracted about the upper sutures and strongly sculp-
tured with coarse subregular, spiral threads separated by inter-
spaces as wide or a little more; the spiral threads and their in-
terspaces are crossed by fine lines of growth; sutures distinct; the
columella with four plaits, the posterior one being the strong-
est.
Length 29, diameter 18, diameter of nucleus 5.75 mm.
The unique type is unfortunately fragmentary and consists
of the large, smooth, mammilate nucleus and part of the 3 suc-
ceeding whorls. ‘The shell is subfusiform and the whole surface
is strongly sculptured with coarse, spiral threads. The columel-
la is provided with 4 strong plaits. The type specimen was found
in the Dentalium zone of the Upper Uscari shales of Cocles
Creek?
Uscart Stage: Cocles Creek.
Genus MITRA Lamarck
Mitra Swainsoni Broderip, var. fimonensis, n. var. Plate 6, fig. I.
ef. Witra Swainsont Roderip, 1835, Proc. Zool. Soc., p. 193.
ef. Mitra Swainsoni Reeve, Conch. Icon., (Vitra, pl. 1, fig. 4.
ef. Mitra Swainsont, var. antillensis Dall, 1589, Bull. Mus. Comp.
Zool., vol. 18, p. 158, pl. 38, fig. 7.
Shell large, of about 6+ whorls; the spire whorls are slight-
ly convex, the last distinctly shouldered above, depressed or
slightly concave about its middle; sculpture consisting on the
spire-whorls of about 6, heavy, spirals cords with sculptured
interspace as seen on Mitra Henekeni and longa, on the later
whorls the spiral cords become more widely spaced and on the
273 Costa Rica MrocENE—OLSSON IOI
last are in the form of broad, subobsolete, smooth bands without
the sculptured interspaces; there are about 15 spiral cords on
the last whorl in addition to those of the anterior canal; anterior
canal of moderate length, strongly twisted; columella with 4
plaits, heaviest above; aperture narrow.
Length 74 (apex broken), diameter 23, last whorl 52,
spire 35 mm.
A single, large shell with rudely sculptured whorls was col-
lected from the coralline limestone near Port Limon. It agrees
closely with Reeve’s figure of A%ttra Swainsont, except that the
spiral bands are larger and heavier.
The Mittra Swatnsont is a West Coast shell belonging to the
Panamic province. It is distinguished by its large size, rudely
sculptured whorls and dark colored epidermis. Dall has des-
cribed as the variety azftillensis, specimens dredged off of Cape
Lookout, N. C., Colombia and Yucatan.
Gatun Stage: Port Limon.
Mitra longa Gabb Plate 6, figure ro
Mitra longa Gabb, 1873. Trans. Amer. Phil. Soc., vol. £5, p. 219.
Mitra longa Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., vol.
Osh pa346. pls 240 hig. Ta.
Mitra longa Maury, 1917, Bull. Amer. Pal., vol. 5, p. 238, pl. 11, figs.
TATA
A long, slender species, sculptured with strong, revolving
cords and fine, neatly engraved interspaces. We have collected
this Dominican species only inthe Canal Zone. The figured spec-
imen has the following dimensions:
Length 4o, diameter 10.5, last whorl 26, spire 21 mm.
Gatun Stage: Gatun, C. Z.
Mitra dariensis Brown ond Pilsbry Plate 6, figure 25
Mitra dariensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila.,
vol. 60, p. 346, pl. 24, fig. 11.
102 BULLETIN 39 274
A single specimen from the Island of Bocas (Columbus or
Colon Island) is here figured. It differs from the W/tra longa,
in being shorter and broader. ‘The sculpture is less elegant.
Length 21, diameter 7, last whorl 14, spire 11 mm.
Gatun Stage: Gatun, C. Z.
Bocas del Toro, Panama.
Mitra aff. rudis Gabb | Plate 6, figure 13
Mitra rudis Gabb, 1873, Trans. Amer, Phil. Soc., vol. 15, p. 220.
Shell subfusiform, biconic; whorls about 7, coarsely sculp-
tured with strong, revolving cords, separated by interspaces of
about twice their width; These interspaces are finely longitudi-
nally sculptured with raised threads above, obsoletely so below;
the spire-whorls have 5 cords, the last whorl with 11 and several
smaller ones on the canal; aperture subelliptical, with a thin out-
er lip; columella with 4 plaits, largest above; anterior canal of
medium length, strongly twisted below.
Length 33.5, diameter 12.5, last whorl 24, spire 16 mm.
It is with much uncertainty, that I have identified this rare
species with Gabb’s unfigured Mitra rudis from Santo Domingo.
Gabb,s descriptions and measurements indicate a shell of much
the same characters as well as dimensions.
Gatun Stage: Coll. 3, Hone Walk Creek, C. R.
Mitra poas, n. sp. Plate 6, figures 20, 21
Shell small, nearly smooth and columbelloid in appearance;
whorls 8, with straight sides; last whorl widest just above the
base, which is contracted to the short anterior canal; the early
spire-whorls have 5, low, smoothish spiral bands, with longitudi-
nally sculptured, narrow interspaces; these spirals quickly be-
come obsolete, leaving the whorls smooth except for one or two
spirals bordering the upper sutures; the last whorl has about 4
spirals around the base and smaller ones on the canal; aperture
subellipical, with a thin outer lip; columella with 4 plaits, larg-
275 Costa RicA M1ocENE—OLSSON 103
est above; anterior canal of medium length and slightly bent to
the left
Length 21, diameter 7.5, last whorl 13, spire 11 mm.
A smooth species of Columbelloid aspect. It is fairly abund-
ant in the coral limestones near Port Limon.
Gatun Stage: Port Limon,
Mitra Almagrensis Toula var. coralliophila, n. var. Plate 6, figures 18, 24
of Mitra Almagrensis Toula, 1911, Jahrb, der K-K Geol. Reichsanstalt,
VOl VOID. 40m ple 23, fei 13:
Shell small with a long spire and a shorter anterior canal;
whorls about 8, with straight or slightly convex profile; sculp-
ture of low, slightly elevated spiral cords, separated by inter-
spaces of about their own width; these interspaces are finely
sculptured by regular, raised, longitudinal threads; the spire-
whorls have about 5 spiral cords, the last whorl with about 15;
and smaller ones on the anterior canal; aperture subelliptical; a
short canal, slightly bent to the left; pillar with 3 folds and a
very small faint one below.
Length 17, diameter 6, last whorl 11, spire 10 mm.
This is a common species in the coral limestones of Limon.
Toula’s Mitra Almagrensis was described from Tehuantepec.
Our shells differ from Toula’s figure in being more slender and in
having the body-whorl less contracted below.
Gatun Stage: Port Limon.
Genus FUSUS Lamarck
Fuscs miocosmius, n. sp. Plate 8, figure 5
Shell long and slender, with the spire and canal of nearly
equal length; whorls about 11 plus, very gradually tapering from
the small nucleus to the body-whorl; whorls convex with indis-
tinct, appressed sutures; the sculpture consists of large, swollen
ribs, separated by equally wide interspaces, and the whole
104 BULLETIN 39 a7
crossed by strong spirals with finer threads in between; on the
last whorl there are 7 ribs, which commence on the base, just
above its union with the long canal, and continue across the
whorl to the suture; the early spire-whorls have 6 or 7 strong
spirals with 1, 2 or 3, fine threads between; on the penultimate
whorl, the earlier spirals have become a primary set of 6 or 7,
with their intervals occupied by asecondary set, nearly as large
as the primary, and still smaller tertiary threads; the last whorl
has about 18 strong spirals in addition to those on the canal; on
the canal the division into primary and secondary spirals is bet-
ter shown, there being about 16 primary and the same number of
secondary spirals; on the extreme tip of the canal the spirals are
very small and numerous; aperture small, rounded, with the out-
er lip internally lirated and three or more oblique plaits on the
columella.
Height roo, diameter 24 mm.
This fine species is closely related to the recent Fusus eucos-
mtus Dall, from the Florida coast and the West Indies. The spire
of ezcosmius is more slender, has 8 ribs instead of 7, and the out-
er lip is always smooth, while it is strongly lirated in szocos-
meus.
Gatun Stage: Bocas del Toro, Panama.
Fusus honensis, nu. sp. Plate 8, figure 6
Shell resembling in general features the /. mzocosmius, and
with the same number of ribs; the spire is somewhat longer than
the anterior canal; the whorls are somewhat more convex than
those of mzocosmtius due to the more strongly knobbed ribs; the
ribs are crossed on the penultimate whorl by about 6, strong,even
spirals, there being no secondaries and only occasionally a small
thread may appear in their intervals; the last whorl, exclusive
of the canal shows 10 or i1 spirals to which are added 17 or 18
on the canal; the canal is relatively short and not perfectly
straight; the aperture is probably rounded (largely broken on the
type specimen) and with its outer lip internally lirated.
Height 63, diameter 20 mm.
277 Costa Rica MrocekNE—OLSSON 105
Similiar in general form and in the number of its ribs to the
fF. miccosmius from Bocas, this species differs in its smaller,
heavier shell, and in the greater coarseness of its sculpture. The
spirals are heavier and primary in character throughout and the
ribs are more knobbed and sharp on their crests. From the Do-
minican /, Hlenekent Sowerby, it differs in being much more
slender and with fewer, persistent ribs.
Gatun Stage: Coll. 4, Home Walk Creek.
Genus FASCIOLARIA Lamarck
Fasciolaria Gorgasiana Brown and Pilsbry Plate 8, figure 9
Fasciolaria gorgasiana Brown and Pilsbry, 1912, Proc. Acad. Nat.
Sci. Philia., vol. 64, p. 506, pl. 22, fig. 5.
This large Fasciolaria is fairly abundant in the Gatun beds
of the Canal Zone and is recognized by its shouldered whorls and
and strong knob-like ribs. The figured specimen is a large shell
from the Gatun of Rio Betey, Costa Rica.
Gatun Stage: Gatun, C. Z.
Water Cay.
Rio Betey.
Fasciolaria tulipa Linnzeus, variety Plate 13, figure 4
Murex tulipa Linnzeus, 1758, Syst. Nat.,ed. 10, p. 754.
Fasciolaria tulipa Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8,
2nd series, p. 354.
Fasciolaria tulipa Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 3,
DEIN LOL apl yy 7 uuies on
The Tulip shell is the common, recent /asctolaria, in the
Caribbean area, and it is distinguished from its more northern re-
lative, the dzstans, by its darker coloration and sulcated sutural.
band. It occurs as fossil in the Pliocene of Florida.
From the coralline limestones, near Port Limon we collected
three young specimens and a portion of the spire of a large in-
dividual of Fasctolaria. ‘The young shells show the nucleus and
106 BULLETIN 39 27%
early sulcated whorls of typical ¢ulipa, as fignred by Dr. Dall.
They however become smooth somewhat earlier and the fragment
of the spire shows the succeeding whorls to be smooth and lack-
ing the sulcated sutural band as in dsfans,
The fasciolaria semtstriata Sowerby is an allied but distinct
species from the Miocene of Santo Domingo, and likely to be dis-
covered in the Gatun beds of Costa Rica and Panama. ‘This as
may be seen from the new figure of Dr. Maury, (Bull. Amer.
Pal., vol. 5, p. 244, pl. 13, fig. 1), differs conspicuously from the
distans and tulipa, with which it was united by Gabb, by its
deep sutural depressed band, which gives to the whorls, a very
convex or even shouldered appearance. The early whorls are
spirally sulcated as in ¢u/ija, but have in addition knob-like ribs,
like those seen on /. gorgastana.
Useart Stage: Port Limon.
Fasciolaria MacDonaldi, n. sp. Plate 8, figure 1
Shell large, (type specimen imperfect, with only the last two
whorls preserved); number of whorls unknown; the spire-whorls
are angled about the middle, forming a board, sloping shoulder
above; the last whorl has the shoulder about the upper third; the
sculpture is predominantly spiral, consisting of numerous, fine,
alternating threads, which are slightly roughened by the growth
lines; the angle of each whorl is longitudinally undulated by 12
faint ribs; anterior canal is nearly straight with two faint plice;
outer lip sharp.
Length (2 whorls) 70, diameter 37 mm.
Of this large and elegant species, only a single imperfect
specimen with less than two complete whorls preserved, was col-
lected from the Gatun beds of the Banana River, Costa Rica. It
differs from the more common /. Gorgaszana, by its persistent
and rough spiral sculpture and less heavily tuberculated should-
er.
This species is named in honor of Dr. D. F. MacDonald, well-
known for his geologic work on the Isthmus, during the con-
270 Costa Rico MrocENE—OLSSON 107
struction of the Panana Canal, and former Chief Geologist of
the geological force of the Sinclair Oil Corporation in Panana and
Costa Rica.
Gatun Stage: Hill No. 2, Banana River.
Genus LATIRUS Montfort
Latirus infundibulum Gmelin, variety Plate 8, figure Io
Latirus infundibulum Gmelin, Lamarck, Anim. sans, Vert. (ed, Desh. )
vol. 9, p. 386.
Latirus infundibulum Guppy, 1866, Quart. Journ. Geol. Soc. Loudon,
vol. 22, p. 288.
Latirus infundibulum Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15,
pe2t7.
Latirus infundibulum Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. '
8, 2nd series, p. 354.
Latirus infundibulum Maury, 1917, Bull. Amer. Pai., vol. 5, p. 246,
pla he. 3
A common recent species of the West indian and Costa Rican
coasts. Only an imperfent specimen was collected from the Gatun
beds of lower Pumbri Creek, a small tributary of the Estrella
River. From recent examples of znfundibulum, it differs by its
shorter canal, more numerousribs (9 instead 6 or 7),and heavier
spiral sculpture. The specimen is, however, to fragmentary to
serve as a type for a new variety or related species.
Gatun Stage; Coll. 7, Pumbri Creek; GC. R:
Latirus irazu, n. sp. Plate 8, figure 12
Shell subfusiform; solid, with a long spire and shorter can-
al; whorls about 9, with heavy, sharp, knob-like ribs and finer
spiral threads; the last whorl shows 7, sharp, pointed ribs, which
are continuous from suture to suture, but only feebly across the
base of the last whorl; the suture is bordered anteriorly by a
prominent, frilled band or cord, formed by the elevated edges
108 BULLETIN 39 280
of the growth lines: spirals consisting of low, raised threads with
wide interspaces; canal short, straight above but bent below, and
with three small, smooth folds on the columella above; aperture
with a thin outer lip.
Length 46, diameter 17, last whorl 29, spire 24 mm.
A smaller and more delicate species than the preceding ZL. zz-
fundtbulum Gmelin. In the present shell, the ribs are sharp-
er and more pointed, and the canal is shorter and more strongly
reflected backwards. The strongly frilled sutural band is lacking
or only imperfectly developed on znfundibulum.
Gatun Stage: Port Limon.
Latirus taurus, 0. sp. Plate 8, figure 4
Shell large, slender, solid with the spire more than % the
height of the shell; whorls convex, with indistinct sutures; sculp-
ture of very regular, narrow ribs with deep interspaces, and cross-
ed by strong, ridge-like spiral cords; whorls about 9; axial sculp-
ture on last whorl with about 11 ribs; the ribs commence well
down on the base and continue to the upper suture and are
nearly in line acrossthe spire-whorls to the apex; the spirals
consist of ridge-like cords and finer spiral threads; there are
2 principal spirals on the spire-whorls which cross the middle
of the whorls; there are 4 principal spirals on the last whorl
above the edge of the base, below which there are a few small-
er threads; the anterior canal has 2 strong cords and finer
threads are scattered over the whole shell; the anterior canal
is straight and stocky and carries at its base a deep but narrow
umbilical pit; the columella has 4 small folds; aperture subo-
vate, anteriorly extended to form the long narrow canal; out-
er lip smooth within.
Height 60, diameter 25, aperture 32 mm.
A large, solid and strongly sculptured shell. The straight,
narrow ribs are spaced regularly over the whorls of the shell and
are continuous from the base to the upper suture. They are
a8x Costa RicA MiocENE—OLSSON 109
crossed by heavy, ridge-like, spiral cords, which on crossing the
deep interspaces between the ribs, form small, sunken pits, so
that the sculpture appears coarsely trellised.
Gatun Stage: Toro Cays.
Genus PTYCHOSALPINX Gill
Ptychosalpinx ? dentalis, n. sp. Plate 15, figures 14, 18
Shell buccinoid with convex, cancellated whorls; whorls
about 6, with channelled sutures so that the whorls appear nar-
rowly shouldered above; the sculpture is predominantly spiralled,
the penultimate whorl with 7 strong spiral cords with interspaces
2 or 3 times their width; small spiral threads may appear in the
interspaces; the last whorl has 14 or more spirals and the wide
interspaces with one or more finer threads; the spirals are cross-
ed by coarse wavy lines of growth, producing a sub-cancellate
sculpture; the aperture is subovate; a well-developed siphonal
sinus; the columella carries a single sharp plication at its lower
end; the outer lip is slightly thickened and bears within about
12, small, narrow liree.
Height 28, diameter 11, aperture 16 mm.
The generic relation of this interesting species is very much
in doubt. It has the general form of Comznella, but its colum-
ella is provided with a strong anterior plication. In this regard
it agrees with Pfychosalpinx of which there are several species in
the Chesapeake Miocene. The apex of dentalis, appears to have
been pointed and sharp, although all of our specimens have the
tip of the spire very much weathered. True Ptychosalpinx is
characterized by a large, convex and blunt nucleus and the outer
lip is thin and smooth within. The outer lip of deztalzs is slight-
ly thickened and internally finely hrated. .
The P ? dentalis, is a very characteristic fossil of the Denta-
lium zone of the Upper Uscari formation.
I10 BULLETIN 39 252
Uscari formation. Coco Plum, Panama.
Rio Cocles.
Comadre Creek, etc.
Genus PERISTERNIA Morch
Peristernia insuia, n. sp. Plate 8, figure 11
Shell small, with a sharp pointed spire, a little longer than
the aperture; the general form and sculpture of the shell is like
that of Uvosalpinx cinereus Say; nucleus of 2 small smooth
whorls, followed by 7 post-nuclear; the profile of the spire-whorls
is convex and strongly sculptured with ribs and sharp spiral cords;
the last whorl has 8 ribs which are nearly lacking from the base
of the whorl; the spiral sculpture consists of 2 principal cords
about the middle of the earlier whorls above which lie smaller
threads about the suture; on the later whorls, the spirals are
somewhat heavier about the middle but irregular with finer, in-
termediate threads in between the principal ones; base contract-
ed; aperture subcircular; outer and inner lips crenulated or den-
ticulated; canal short and bent.
Height 23, diameter 13, aperture 11 mm.
This and the following fortugera seem correctly referable to
the genus FPeristernta Morch. The P. insula is somewhat like
young specimens of //icata Conrad, from the Chesapeake Mio-
cene of eastern United States, but has a longer and more pointed
spire, and heavier sculpture.
Gatun Stage: Water Cay.
Peristernia tertugera, n. sp. Plate 8, figure 13
Shell elevated with a spire much longer than the aperture;
nucleus of about 2% small, smooth whorls; the post-nuclear
whorls about 7; sutures distinct; whorls strongly convex, should-
ered; sculpture of narrow, heavy ribs, widely spaced and num-
bering on the last whorl about 9; the rib are continued across the
whorls from suture to suture and for a short distance down on
283 Costa Rica MIocENE—O1S¢sON II
the base; the spirals consists of irregular sharp cords; the early
spire whorls carry 2 main spiral cords but bordered above and be-
low by finer threads; the 2 principal spirals are continued on the
later whorls but are nearly equalled in strength by the other
spirals and hence lose their prominence; a short anterior canal;
base strongly contracted.
Height 29, diameter 14.5, aperture 12 mm.
Distinguished from the preceding zmsau/a by its very much
longer spire, more strongly contracted base and different spiral
sculpture.
Gatun Stage: Port Limon.
Genus XANCUS Bolton
Xancus scopulus, n. sp. Plate 11, figure I
Shell large, solid and heavy; spire nearly as long as the aper-
ture; spire-whorls 6 plus (the tip broken), strongly coronate
above and with large, wide, persistant ribs; the earlier spire-
whorls are simply shouldered or angled about the middle, but the
area about the upper sutures rapidly deepens and on the later
whorls is a deep, excavated sutural zone, above which project
the ends of the ribs; there is a strong sutural cord and a rather
wide, ribbon-like band just above, forming at first a strongly ap-
pressed suture; the last whorl has about 7 large, wide ribs, the
areas between appearing as troughs or depressions; the sutural
excavated zone carries several irregular spiral threads which are
crossed by large growth-lines so that the resulting sculpture is
more or less cancellate, more particularily on the earlier whorls;
the growth-lines cross the sutural cord and upon the ribbon
above, become much crowded and strongly bent forwards; the
young shell was sculptured over the whole shell with strong
spirals, but with maturity, the spirals become obsolete and the
shell is smooth and polished; columella with 3, strong plicee as in
the recent scolymus; a long anterior canal, with a narrow, deep
umbilicus behind the spreading inner lip.
m2 BULLETIN 39 284
Height 265, diameter 136, aperture 158, last whorl 195
mm.
This very remarkable species is an extreme development of
the X. scolymus stock in which the upper portion of the whorl
becomes a wide, deep, excavated sutural zone. Above this ex-
cavated zone, project the high, rounded or appressed ribs and the
carinate edge of the whorls. X. scolymus Gmelin, a recent spec-
ice found plentifully along the north Panama coast, has the
whorls simply shouldered, often merely rounded.
The Santo Domingan Miocene contains X. validus Sowerby,
which has been identified by some with scolymus. In validus, the
ribs are more numerous (about 10) and are sharper and more tub-
ercular in form.
Gatun Stage: Banana River.
Genus MELONGENA Schumacher
Melongena consors Sowerby Plate 9, figure 1
Pyrula consors Sowerby, 1849, Quart. Journ. Geol. Soc. London, vol. 6,
Pp. 49.
Melongena melongena Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p-
205. Not of Linneeus.
Pyrula melongena Guppy, 1874, Geol. Mag., vol. 11, p. 438.
Pyrula melongena Guppy, 1876, Quart. Journ. Geol. Soc. London, vol.
32, P- 523.
Melongena consors Dall, 1900, Trans, Wagner Free Inst. Sci., vol. 3, pt
1) 169) bo
Melongena consors Maury, Bull. Amer. Pal., vol. 5, p. 249, pl. 14,
fig. 5.
The Melongena consors is not a common fossil in Costa Rica.
It is closely related to the recent 7. corona Gmelin of the West
Indies, the fossils shells differing mostly in having a longer spire
and somewhat different sculpture above.
The Costa Rican examples are exactly like Miocene specti-
mens from Santo Doingo. As a fossil it occurs in the Miocene
of Jamaica, Santo Domingo and Venezuela.
285 Costa Rica MrockENE—OLSSON 113
Gatun Stage: Coll. 5, Old Man Sam Creek.
Cocles Creek.
Genus SOLENOSTEIRA Dall
Soilenesteira Dalli, Brown and Pilsbry Plate 8, figure 8
Solenosteira dalli, Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 348, pl. 24, fig. 14.
Coralliophila gatunensis Toula, 1911, Jahrb. der K-K Geol. Reichsan-
stalt, vol. 61, p. 502, pl. 30, fig. 9.
A common species in the Canal Zone and figured here for
comparision with the following species. The whorls are strong-
ly shouldered, a feature accentuated by the few, but high,angled
ribs. The spirals are heavy, raised cords, over and between
which are finer secondary and tertiary threads.
Length 34, diameter 23, last whorl 27, spire 15 mm.
Gatun Stage: Gatun, Mt. Hope, C. Z.
Solenosteira Vaughani Dall, var. medioamericana, n. var. Plate 8, fig. 7
cf. Solenosteira Vaughani Dall, 1903, Trans. Wagner Free Inst. Sci.,
vol, 3, pt. 6, p. 1633, pl. 60, fig. 18.
Shell solid, with a small pointed nucleus of 2 smooth whorls,
and 6 post-nuclear; spire conic, of about % the total length of
the shell; whorls convex, or bluntly shouldered about the mid-
dle ; sculpture of numerous low ribs which are rounded, and
heaviest on the shoulder angles; There are 7 to Io ribs on the
last whorl; spirals of raised cords, with finer threads on the in-
terspaces; the spire-whorls have about 6 primary spirals, the last
whorl with 18, including those of the short canal; canal short
with a deep but narrow umbilical pit; aperture oval, a thick in-
ner and outer lip, the outer with about 10, long, entering lire
and the inner with three small denticles above.
Length 42, diameter 24, last whorl 34, spire 17 mm.
34 2S 28 13 mim.
The Solenosteira Vaughani is a Chesapeake Miocene species
T14 BULLETIN 3G 286
from Jackson Bluff and Coe’s Mill, Florida. Dall’s figure in the
Wagner Free Institute, isso like our shell from western Panama
and Costa Rica, that there can be but little doubt of their close
affinities.
The present shell is larger than the S. Dalli, of the Canal
Zone, with more numerous, obtuse and not sharp ribs, and with
a wider and deeper umbilical pit. A canal is developed at the
posterior angle of the aperture asin the recent S. pallida Brod.
while in S. Dadli, the posterior portion of the aperture is round-
ed and there is no canal.
Gatun Stage: Toro and Water Cay, Panama.
fiill ra, Banana River, C. R.
Solenosteira chiriquiensis, n. sp. Plate 8, figure 3
Shell large, heavy; spire about % the height of the shell;
whorls about 7, strongly angled about the middle and carrying
heavy knob-like ribs, crossed by heavy spiral cords; the last
whorl has 8 ribs which are developed only on the middle of the
shell, being lacking from the upper slope and from the base of
the last whorl; the tops of the ribs are crossed by 2 strong,spiral
cords with a wide, trough-like interval between; above the 2
principal spiral cords, there are 4 smaller cords on the upper
slope and on the base and canal ro or 11; base of the last whorl
contracted to form the straight canal which carries a deep, narrow
umbilius; aperture subelliptical.
Height 57, diameter 34, aperture 34 mm.
It is possible that this species belong to the genus Cymza
rather than Solenostetra, but its aperture is so completely filled
with a hard sandstone matrix that the presence or absence of a
columellar fold cannot be determined. Its sculpture however is
more like Solenosteiva than Cymia. ‘The species will be recog-
nized by its large size and characteristic sculpture.
Gatun Stage: Water Cay.
287 Costa Rico M1ocENE—OLSSON II5
Genus METULA H. and A. Adams
Metula cancellata Gabb Plate 10, figure 12
Metula cancellata Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, Pp.
205.
Metula cancellata Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd
series, p. 351.
Metula cancellata Dall, 1903, Trans. Wagner Free Inst. Sci., vol, 3, pt.
6, p. 1584.
Metula cancellata Maury, 1917, Bull. Amer. Pal., vol. 5, 249, pl. 14,
fig. 19.
Less common in Costa Rica and Panama than the following
species and from which it is easily distinguished by its usual
smaller size, more delicate shell, and fine sculpture.
The spiral threads on the last whorl number about 37 and
are crossed by nearly as fine longitudinal ribs. The resulting re-
ticulate sculpture is fine and neat. Our largest shell, a specimen
from Bocas with 6 whorls measures:
Length 19, diameter 6.5, last whorl 14, spire 9, aperture
IO mm. e
Gatun Stage: Bocas del Toro, Panama.
fiill ra, Banana River, C. R.
Metula Gabbi Brown and Pilsbry Plate 10, figure 11
Metula Gabbi Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila.,
vol. 63, p. 351, pl. 25, figs. 4, 8.
This beautiful species is fairly common in the Canal Zone,
but usually in a fragmentary condition. It related to the W/etula
cancellata Gabb, differing mainly in its larger size and heavier
sculpture. Ona speciman of 8 whorls from Gatun, the spirals on
the spire whorl number about 8 and 39 on the last whorl and an-
terior canal. The 2 upper spirals are noticeably stronger than
the others. Aperture long and narrow and finely denticulated
along the interior of the outer lip.
116 BULLETIN 39 288
The figured specimen from Gatun has the following meas-
urements:
Length 29, diameter 10.5, last whorl 22, spire 12.5, aper-
ture 16 mm.
Gatun Stage: Gatun and Mt. Hope, C. Z.
Metula Harrisi, n. sp. Plate to, figure Io
Shell solid and more coarsely sculptured then Wetula Gabéz,
and with a longer and broaded spire and shorter aperture; whorl
6 plus, (apex lost); sculpture of spiral cords and finer, curved
longitudinal ribs; the spire whorls with 8 spiral cords which are
finest next to the lower suture, become progressively stronger
above; on the last whorl and canal there are about 25 spirals;
the spirals are slightly nodulated by the ribs of which there are
about 40 on the last whorl; certain of the spirals on the middle of
the whorl are double; aperture ovate-elliptical with a strongly
thickened outer lip, internally with about 19 short lire; inner
lip smooth; canal short and twisted.
Length 26.5, diameter 11, last whorl 19.5, aperture 13.5,
spire 14 mm. ¢
A solid and more coarsely sculptured species than the A/etula
Gabbi. ‘The spire is broad and porportionately longer, due to the
shorter aperture. In Gaddz the spirals are of nearly equal
strength over most of the shell surface with exception of those
immediately bordering the upper suture. In the present species
the spirals are finer below and progressively become stronger post-
eriorly.
The type specimen was collected in the lower Gatun sand-
stones of Coco Plum, Panana.
Gatun Stage: Coco Plum, Bocas del Toro, Panama.
Metula Harrisi var. limonensis, n. sp. Plate 10, figures 5, 6
Shell more slender; a narrow and longer aperture more
nearly the length of the spire; sculpture coarsely reticulate,
289 Costa RicA MrocENE—OLSSON 117
with ro spirals on the penultimate whorl, about 26 on the last;
aperture long, with a thickened outer lip, internally lirated in
unison with the external spiral cords; inner lip smooth witha
long, bent, beak-like canal.
Length 27, diameter 9.50, last whorl 19, aperture 14,
spire 13 mm.
Several specimens from the coral limestones of Port Limon.
They are most closely related to Harrisz, in their heavy shell,
coarse sculpture, and thickened outer lip. They differ in being
more slender, a longer aperture and in minor details of their
sculpture.
Gatun Stage: Port Limon,
Genus PHOS Montford
Phos gatunensis Toula Plate 9, figures 4, 5
Phos gatunensis Toula, 1909, Jahrb. der K-K Geol, Reichsanstalt, Wein
VOlNsop. 7Ol, pl-i28, tig 6: plo 25 (hee T,
Phos gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila.
vol. 63, p. 349, pl. 25, figs. 1, 2.
The Phos gatunensis is possibly the most common fossil in
the Gatun beds of the Canal Zone and the adjacent sedimentary
areas in the Province of Colon, but it still remains to be collected
outside of this, its type area.
The shell is typically rather slender, with a sub-reticulate
sculpture formed by the intersection of its numerous, straight
longitudinal ribs, and the nearly equal, regular, strap-like, spiral
cords.
A typical specimen measures as follows:
Length 30, diameter 13, last whorl 20, spire 15 mm.
Gatun) Stage: (Gatun, CZ.
Phos mexicanus Bose Plate 9, figures Io, II
Phos mexicanus Bose, 1906, Bol. de Inst. Geol. de Mexican, numero
22, p. 38, pl. 4, figs. 18-21.
118 BULLETIN 39 290
Shell more broad and solid than gatumensis of the Canal
Zone; whorls about 8, those of the spire with nearly straight or
only slightly convex sides; last whorl large, broad, convex and
widest about the middle; the sculpture is much more coarsely re-
ticulate than in gatunens?s, with the intersection points of the
ribs and spirals more or less subnodulose; the last whorl has
about 18 ribs, which on the back of the outer lip are more close-
ly spaced than elsewhere; the spiral cords are not so heavy as the
ribs, separated by wider interspaces which carry one or some-
times two very, fine, spiral threads; in addition, the spiral inter-
spaces are very finely and neatly sculptured with fine, raised,
longitudinal lines, corresponding to the growth-lines; on the
spire-whorls there are 5 spiral cords, 13 on the last whorl and 3
more on the canal; canal short, twisted; aperture sub-ellip-
tical, with a moderately heavy, but not thickened outer lip and
internally lirated.
Length 30, diameter 15.5, last whorl 21, spire 14.5 mm.
A broader, more solid and more coarsely sculptured shell
than the Phos gatunensts of the Canal Zone,
It was described from the Miocene of Paso Real, near Tux-
tepec, in the State of Oaxaca, Mexico, by Dr. Bose, whose excel-
lent figures agree exactly with specimens from Water Cay. The
several species of Phos, are often extremely abundant but local
in their distribution. Such is the case of Phos gatunensts of
the Canal Zone, the Phos Mooret var. costaricenis of the Banana
River. Likewise in the present instance; the mextcanus, is a very
abundant fossil of the Gatun beds of the islands of the Chiriqui
Lagoon and the adjacent Valiente Peninsula. A few specimens
have also been collected in Costa Rica.
Gatun Stage: Water Cay. Toro Cay. Coco Plum, ete.
Rio Cocles, C. R.
Phos Moorei Guppy, var. costaricensis, n. var. Plate 9, figures 8, 9
Shell of medium size, solid; spire long about one-half the to-
tal length of the shell; whorls 10, of which the first two belong
2QT Costa Rica MrocENE—OLSSON 119
to a small smooth nucleus; succeeding whorls are sculptured with
heavy persistent, straight ribs (13 or 14) on the last whorl; the
ribs are crossed by even, raised, spiral cords; the spire-whorls
have 4 spiral cords and a small one close to the upper suture; an-
terior canal short and strongly twisted; aperture subelliptical,
the outer lip with about Io, strong and regular internal lire.
Height 27, diameter 12, last whorl 17, spire 14 mm.
This is the common /hos of the Banana River. Although
closely related to Mooret, from Jamaica, the Costa Rican fos-
sils are smaller and more slender, as compared with Guppy’s
nearer they Ouarterly, Journal, volw22s\plario. ig.) 1:
Dr. Maury’s Phos Moorez, from the Miocene of Santo Do-
mingo probably represents a distinct species, differing by its
more slender spire, and in sculpture. The spiral intervals of
costaricensis are smooth, while in the Santo Domingan shells,
they carry fine spiral threads, which are best seen on the spire-
whorls, sometimes becoming obsolete on the later.
Gatun Stage: Banana River,
Old Man Sam Creek, r mile from shore or
beach.
Phos elegans Gupp, var. limonensis, n. var. Plate 9, figures 12, 13
Shell usually smaller, more slender and delicate than the pre-
ceding species; the spire is long, composed of about 9 whorls:
the nucleus consists of 4 whorls, the first 2 are small,
smooth and convex, the last 2, smooth, but sharply keeled about
the periphery on the last 1-4 of the last nuclear whorl, faint
curved lines appear above the peripheral keel, they gradually in-
crease in strength and on the Ist post-nuclear whorl become the
longitudinal ribs; the sculpture consists of fine, longitudinal ribs
(23 or 24 on the last whorl); occasionally a rib may become
greatly enlarged, forming varix-like thickenings corresponding
to resting stages; the ribs are overrun by primary, secondary and
tertiary spiral threads; the later spire whorls have about 6 spirals,
120 BULLETIN 39 202
with about 17 on the last whorl andthe canal; aperture suboval,
with a heavy but not thickened outer lip, internally lirated.
Length 27, diameter 11, last whorl 17, spire 4.5 mm.
This shell is related to the Phos elegans Guppy of the Mio-
cene of Jamaica and Santo Domingo. Our shells differ from
typical examples of elegans from Santo Domingo, in being larg-
er, and more coarsely sculptured.
Varix-like thickenings occur scattered along the spire-whorls.
The protoconch of typical elegans and limonensts are practically
identical. The recent West Coast Phos Veraqguensits Hinds isa
related but more finely sculptured species.
Gatun Stage: Port Limon.
Bocas del Toro.
Phos estreilensis, n. sp. Plate 9, figures 17, 18
Shell small, slender, solid; spire much longer than the ap-
erture, pointed; whorls of the spire 6 or more, convex in profile
with deep sutures; sculpture of narrow, straight ribs which com-
mence on the columellar area and pass across the whorl to the up-
per suture; there are 12 or 13 ribs on the last whorl which are
evenly and widely spaced except on the back of the outer lip
where they are smaller and more numerous; the spirals consist
of a primary set of threads or cords with a single, smaller second-
ary thread in each interspace except on the base and the canal;
the primary threads crossing the ribs form small, whitish tuber-
cles; the spire-whorls have 4 primary spirais and there are 11 on
the last whorl, exclusively of the canal; aperture sub-elliptical,
produced anteriorly to form the straight anterior canal; outer lip
thickened and internally lirated.
Height 20, diameter 8, aperture 9 mm.
A small, slender species, coarsely sculptured like the larger
and broader Phos Moorei variety costaricensis. It is limited in
our collections to the Gatun beds of the Estrella River.
Gatnu Stage: Coll. 6, 7, Estrella River.
293 Costa Rica MrockNE—OLSSON 121
Phos subsemicostatus Brown and Pilsbry Plate 9, figure 15
Phos subsemicostatus Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
hilary Vols) 6351 p41350.) ple 25) figai 3.
A larger and finer sculptured species than the Dominican
Phos semicostatus of Gabb. A single imperfect specimen was col-
lected from Zone E of Saury Creek.
Gatun Stage: Gatun (Brown and Pilsbry)
Zone E,\ Saury (Creeks Oak.
Phos Veatchi, n. sp. Plate 9, figures 2, 3
Shell large and heavy, with a moderately long, coarse spire
of about 9 whorls; nucleus small, of 2 smooth whorls; whorls of
the spire moderately convex, with strong ribs continuous from
suture to suture; sutures appressed and bordered below by a
small spiral; the last whorl is widest about the middle and con-
tracted below to the short anterior canal; sculpture of heavy,
persistent ribs and regular raised spirals; the last whorl has 8
ribs; the spirals consists of raised threads, separated by wide, in-
terspaces; an occasional secondary spiral will appear in the in-
terspace but they are generally smooth or only obsoletely sculp-
tured by the growth lines; there are 8 spirals on the spire-whorls,
16 or 17 on the last whorl; aperture ovate, with the outer lip
thickened by the last rib, and internally with 9, long, entering
liree; inner lip formed by a raised, spreading callus, armed above
by a tooth-like denticle and a strong fold below on the edge of
the canal.
Length 43, diameter 22, last whorl 28, spire 25 mm.
A large, robust species from Hill 1a of the Banana River. It
differs from the szbsemzcostata by its strong persistent ribs and
coarser spirals.
It is named for Dr. A. C. Veatch, Director of the Explora-
tion work of the Sinclair Oil Corporation, under whose direction
the Costa Rican collections were made.
122 BULLETIN 39 204
Gatun Stage: Hill ra Banana River.
Phos beteyensis, n. sp. Plate 9, figure 6
Shell of the type of subsemicostata; whorls 3, moderately
convex; sculpture with the axial element very weak but present
on the penultimate whorl as fine, sub-obsolete, narrow ribs or ir-
regular wrinkles on the shell surface; they are absent from the
last whorl; the spiral sculpture consists of a broad zone, about
the upper 1-3 of the spire whorl on which are irregularly distri-
buted about 5 raised spiral threads; below this zone, the sculp-
ture is largely in the formof broad uneven bands formed between
incised spiral lines; on the last last whorl, this later zone occup-
pies the middle of the whorl; the zone of raised spirals above the
more smooth area of banded spirals gives to the whorls a slightly
coronated appearance.
Length 27 (2% whorls), diameter 15, last whorl 21 mm.
The type specimen is very incomplete, but its characters are
so distinctive and unlike any other species of Phos which we
have seen that its discription is advisable. Like the swédsemzcost-
atus the longitudinal ribs or coste fade out on the later
whorls.
Gatun Stage; Betey Creek, C. R.
Phos meiuloides Dall Plate 9, figure 16
Phos metuloides Dall, 1896, Proc. U. S. Nat. Mus., vol. 19, p. 310, pl.
28, fig. 15.
Phos metuloides Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila.,
vol. 63, p. 350.
Phos metuloides Maury, 1917, Bull. Amer. Pal., vol. 5, p. 252, pl. 14,
fig, 17.
A small species with a neat sculpture recalling that of J/e-
tula cancellata and gabéi. It isnot rare in the Gatun beds of
Mt. Hope in the Canal Zone. Dall also records it from the Mio-
cene of Santo Domingo.
Length 19, diameter 9, spire whorl 13.5, spire 10 mm.
Gatun Stage: Mt. Hope.
295 CosTA Rico MiockENE—OLSSON 123
Genus ALECTRION Montfort
Alectrion ranuncula, n. sp. Plate 9, figures 20, 22
Shell of medium size, with a large body-whorl and a sharp
pointed spire; the general form and sculpture recalls the recent
A. acuta Say; spire about the same length as the aperture and
canal and composed of about 7 whorls, of which the first 2 belong
to the small, smooth nucleus; the profile of the spire-whorls is
convex with deep, excavated sutures; sculpture of narrow ribs,
the tops of which are crossed by spirals and separated by wide,
smooth interspaces; the last whorl has about g ribs which are
large and humped on the ventral face, finer and more numerous
on the back; there are 3 spirals on the spire-whorls and 7 on the
last with several more on the short canal; aperture subcircular
with a large, heavy outer lip, internally denticulated.
The general form and sculpturing recalls the recent East
Coast <Alectrion acuta Say, but the Costa Rican shell is much
broader as indicated by the following comparative measurements
with 4. acuta Say, of the same height.
Height 14.00, diameter 6.25, last whorl 8, aperture 6 mm.
(Alectrion acuta Say)
Height 14.50, diameter 8.25, last whorl 9g, aperture 7.5 mm.
(Alectrion ranuncula, n. sp.)
It is not uncommon in the Gatun beds of the Banana
River.
Gatun Stage: Hill ra, Banana River.
Coll. 7 Estrella.
Alectrion losquemadica Maury Plate 9, figure 21
Alectrion losquemadica Maury, 1917, Bull. Amer. Pal., vol. 5, p. 255,
pls wigsy 225 23%
The Costa Rica fossil is similiar in its sculpturing and form
to the common recent West Indian A. ambigua Montfort. As
Maury has pointed out in regard to the Dominican examples, the
124 BULLETIN 39 296
difference between losguemadica and ambigua, liesin the proto-
conch, which in typical amdéigua is short and blunt, but pointed
and composed of several whorls in /osquemadica. Our shells are
larger than typical lJosquemadica from Santo Demingo.
Height 11.50, diameter 7.00 mm.
Gatun Stage: Port Limon.
Coll. 7, Estrella River.
Alectrion merenda, n. sp. Plate 9, figure 19
Shell small, a large body whorl anda sharp pointed spire;
nucleus of 2 small, convex whorls followed by 5 post-nuclear
whorls; sutures distinct but not excavated; the sculpture con-
sists of about 12 ribs which are variable in their strength and
spacing; the ribs are crossed by subequal spiral cords separated by
wide, interspaces in which there may appear a smaller intermed-
iate spiral; the suture is bordered by a set of 2 or 3 finer spirals;
there are 3 spirals on the spire-whorls and 8 on the last whorl
above the contracted zone of the canal; aperture subcircular with
a thick, heavy outer lip.
Height 7, diameter 4.5 mm.
The aperture is completely filled with a hard matrix which
completely conceals the characters of the inner and outer lips.
Uscart Stage: Old Man Sam Creek.
Genus NORTHIA Gray
Northia northiae Gray, var. miocenica, n. var. Plate 9, figures 7, 14
ef. orthia northie Gray, 1834, in Griffith’s Cuvier, pl. 30, fig. 2.
ef. Northia northie Kiener, 1832 Icon., Buccinum, p. 23, pl. 9, fig.
28.
The Northia northie Gray, (NV. serrata Dufresne) is a
common and characteristic West Coast or Pacific species, ranging
from the Bay of Panana to Guayaquil Ecuador. The fossils rep-
resented by three incomplete specimens differ only from recent
examples from Panama in lacking the hump-like thickening in
2907 Costa Rica MIocENE- OLSSON 125
back of the outer lip and the corresponding shoulder to its whorls.
But this is not an importnnt character and recent examples may
sometimes lack this character.
But few investigators who have studied the West Iadian
Miocene have failed to note the large percentage of species, close-
ly related or even identical with recent Pacific species, a condition
indicative of the union of the Atlantic and Pacific, permitting a
free intermingling of their respective faunas. Late during the
Miocene, this union was brought to a close or largely restricted,
and since then the Pacific element has gradually given way before
the encroachment and development of the West Indian. That
this extinction has been a gradual one is shown by the Pleisto-
cene deposits of Panama containg several common West Coast
Species not known from the recent Caribbean. Such species are
the Pecten veniricosus Sowerby and Norihia northie both record-
ed by Dall from the Pleistocene of Panama or Costa Rica.
Gatun Stage: Middle Creck.
Banana River, Hill No. 3.
Goll OmiRed Glog Greek:
Genus COLUMBELLA Lamarck
Columbelia submereatoria, n. sp. late 10, figures 33, 34
Shell ovate, solid, a small conic spire and a large body-
whorl; whorls about 7; spire-whorls but slightly convex so that
the profile of the spire is nearly plane; last whorl large, broadly
convex about the upper 1-3 and sloping evenly below to the
short, produced anterior canal; sculpture of subobsolete spirals,
there being about 6 on the spire-whorls and about 22 on the last
whorl; aperture linear with a thickened outer lip, expanded in
the middle and finely and evenly crenulated throughout; anteri-
or canal short and armed with 8 strong denticles.
Length 17, diameter 10.5, aperture 11, spire 5.5 mm.
Closely related to the recent C. mercatoria Linn., common
along the northern Costa Rican ccast, but differs in its more
126 BULLETIN 39 298
globose form and finer spiral sculpturing. Traces of coloration
are still preserved as faint blotches of yellow scattered over the
back of the shell.
Typical Columbella has heretofore not been recorded from
beds older then the Pliocene either in America or Europe.
Gatun Stage: Zone 5, Red Cliff Creek.
Genus STROMBINA Moerch
Strombina ambigua Guppy Plate 10, figure 9
Columbella ambigua Guppy, 1866, Quart. Jour.Geol. Soc. London, vol.
22, p. 288, pl. 16, fig 8.
Strombina ambigua Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3,
pt. 6, p. 1584.
This species was described from the Bowden beds of Jamai-
ca, of Miocene age. Our shells from Costa Rica agree exactly
with Guppy’s figure in the Quarterly Journal.
A large species of Columbelloid aspect due in large measure
to the slight thickening of its outer lip. The sculpture consists
of about 25 ribs, which are smooth on the spire whorls and upper
half of the last. The base of the last whorl and the canal have
in addition about 14 spiral cords.
Length 27 (7 plus whorls), diameter 10, spire 15, aper-
ture 13 mm.
Gatun Stage: Banana River.
Strombina Lessepiana Brown and Pilsbry Plate 10, figures 13, 20
Strombina lessepiana Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 352, pl. 25, figs. 11, 12.
Columbella (Strombina) gatunensis Toula, 1911, Jahrb. der K-K,
Geol. Reichsanstalt, Wien, vol. 61, p. 501, pl. 30, fig. 8.
An abundant species in the Canal Zone and with the excep-
tion of S. ambzgua, the largest species of Stvombina in the Mio-
cene beds of Panama and Costa Rica. The spire is long and
smooth, with the exception that the upper sutural zone is usual-
299 Costa Rica M1ocENE—OLSSON W277
ly distantly raised into small, longitudinal plicsee which is most
characteristic. An average specimen from the Canal Zone will
measure:
Length 25, diameter 10, spire 13, aperture I2.50 mm.
We collected from the lower Gatun sandstones of Cocles
Creek, near Old Harbor, Costa Rica, several small Strombinas
which except for size are typical Lessepiana.
Gatun Stage: Gatun and Mt. Hope, C. Z.
Cocles Creek, C. R.
Strombina matima, n. sp. Plate 10, figure 7
Shell ovate, moderately solid with a long evenly tapering and
sharp-pointed spire; nucleus small, pointed and of three, smooth
whorls; post-nuclear whorls 6; suture distinct and bordered an-
teriorly by a small sutural ridge; the sculpture consists of about
18, smooth, narrow ribs, which are lacking from the back of the
last whorl, which is smooth; the ribs are oblique in direction and
along the upper sutures are slightly knobbed by the sutural ridge;
the base of the last whorl and canal carries about 12 spirals; aper-
ture subelliptical with a heavy outer lip internally provided with
3 or 4 denticles about the middle; anterior canal straight and
smooth within.
Height 12, diameter 5.25, aperture 6 mm.
This is a species witha long, evenly-tapering spire and
small nucleus. The spire-whorls are sculptured with numerous,
small, smooth riblets. The riblets are obsolete on the back of
the last whorl which is entirely smooth except for the spirals
about the base and canal. .S. matima resembles S. Basst Maury,
from the Miocene of Santo Domingo but is more slender and has
a great many more ribs.
Gatun Stage: Hill ra, Banana River.
Coll. 7, Estrella River.
Strombina cricamola, n. sp. Plate 10, figure 18
Shell ovate, solid, with a large, convex and spirally sculp-
128 BULLETIN 39 300
tured body-whorl and a long attenuated spire; whorls 6 plus
(apex broken so that at least one complete whorl is lost from the
type specimen); the first 4 whorls are smooth, after which a
few incised spiral lines appear just below the upper suture; these
spirals gradually increase in strength and in number and on the
following whorls have become strong, raised spiral cords; the pen-
ultimate whorl has 5 spirals, the last about 24 (including those
of the anterior canal); the upper half of the last whorl and some-
times the penultimate, are longitudinally plicated with 18 rib-like
costz; aperture narrow, with a thickened outer lip, notched
above and obsoletely denticulated below; the inner lip is smooth;
canal short, twisted.
Length 16, diameter 8.5, aperture 9, spire 7 mm.
An unusually short and broad species, distantly related to
the C. pulcherrima Sowerby, recent from the Gulf of Dulce. Two
specimens were collected from the Gatun sandstones of Water
Cay.
Gatun Stage: Water Cay.
Strombina costaricensis, u. sp. Plate 10, figures 8, 17
Shell solid, ovate, with a long attenuated spire; last whorl,
large, flattened or contracted about the middle; whorls about 9%;
the first 6 whorls are smooth and form the long attenuated spire;
faint longitudinal plicee appear on the 7th, which increase in size
and form the rib-like costee on the later whorls; axial ribs persist-
ent and number on the last whorl about 15; the spirals begin
shortly after the first appearance of the longitudinal plice as
faint incised lines near the upper suture; these spiral lines are
further increased by the introduction of more lines below and be-
come on the following whorls raised spiral cords; the penultt-
mate whorl has 5 or 6 spirals, the last with about 20; the spirals
vary in strength on individual shells and on some specimens may
be lacking from the middle of the whorl, forming a smooth me-
dian band; aperture linear, small and coutracted and with a cal-
loused inner and outer lip ; outer lip strongly thickened, notched
above and feebly dunticulated below; canal short, straight.
301 Costa Rica Mr1ockNE—OLSSON 129
Length 15.5, diameter 7.5, spire 8, aperture 8 mm.
A species with an Azachis-like sculpture, the ribs and gen- -
erally the spirals being persistent. The body-whorl is flattened
or slightly contracted about the middle, but less so than in the
following variety. A long, tapering and nearly smooth spire as
in the preceding S. cricamola but much more slender and at-
tenuated.
Gatun Stage: Headwater of Middle Creek, C. R.
Strembina costaricensis, var. musanica, n. var. Plate 10, figures 26, 27
Shell ovate, with a long, conic spire of 8 or more whorls;
last whorl strongly contracted or indented about the middle; the
first 3 whorls or more (apex broken) are smooth or sculpturless;
on the following 3 whorls, the sculpture is largely of numerous,
straight rib-like coste, the spirals being only very faint and in-
distinct; on the last whorl the ribs number about 14; spirals
about 9 on the penultimate whorl as faint raised threads and on
the last whorl may be lacking from the middle or contracted
zone; aperture linear-ovate, a thickened outer lip with 3 faint
denticles below the upper notch; canal straight.
Length 15, diameter 6.5, spire 13, aperture 8 mm.
Probably a distinct species from the preceding and differing
in its more evenly tapering and more strongly sculptured spire,
and deeply contracted body-whorl. It resembles somewhat the
figure of S. carfagensis Brown and Pilsbry from Colombia, but is
smaller and has more numerous ribs. ‘The periphery of S. car-
tagensis is rounded.
Gatun Stage: Banana River, C. R.
East fork of Red Cliff Creek.
Strombina sincola, n. sp. Plate 10, figures 19, 21
Shell small, with a large, conic spire and a dorso-ventrally
flattened body-whorl; whorls about 8, the first 3 belonging to the
small, smooth, pointed nucleus; sculpture of the post-nuclear
whorls of numerous, straight, strong, longitudinal ribs, which are
730 BULLETIN 39 302
persistent to the penultimate whorl, but are laking from the
last whorl which is smooth and polished; the penultimate whorl
has about 15 ribs; the ribs are generally beaded about their up-
per extremities; sutures distinct; last whorl large and strongly
laterally flattened between a prominent ridge or hump on the left-
hand side of the whorl and the large outer lip; aperture sub-
elliptical with a large, expanded outer lip, deeply indented be-
hind; canal short and beak-like and spirally striated with about
8 threads on the back.
Length 7, diameter 3.25, spire 3.50, aperture 3.75 mm.
A small species abundant in the Gatun beds of Hill 1a, of the
Banana River. The last whorl is rather small and strongly flat-
tened dorso-ventrally. The spire is large and about % of the to-
tal length of the shell.
Gatun Stage: Hill ra Banana River.
fiill 3, Banana River.
Strombina chiriquiensis, n. sp. Plate 10, figures 14, 24
Shell small and in genral characters like the preceding but
with a larger body-whorl and a shorter, more conic spire; whorls
about 8, of which the first 3 are smooth and belong to the small
pointed nucleus; the succeeding whorls until the penultimate are
sculptured with fine,straight, longitudinal ribs,there being 19 on
the penultimate whorl; the last whorl israther large, smooth and
dorso-ventrally flattened between a hump-like ridge on the left
side and the large, expanded outer lip; canal short, beak-like and
spirally striated; aperture linear-ovate, deeply indented behind;
inner and outer lips are both heavily calloused, especially about
their posterior portions; over thiscallous and between the outer
and inner lips passes the small posteriorly directed canal; lower
half of the inner and outer lips denticulated with 8 or 9 strong,
denticles, heaviest above.
Length 7.50, diameter 4.25, spire 2.50, aperture 4.50
mm.
Of much the same size and general characters as S. s¢zcola of
303 Costa Rico MiocENE—OLSSON 13k
Costa Rica, but with a larger and broader body-whorl, and a
shorter, more conic spire. ‘The porterior portion of the inner
and outer lips are heavily calloused, a feature not developed in
S. sincola.
Gatun Stage: Water Cay.
Genus MUREX Linnzus
Murex messorius Sowerby
Murex messorius Sowerby, 1840, Proc. Zool. Soc. London, p. 137.
Murex rvecurvirostris Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
201. Not J. recurvirostris Broderip.
Murex messorius Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila.
vol. 63, p. 353.
Murex messorius Maury, 1917, Bull. Amer. Pal., vol. 5, p. 265, pl. 16,
figs. 1, 2.
This is the common Murex in Costa Rica. It is generally
small, seldom exceeding 40 mm in height. Each whorl carries
three primary varices, between which are 3 or 4 smaller ribs or
costz. It is arecent species in the West Indies.
Cammy Stage: Gatun, (GZ.
Water Cay, Panama.
Zone 5, Red Cliff Creek.
Coll. 7, Pumbri Creek.
Banana River.
Subgenus PHYLLONOTUS Swainson
Murex cornurectus Guppy
Murex (Chicoreus) megacerus Gabb, 1873, Trans. Amer, Phil. Soc.,
vol. 15, p. 202. Not WZ. megacerus Sowerby.
Murex cornurectus Guppy, 1876, Quart. Journ. Soc. London, vol. 32,
(OS) aig jolly keh pavers ite
Murex (Phyllonotus) cornurectus Maury, 1917, Bull. Amer. Pal., vol.
St DOV pI DOs Sey TO:
132 BULLETIN 39 304
A large species, possibly identical with the recent Murex
brevifrons Lamarck. ‘The varices are large and cross the spire
whorls in a spiral direction. Between the varices, there is gen-
erally a single rib-like elevation in the middle of the whorl.
Gatun Stage: Coll. 4, East Grape Point Creek.
Coll si ibed \Gligy Greer:
Rio Betey.
Soust Creek.
Genus TYPHIS Montfort
Typhis alatus Sowerby Plate 10, figure 15
Typhts alatus Sowerby, 1849, Quart. Journ. Geol. Soc. London, vol. 6,
p. 48, pl. 10, fig. 4.
Typhis alatus Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 203.
Typhis alatus Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 32,
Pp. 522.
Typhis alatus Brown and Pilsbry, 1911, Proc. Acad. Nas. Sci. Phila.,
vol. 63, p. 354.
A very characteristic species distinguished by the large wing-
like expansion of its outer lip. It is the largest species in the
Gatun and measures about 23 mm in height by 17 mm in diame-
ter. Itisfound also in the Miocene of Santo Domingo and in
the Bowden beds of Jamaica.
Gatun Stage: Gatun, C. Z. (Brown and Pilsbry)
Toro Cays, Panama.
Typhis linguliferus Dall, var. costaricensis, n. var. Plate r0, figs 22, 29
cf. Zyphis linguliferus Dall, 1890, Trans. Wagner Free Inst. Sci., vol.
By EP ta Ps r5 25 ployee mio
This is a smaller and more delicate speciesthan the 7. Gabdz
Brown and Pilsbry from the Canal Zone. They agree with the
Chipolan “inguliferus Dall in their general form but differ in their
uniformly smaller size and in nearly lacking the spine-like pro-
305 Costa RicA MrocENE—OLSSON 133
cess on the shoulder of primary varices. It is fairly abundant in
the Gatun beds of the Banana River.
Gatun Stage; Ffill ra, Banana River.
Genus S!STRUM Montfort
Sistrum nodulosum C. B. Adams Plate 10, figure 23
Purpura nodulosa ©. B. Adams, 1845 Proc. Bost. Soc. Nat. History,
Dens
Ricinula nodulosa Tryon, 1880, Manual of Conch., vol. 2, p. 190, pl.
59, fig. 275.
Sistrum nodulosum Dall and Simpson, tgo1, Bull. U. S. Fish. Com.,
VOI Te Penal.
Worn specimens of the species occur in the transitional beds
of Red Cliff Creek. They cannot be separated from recent ex-
amples from the present coast. It is very common as a recent
species on the rocks and reefs along the northern Costa Rican
and Panama coast, where it occurs about high-water level, asso-
ciated with the equally as abundant Lxgina turbinella Kiener
and Phasianella affinis C. B. Adams.
Uscari and Gatun transitional beds; Zone 5 and 6,
Red Cliff Creek.
(C) SUPER-FAMILY T4NIOGLOSSA
Genus DISTORTRIX Link
Distortrix similfima Sowerby
Triton simillimus Sowerby, 1849, Quart. Journ. Geol. Soc. London,
vol. 6, p. 48.
Persona simillima Guppy, 1866, Quart. Journ, Geol. Soc. London, vol.
22, p. 288, pl. 17, fig. 13.
Distortia simillimus Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
QU2 10%
Fersona simillima Guppy, 1874, Geol. Mag., p. 439.
134 BULLETIN 39 306
Persona simillima Guppy, 1876, Quart. Journ. Geol. Soc, London,
vol. 35, p. 522.
Distorsio ( Distortrix, Persona) gatunensis Youla, 1908, Jahrb. der K-K
Geol, Reichsanstalt, Wien, vol, 58, p. 700, pl. 25, fig. 10.
Distorsio gatunensis Brown and Pilsbry, 1g1t, Proc. Acad. Nat. Sci.
Phil., vol. 63, p. 356, pl. 29, fig. 8.
Distortrix simillima Maury, 1917, Bull. Amer. Pal., vol. 5, p. 271, pl.
17, Hes. 4,5.
Not common in Costa Rica. It also occurs in the Miocene
of Jamaica and Santo Domingo.
Gatun, Stages Gatun, Ga Ze
Banana River.
Port Limon.
Genus BURSA Bolten
Bursa crassa Dillwyn Plate 15, figure 15
Ranella crassa Dillwyn, Reeve, Conch. Icon.,fig. 18.
Ranella crassa Guppy, 1866, Quart. Journ. Geol. Soe. London, vol. 22,
p. 288, pl. 18, fig. 9.
Bursa crassa Gabb, 1873, Trans, Amer. Phil. Soc., vol. 15, p. 212.
Ranella crassa Guppy, 1874, Geol. Mag., p. 438.
Ranella crassa Guppy, 1876, Quart. Jouru. Geol. Soc. London, vol. 32,
Pen522:
Bursa crassa Maury, 1917, Bull. Amer. Pal., vol. 5, p. 272, pl. 17
figs. 6, 7.
This is a recent species in the West Indies. It occurs also
as a Miocene fossil in Jamaica and Santo Domingo.
Gatun Stage: Banana River.
Saury Creek.
Genus CASSIS Lamarck
Cassis sulcifera Sowerby
Cassis sulcifera Sowerby, 1849, Quart. Journ. Geo 1. Soc. London, vol
OF P47 supe ow sion i
307 Costa Rica MiocENE—OLSSON 135
Cassis sulcifera Maury, 1917, Bull. Amer. Pal., vol. 5, p. 274, pl. 18,
TONERS, gh Ad ey
This is a large species very common in Miocene rocks of
Santo Domingo but very rare in Costa Rica. Our record is bas-
ed on a single imperfect specimen from the Banana River and for
a figure of this species the reader is referred to Dr. Maury’s Do-
minican Fossils.
Gatun Stage: Flill No. 3, Banana River.
Cassis flamimea Linnaeus
Buccinum flammeum Vinneeus, 1758, Syst. Nat., ed. 10, p. 736.
Cassis filammea Reeve, 1848, Conch. Icon., vol. 5, pl. 5, fig. 12.
Cassis flammea Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd
series, p. 356.
This is a recent species recorded by Gabb, from his so-called
Pliocene clay beds of Costa Rica. Under this term, Gabb con-
fused two formations, one composed largely of coralline lime-
stones but with subordinate amounts of clays and sands equiva-
lent in large measure to the Gatun formations,and an overlying,
mostly sand and clays of true Pliocene age. Itis most likely
that the single example of Cassis fammea collected by Gabb,
was obtained from the upper or Pliocene formation.
Subgenus PHALIUM Link
Phalium moniliferum Guppy Plate 12, figure 11
Cassis monilifera Guppy, 1866, Quart. jouru. Geol. Soc. London, vol.
224) pi 287, pla 17, fg. 8.
Casidea granulosa Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 222.
Not C. granulosa Bruguiere.
Cassis reclusa Guppy, 1874, Geol. Mag., London. pp. 434, 439.
Cassis reclusa Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 32,
Pp. $25.
Cassis monilijera Guppy, 1876, Quart. Journ. Geol. Soc. London, vol.
32, P- 525.
Phalium moniliferum Maury, 1917, Bull, Amer. Pal.. vol. 5, p. 274, pl.
DOM MUSSe AS pl TOs) fol.
136 BULLETIN 39 308
The Costa Rican specimens like those of Santo Domingo are
extremely variable in their sculpture. The typical mzonzleferum
of Jamaica carries on the shoulder of the body-whorl one or two
rows of large tubercles or granules and Guppy’s figure shows the
spirals below as finely granulated.
In Costa Rica, the shells may vary from those with nearly
smooth spirals, through transitional forms in which only the su-
perior bandsare granulated, to the extreme in which all the spirals
are finely granulated over the whorl shell. A typical specimen
will average:
Length 35, diameter 22 mm.
Gatun Stage: Middle Creek.
Rio Betey.
Lone 5, Red Cliff Creek.
Genus SCONSIA Gray
Sconsia laevigata Sowerby, var. Gabbi, n. var. Plate 12, figure 3
ef Cassidaria laevigata Sowerby, 1849, Quart. Journ. Geol. Soc. Lon-
don, vol. 6, p. 47, pl. Io, fig. 2.
cf Cassidaria sublaevigata Guppy, 1866, /demm, vol, 22, p. 287, pl. 27,
fig. 9.
Cassidaria laevigata Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8,
2nd series, p. 356.
cf Sconsia sublaevigata Bose, 1906, Bol. Inst. Geol. de Mexico, Numero
22, p. 36, pl. 4, figs. 9, Io.
Sconsia laevigata Prown and Piisbry, 1911, Proc. Acad. Nat. Sci. Phila.
vol. 63, p. 356.
cf Sconsia laevigata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 275, pl.
19, fig. 2.
As Gabb noted years ago,the Sconsias of Costa Rica are not
the typcal Jaevigata of Santo Domingo but appear to approach
more closely in their striated whorls, the Jameican sublaevigata
_of Guppy and the recent s¢vzata of Lamarck.
Typical /aevigata, has generally a distinct shoulder to its
309 Costa Rica MioceNE-—-OLSSON Way
body-whorl and a lower spire. The spire-whorls are striated but
the spirals are lacking from the generally polished body-whorl.
The aperture of /aevigata is subrectangular in outline, rather
wide anteriorly and flaring posteriorly, while in Gaddz, the aper-
ture is narrow and but little larger anteriorly. In daevigata, the
callus of the inner lip is large and spreads widely over the colu-
mellar area, but its outer edge is raised and shelf-like, instead of
rounded and appressed as in Gaddz. Mature shells of laevigata,
have three large, heavy varices, marking the resting stages but
such varices are small and sometimes entirely lacking from the
Costa Rican shells.
Typical /aevigata will probably be found to be largely limit-
ed to Santo Domingo. ‘The Jamaica sud/acvigata (possibly a var-
iety) has more strongly sculptured whorls, a short spire and less
complete varices. Its aperture is that of true /aevigata.
The Sconsia laevigata was found by Dr. Maury, in her work
on the Paleontology and Stratigraphy of Santo Domingo to be
limited to her upper or Gurabo formation (the-Sconsza laevigata
zone). In Panama and Costa Rica, the Sconsza laevigata variety
Gabi, is frequently very abundant in the Gatun beds, but it is
apperently lacking from the underlying Uscari shales where it is
replaced by the new Scozsza cocleana and probably dbocasenszs.
Carn Stage Gatun, CZ,
Toro Cay, Water Cay, Panama.
Comadre Creek near Cahutta C. R.
TRAD: SEU er Tao (Os, SKE:
Sconsia bocasensis, n. sp. Plate 12, figures 12, 13
Shell like that of /aevigata,but narrow and with coarsely sculp-
tured whorls; spire more evenly conic, the apex of which is not
produced or attenuated; whorls coarsely sculptured with heavy,
persistent spirals threads, of which there are about g on the spire-
whorls and nearly 50 onthe last whorl; sutures appressed and
bordered anteriorly by a strong ridge-like cord or by stronger
spirals; the varices are but slightly developed; aperture subellip-
138 BULLETIN 39 310
tical, widest about the anterior 1-3; outer lip thickened and with
about 18, fine, liree-like denticles ; the inner lip with a thin spred-
ing callus and with about 16 long, narrow rugations.
Length 35 (last 2 whorls only) diameter 25 mm.
Differs from the variety Gadd of laevigata by its more narrow
shell, coarser sculpture and larger aperture. The suture is bord-
ered by a cord-like ridge.
Gatun Stage: Bocas del Toro.
Sconsia cocleana, n. sp. Plate 12, figure 7
Shell globose; spire low of about 7 whorls, the apex slightly
projecting; sutures distinct and deep, but not appressed; surface
coarsely sculptured with large, irregular, but narrow spirals
bands separated by deep interspaces which may in addition carry
one or more finer spiral threads; there are 7 or 8 spirals on each
spire whorl; on the last whorl the spirals are nearly twice as wide
as the upper third of the whorl and more anteriorly; no varices;
the outer lip is broken from the single specimen; inner lip is a
wide but thin wash of callus, without rugations or denticles of
any sort.
Length 46, diameter 32 mm.
The Uscari shales which everywhere in Costa Rica appear to
underlie the Gatun sandstones is very rich in the smaller types
of foraminifera, but mollusks are rare, fragmentary and difficult
to collect. Fragments of Scozsta, were noted by us at several
places which we believe belong to this species or to the docas-
Ensts.
his species will be recognized by its broad, globose shell
and small spire. It lacks the sutural cord so strikingly develop-
ed on docasensis.
Uscari Stage: Rio Cocles near Old Harbor, C. R.
gir Costa Rico MrocENE—OLSSON 139
Genus DOLIUM Lamarck
Subgenus MALEA Valenciennes
Matea elliptica Pilsbry and Johnson Plate 12, figure 2
Malea elliptica Pilsbry and Johnson, 1917, Proc. Acad, Nat. Sci. Phila.,
vol. 69, p. 169.
The true Malea camura Guppy, so common in the Miocene
of Jamaica and Santo Domingo still remains to be collected in
Costa Rica. It is usually a smaller species (height 60 mm) with a
heavier shell and high, narrow, special bands numbering about
16 or 17 rather widely separated.
The 1/. elliptica is larger (height 60-80 mm or more), more
globose and thin. The spiral bands numbering about 22 are
closely spaced and ribbon-like. The coiling is regular, while in
camura the coiling is irregular and descending in the latter
stages of growth.
The Malea elliptica was described but not figured by John-
son and Pilsbry from the Miocene of Santo Domingo.
Height 63, diameter 54 mm.
Uscart Stage: Red Cliff Creek,
Gatun Stage: Bocas del Toro. Estrella River.
fill ra, Banana River.
Genus CYPRAEA Linné
Cypraea parisimina, n. sp. Plate 12, figure 10
Shell small, more or less depressed and with expanded, an-
gulated sides; dorsal surface convex, but with a deep pit or
depression at its posterior end, beyond which project the 2
short, pouting ends of the lips of the aperture; ventral sur-
face flat, with a narrow aperture which is straight except near
at its posterior end where it is slightly curved; lips strongly
but regularly crenulated (about 21 on each lip) posterior sinus
140 BULLETIN 29 312
small and curved to the left; anterior sinus small and round-
ed.
Length 39, basalt diameter 25, height or vertical diame-
ter 17 mm.
A small shell with strongly depessed and angulated sides.
Collected from the coral limestones of Port Limon.
Gatun Stage: Port Limon,
Cpyraea cf. dominicensis Gabb
Cypraea Dominicensis Gabb, 1873, Amer. Phil. Soc., vol. 15, p.
236.
Cypraea dominicensis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 280,
pl. 19, fig. 11
Our collection contains a single, fragmentary shell from the
coralline limestones near Limon. ‘This specimen agrees closely
with the Dominican shell figured by Dr. Maury as C. dominicen-
sts ?, It is characterized by its very thin shell, cylindroid form
and straight, evenly denticulated apertrue.
Length 28, height 12 mm.
Gatun Stage: Port Limon.
Cypraea cinsrea Gmelin Plate 12, figure 8
Cypraea cinerea Gmelin, 1792, Syst. Nat., p. 3404.
Cypraea cinerea Reeve, Icon., vol. 3, pl. 22, fig. 124.
Cypraea cinerea Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd
series, p. 360.
The Cypraeca cinerea is a common, small species of Cowry
shell along the north coast of Panama and Costa Rica. Several
small Cypraeas were collected from the Gatun sandstones of
the Rio Blanco which cannot be distinguished from recent ex-
amples of cinerea.
Gatun Stage: Rio Blanco.
Cypraea almirantensis, n. sp. Plate 12, figure 9
Shell large, heavy and rather high; the outlines of the shell
313 Costa Rica MIocENE—-OLSSON 141
from below, the side and from above is nearly rectangular, but
with the anterior extremity more pointed; the shell is high (about
¥% that of the length), with nearly flat or slightly convex sides;
dorsal surface convex, a deep depression or pit about the posteri-
or 1-4 and low, but large tubercles on each side of the posterior
sinus; aperture narrow, curved, with the lips coarsely but regu-
larly dentate (about 20 on each lip); posterior sinus is long and
vertical; anterior sinus small and rounded; basal surface flat.
Length 60, basal diameter 39, vertical diameter 30 mm,
This is a species but distantly related to the C. Henkeni So-
werby, differing by its longer and higher shell, straight sides and
smaller dorsal tubercles. The aperture is strongly curved and
very deep on the posterior extremity, ending in the long, vertical
sinus.
Gatun Stage: Water Cay, Panama.
Genus STROMBUS Linnzeus
Strombus gatunensis Toula Plate 13, figures 5, 6
Strombus gatunensis Toula, 1909, Jahrb. der K-K Geol. Reichsanstalt
Wien, vol. 58, p. 673, pl. 25, fig. 7.
Strombus gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 355, pl. 26, figs. 3, 5.
But two species of Stromdbus are known from the Gatun beds
of Panama and Costa Rica, while five species occur in the synch-
ronous Miocene strata of Santo Domingo and four in Jamaica.
Other species may therefore be expected with further explora-
tion work in Costa Rica.
In S. gatunensis, the coiling of the whorls follow closely the
shoulder angle so that the last whorl appears very large and the
spire low but wide. Young shells occasionally have the shoulder
armed with few, large tubercles which later become nearly cov-
ered by the close coiling and appressed sutures. The last whorl
is large, nearly smooth and with a rounded shoulder angle.
Occasionally traces of the original coloration is preserved
142 BULLETIN 39 BIA
and consists of narrow, yellow V-shaped lines, the acute base of
which are directed backwards or away from the aperture and lie
along the center of the whorls. Similiar markings may be seen
on Veatcht and Guppy* has described the same markings for
pugiloides from Jamaica. Length 63, diam. 42 mm.
Gatun Stage: Gatun (OF
Water Cay, Panama.
fleadwaters of Middle Creek, C. R.
Strombus pugifoides Guppy Plate 13, figures I, 2, 3
Strombus pugilis Guppy, 1866, Quart. Geol. Soc. London, vol. 22, p.
287.
Strombus pugilis Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.-
233"
Strombus pugiloides Guppy, 1874, Geol. Mag., new series, vol. I, p.
433.
Strombus pugilis Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, p.
340.
Strombus pugilis Bose. 1906, Boll. de Inst. Geol. de Mexico, numero
22, p. 35, pl. 4, figs. 1-6.
Strombus pugiloides Maury, 1917, Bull. Amer. Pal., vol. 5, p. 284, pl.
20, fig. 6.
A smaller species than the gatunensts, from which it also
differs by its longer spire and more strongly sculptured shell.
The coiling follows below the shoulder of the whorls, which is
either simply rounded or bears several large, spine-like tubercles
asin the recent pugilzs. Resting stages develop large, hump-
like areas irregularly distributed on the spire-whorls. Varieties
occur which may be spirally sculptured over the whole shell or
the last whorl may be largely smooth. ‘The original coloration
is still preserved on some shells and consist as in gatumnenszs of
narrow, yellow, V-shaped lines, the apex of which lie about the
middle of the shell and directed backwards. It is very distinct
from the recent West Indian pugz/7s,which does not appear to de-
scend below the Pliocene.
* Proc. Sci. Assoc. Trinidad, 1873, vol. 2, p. 82,(reprint: Bull. Amer
Pal., vol. 8, p. 214).
OLSSON 143
315 Costa RicA MIocENE
Uscari Stage: Sapote, C. R. (Gadd)
Gatun Stage: Banana River.
Zone FE, Saury Creek.
Zont 5, East Grape Point Cretk.
Genus CREPITACELLA Guppy
Crepitacella limonensis, n. sp. Plate 15, figure 17
Shell small, melanoid in form,with a large body-whorl and
shorter conic spire; nucleus of 2 small, smooth whorls followed
by 5 post-nuclear whorls; sutures distinct; the whorls are slight-
ly coronated and crossed by oblique and slightly curved riblets
which become headed on the coronate edge of the spire-whorls
just below the upper sutures; on the last whorl, the ribs do not
pass below the middle; the surface is covered with faint spiral
threads, most distinct on the base of the body-whorl; aperture
elliptical, with a thin, oblique outer lip and a subobsolete anter1-
or canal; pillar straight.
Height 8, diameter 4.5, aperture 4 mm.
The Costa Rican shell, although possibly not mature,is much
smailer than the C. cepula Guppy from the Miocene of Venezuela
and Santo Domingo of the same number of ribs. It differs also
in having but 2 nuclear whorls, while cepuda has 3. The corona-
tion of the whorls which is such a striking feature of the Domin-
ican shell, is very much less and inconepicuous. The longitud-
inal costee are more numerous and produce a fine circle of beads
as they cross the small coronate edge of the spire-whorls.
Gatun Stage: Port Limon,
Boras del Toro.
Genus CERITH!UM Adanson
Cerithium costaricensis, n. sp. Plate 10, figure 28
Shell with a long, slender spire of about 12 whorls (tip brok-
ed); sutures deep but somewhat appressed; the profile of the
144 BULLETIN 39 316
spire-whorls is convex with an occasional hump-like resting mark;
the sculpture consists of primary spiral threads between which
lie 1, 3, or more finer secondaries; the earlier spire-whorls have
heavy ribs but they become replaced on the later by simple gran-
ulations on the primary spirals; a narrow, sloping band borders
the anterior side of the suture and carries finer, nou-beaded spiral
threads: below this band, the later spire-whorls bear about 4
beaded, primary spirals and finer secondaries; the last whorl shows
6 primaries together with other spirals on the base and canal;
aperture subcircular, with a moderately expanded outer lip, obli-
quely produced anteriorly.
Height 28, diameter 10.5, aperture 8 mm.
The general form and sculpture of this species is that of the
recent West Indian C. eburneum Bruguiere. ‘The fossil shell is
larger, more slender and differently sculptured.
Gatun Stage: Coll. 6, Red Cliff Creek.
Genus CLAVA Martyn
Clava costaricana, n. sp. Plate 10, figures 3, 4
Shell of medium size, cerithoid; whorls about 15, the early
spire-whorls very small; the last whorl carries a large hump on
the ventral side of the shell to the right of the aperture, so that
the shell appears flattened on this side; the sculpture consists of
strong spiral cords which are coarsely granulated by small, close-
ly spaced longitudinal cost; the upper spiral cord bordering the
suture is somewhat larger than the other; there are 3 spirals be-
tween the sutures of the spire-whorl and 6 on the last whorl with
additional smaller ones on the base of the shell; the outer lip is
broken on all our specimens; the columella carries 2 strong
folds.
Height 40, diameter 15 mm.
Al, I4 mm.
This species bears some resemblance to the C. caloosaénsts
Dall from the Pliocene of Florida. The Costa Rican shell is
317 Costa Rica MiroceNE--OLSSON 145
very much larger, broader and the longitudinal costze are more
rib-like, continued across the whorls, crossing the interspaces as
well as the spiral cords.
Gatun Stage: Banana River.
Clava alajuela, n. sp. Plate 10, figure 30
Shell small, slender; whorls 12 plus, the apex broken; the
sides of the whorls are straight and form the narrow, tapering
spire; sutures indistinct; the sculpture consists of nearly equal
ribs and spiral threads; the early spire-whorls have 3, equal,
spiral threads, separated by interspaces as wide as the spiral
threads and deep; on the later whorls fine, intermediate threads
appear in the interspaces and a fourth large spiral in the lower
suture; the spirals are crossed and strongly beaded by straight or
slightly curved, longitudinal ribs, which number on the last
whorl about 25 or 24; heavy and thickened resting marks ap-
pear on the later whorl on each volution; the aperture is broken,
the columella straight and witha single small fold.
Height 17 mm, diameter 6 mm.
A small and finely sculptured species, represented by the
single type specimen.
Gatun Stage: Middle Creek.
Genus SERPULORBIS Sassi
Serpulorbis papulosa Guppy Plate 12, figure I
Vermetus papulosus Guppy, 1866, Quart. Journ. Geol. Soc. London,
vol. 22, p. 292, pl. 17, fig. 3.
Vermetus papulosus Guppy, 1876, Quart. Journ. Geol. Soc. London,
vol. 32, p. 519.
Sespulorbis papulosa Maury, 1917, Bull. Amer. Pal., vol. 5, p. 291, pl.
22, fig. Io.
Considerable variation may be seen in the Costa Rican shells,
from specimens in which the sculpture is quite coarse and with
the interspaces between the main cords carrying only one strong
146 BULLETIN 39 318
intermediate cord, to more delicately sculptured forms with finer
tuberculated cords separated by wide and finely sculptured inter-
spaces.
This species occur also in the Bowden beds of Jamaica and
in the Gurabo formation of Santo Domingo.
Gatun Stage: Fiill 1, 2, Banana River.
Genus PETALOCONCHUS H. B. Lea
Petaloconchus sculpturatus, H. C. Lea Plate 14, figures 10, 15
g
Petaloconchus sculpturatus A. C. Lea, 1845, Trans. Amer. Phil. Soc.,
Vol./o,)\p-)2230 pl) 34 hig. ia!
Petaloconchus domingensis Sowerby, 1849, Quart. Journ. Geol, Soc.
London, vol. 5, p. 51, pl. 10, fig. 8,a be.
Patalocouchus sculpturatus Gabb, 1875, Trans. Amer. Phil. Soc., vol.
25, Pp. 240.
FPataloconchus sculpturatus Guppy, 1876, Quart. Journ. Geol. Soc. Lon-
don, vol. 32, p. 5109.
Vermetus (Petaloconchus) sculpturatus Dall, 1892, Trans, Wagner
Free Inst. Sci., vol. 3, pt. 2, p. 305.
Vermetus (Petaloconchus) pulcher Bose, 1906, Bol. Inst. Geol. de Mex-
ico, Numero 22, p. 32, pl. 3, figs. 22, 23.
Petaloconchus domingensis Brown and Pilsbry, 1911, Proc. Acad. Nat.
Sci. Phila., vol. 63, p. 359.
Petaloconchus domingensis Pilsbry and Brown, 1917, Proc. Acad. Nat.
Sci. Phila., vol. 69, p. 37.
Pctaloconchus domingensis Maury, 1917, Pull. Amer. Pal., vol., 5, p.
2O2s Pl, 22s) ite ME.
Pataloconchus domingensis Hubbard, 1920, New York Academy of
Science! Scei)(Sur. Py Ray voluaa pt 2 ip ail39:
The Antillean fossil was described by Sowerby as domingen-
sis. It has been by some writers, considered synonymous with
sculpturata, described a few years earlier by H. C. Lea from the
Chesapeake Miocene of the eastern United States, while others
have given domingensis the rank of a full species. A close com-
parative study of the Costa Rica fossils with several sets of typi-
319 ' Costa Rico MrocENE—OLSSON 147
cal sculpturata reveals no distinction between them. Both are
variable in their coiling from closely wound, narrow cylinders to
more open, umbilicate coils. The sculpture is much the same.
In the United States, this fossil is limited to the Upper
Chesapeake Miocene. In the Antillean and South American re-
gions, itis found in the Bowden beds of Jamaica, the Gurabo
formation of Santo Domingo, the Quebradillas limestone of Por-
to Rico, the Springdale beds of Trinidad, fromm the Gatun beds
near Cartagena, Colombia and Mexico.
Gatun Stage: Gaiun Stage, GC.) Z.
Water Cay, Panama.
Coll. 4, 5, East Grape Point Creek.
Comadre Creek.
Genus SILIQUARIA Lamarck
Siliquaria modesta Dall, var. limonensis, n. var, Plate 12, figure 4, 5, 6
ef Siliquaria modesta Dall, Mus. Comp. Zool., vol. 9, p. 39.
ef Siliguaria modesta Dall, 1889, Bull. Mus. Comp. Zool., vol. 18, p.
260, pl. 26, fig. 4.
This shell is very abundant in certain marly and sandy beds
in the coralline limestones of the Limon Peninsula and elsewhere.
Specimens in perfect condition are very difficult to obtain, the shell
being very fragile and delicate. There are no longitudinal threads
and the shell is smooth, except for the growth lines which are
sometimes a little irregular.
We have associated this shell with the .S. szodesta of Dall, a
recent species and recorded by Dr. Dall from several stations in
the West Indies and the Gulf of Mexico, in waters ranging from
94 to 805 fathoms in depth. The fossils differ in being more
loosely coiled and at the later stages long drawn out.
Dr. Maury’s S. guarabensts from the Miocene of Santo Do-
mingo is more irregular in its habitus of growth and the surface
of the shell carries faint longitudinal threads. Gabb has des-
148 BULLETIN 39 320
eribed as sculpturata a Siliquaria from the Limon Peninsula.
This is united by Dall with the recent West Indian sguamata
Blainville. I have seen no spicimens and Gabb may have collect-
ed it from the true Pliocene beds of the Limon Peninsula.
Gatun Stage; Island of Bocas del Toro.
Port Limon.
Genus TURRITELLA Lamarck
Turritelia gatunensis Conrad Plate 14, figures 12, 13
Turritella gatunensis Conrad, 1857, Pacific R. R. Report, vol. 6, p. 72,
PLS whe20:
Turritella conradi Toula, 1909, Jahrb. der K-K Geol. Reichsanstalt,
vol. 58, p. 694, pl. 25, fig. 4.
Turritella gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci-
Phila., vol. 63, pl. 27, figs. 4, 5, 9.
The Zurritella gatunensis is a medium sized shell seldom
over 40 mm in length. The base of the whorl overhangs the next,
and each whorl is encircled by a median concave or constricted
band. The whole surface it finely sculptured with small spiral
threads. The spirals defining the median band are usually slight-
ly heavier as well as a few about the upper part of the base.
The 7. gatunensts is a common and widely distributed
species in the Gatun beds, but always less abundant than the
T. altilira or its varities. Like the altdliva, zones of 7. gatun-
ensis are frequent in some localities.
The 7. atacta Dall of the Tampa Silex beds, the 7. acropo-
ra fossilin the Pliocene of Florida and recent along the east
coast of United States and the West Indies, are related
species.
Gatun Stage: Gatun, C. Z.
Water Cay, Panama.
Boucary Creek, C. R.
32K Costa RicA MrocENE—OLSSON 149
Turritella mimetes Brown and Pilsbry Plate 14, figure 5
Turritella mimetes Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 357, pl. 27, fig. 1.
Turritella (Haustator) aff. Hanleyana Reeve- 7. lineolata (Kiener)
Toula, 1911, Jahrb. der K-K Geol, Reichsanstalt, vol. 61, p.
401, pl. 30, figures 6a and 6b,
A large, robust species in form and sculpture like the 7.
variegata Linnzeus, recent in the West Indies.
The whorls are straight and usually slightly overhanging
the lower sutures. Surface sculptured with primary, secondary
and tertiary spiral threads. The primary threads are irregularly
disposed and usually number about 7 or 8 to each whorl. Be-
tween the primaries are the finer secondaries and tertiaries.
Small shells may be mistaken for the gazunensis, but the
whorls are flat, without the median concave and constricted zone
and the sculpture is more coarse. The figured shell of 14 whorls
has a length of 70 mm and a diameter of 18 mm. A larger but
more imperfect specimen of but 4 whorls measures:
Length 50, diameter 20 mm.
Gatun Stage: Gatun, C. Z.
Turritella altilira Conrad, and varities.
Turritella attilira Conrad, 1857, Pacific R. R. Reports, vol. 6, p. 72,
pl. 5, fig. 19.
Turritella gabbi Toula, 1909, Jahab. der K-K Geol. Reichsanstalt, p.
695, pl. 25, fig. 5.
Turritella altilira Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 358, pl. 27, figs. 2, 3.
Of the species of 7urritella in the Gatun beds of Costa Rica
and Panana, the 7. adé/zra of Conrad and its varieties is the most
common and characteristic. The 7. fornata Guppy, from the
Miocene of Trinidad and Venezuela is a closely related species.
These magnificent 7urrztelle are preminently characteristic
of the West Indian and Caribbean Miocene, but probably deriv-
ed from earlier forms in the Oligocene. The stock continued in-
150 BULLETIN 39 322
to the Pliocene where its acme of development was reached in
the elegant 7: preattenuata Heilprin of Florida. It has left no
descendants in the recent fauna.
The shell is long and tapering, each whorl sculptured with 2
prominent beaded spiral cords, between which the surface is con-
cave or deeply channeled and may be smooth or with finer
spiral threads. The upper spiral cord is simple or double. Several
varieties may be recognized which differ in details of their sculp-
ture, size and form of shell.
T. altitira Conrad, typical Plate 14 figures 6, 7
This, the most elegant form is limited to the small Gatun
sedimentary basin of the Canal Zone and the adjacent Province
of Colon.
The shell is long, slender and with numerous whorls; each
whorl is strongly sculptured by the two, primary spiral cords,
each of which is bordered on adjacent faces by a smaller
spiral, between which lies a deeply channelled groove; small-
er beaded spirals occur in the median channel and are best devel-
oped on the spire whorls; the beading or granulation of the
spirals is produced by the arcuated lines of growth, the sinus of
which lies a little above the middle of the shell; the upper spiral
cord is larger than the lower and more strongly beaded and in
general becomes double on the later whorls.
Gatun Stage: Canal Zone and Province of Colon.
Var. chiriquiensis, n. var. Plate 14, figures 4, 8, 9, 14
Shell larger, less attenuated and more coarsely sculptured
than typical altliva; the 2 spiral cords are less strong, of more
nearly the same size and usually remaining simple; the 2 spiral
cords enclose a wide, concave area of about % the width of the
whorl; this concave band usually carries 2, 3, or more fine spiral
threads; the beading or granulation of the spiral cords is feeble
and best developed on the earlier spire whorls and in certain large,
323 Costa RicA MiocENE—OLSSON 151
gerontic shells the spiral cords may become nearly smooth.
This is the common form on the islands of the Chiriqui La-
goon, and of eastern Costa Rica. Like the typical a/tzlira it is
often a very abundant fossil, forming zones of Turritella in the
Gatun beds.
The sculpture is much less elegant than that of true al¢zzra,
and consists mainly of the 2, primary spiral cords, between which
lies a wide concave band. This band is but feebly sculptured by
small, spiral threads, or in some cases, is smooth.
In small matters of detail of sculpture and in the form and
taper of the shells, there is considerable variation. Such varia-
tion is usually of but local signifieance and in general the shells
collected at the same or nearby localities will agree closly with
each other but differ more or less from those of more distant
areas.
The 7. fornata Guppy from the Miocene of Venezuela and
Trinidad should probably be considered asa variety of alézlzra,
most closely allied to the chirzguensts. Its sculpture is like chzr-
Zquensts but with stronger spiral threads on its median band.
Gatun Stage: Water Cay.
Grape Point and Red Cliff Creeks, C. R.
Banana River.
Rio Blanco.
Var. costaricensis, n. var. Plate 14, figure II
Shell small; whorls less numerous and more rapidly increas-
ing so that the shell is porportionately broader and shorter; su-
tures deep; the 2 primary cords are but weakly developed, be-
tween which lies, the concave band as in chiriquiensis,; the band
carries 3, 4, or more strong, irregular spirals, but slightly weak-
er than the primaries; the granulation of the cords is weak and
irregular.
Much smaller than the preceding varieties (usually about 4o
mm) and with wider and more rapidly increasing whorls. The
152 BULLETIN 39 324
cords on mature shells are small and but slightly heavier than the
spiral threads on the concave band.
This type replaces the chiriguiensis in the Gatun sandstones
in the area of Upper Hone Creek and Boucary Creek. It is often
extremley abundant, associated with the Turritella gatunen-
STS.
Gatun Stage: Upper Hone and Boucary Creeks.
Turritella exoleta Linnzeus, var. limonensis, n. var. Plate 14, figs 2, 3
Shell turreted, the whorls increasing more rapidly in diame-
ter than typical evolefa; the nucleus is small, immediately follow-
ed by whorls carrying a median carina which gradually becomes
more anterior in position and later the lower carina of the adult
sculpture; the whorls are deeply concave or excavated about the
middle between 2 principal carinze; on the last whorl there is
seen to be a third carina which is concealed in the suture; the su-
tures are excavated, becoming generally deeper with age; the
surface is nearly smooth, except for very faint spirals and rough
growth lines which may be raised and sublamellose on the medi-
an concave zone.
Height 4o (6 later whorls) diameter 17.50 mm.
Height 35 (10 earlier whorls) diameter 10 mm.
This is probably a distinct species from the recent West In-
dian Turritella exoleta Linnzeus, differing in its more rapidly ex-
panding whorls. Faint spirals may generally be seen covering
the entire shell and the growth lines become raised and sublamel-
lose in the concave medial zone. It is acommon shellin the cor-
alline phase of the Gatun at Port Limon and elsewhere.
Gatun Stage: Port Limon.
Bocas del Toro.
Turritella oreodoxa, n. sp. Plate 14, figure 1
Shell large, solid and very slender; whorls numerous and
each but slightly larger than the preceding, slightly concave or
constricted about the upper 1-3, above which the edge of the
325 Costa Rica Mrocene#—OLSSON 153
whorl is enlarged, forming a cord-like expansion about the upper
suture; sutures indistinct and appressed; sculpture of fine, ir-
regular and somewhat wavy spiral lines and indistinctly alter-
nating in strength; growth lines as typical for Haustator (not
visible on the base of our specimen), commence at the lower
suture and pass straight and parallel to the longitudinal axis
of the shell to the middle of the whorl where they become
bent™and continue obliquely to the left or towards the aper-
ture to the upper suture.
Length (2% whorls), 49, diameter 23 mm.
A very rare species and represented in our collection by
only two imperfect specimens. It appears to be closely related
to the 7. cartagensis Pilsbry and Brown from the Gatun of
Colombia. The full-grown Costa Rican shell was probably
longer and more attenuated and each whorl is prominently
elevated or ridged just in front of the lower suture.
Gatun Stage: Rio Banana beyond Hill No. 5.
Genus ALABA A. Adams
Alaba turrita Guppy, 1896, Proc. U. S. Nat Mus., vol. 19, p. 321, pl.
28, fig. 7.
Guppy’s figure of his type specimen from Jamaica agrees
very closely with our specimen from the Estrella River. The
whorls are finely spiralled with low ridges which are obsolete
from the upper parts of the whorls. Each whorl carries 2 or
more broad, smooth varices formed during resting stages.
The last whorl is strongly angulated by these varices.
Height 5, diameter 2.25 mm.
Gatun Stage: Coll. 7, Estrella River.
154 BULLETIN 39 326
Genus ARCHITECTONICA Bolton
Architectonica granulata Lamarck Plate 13, figure 10 ,1I, 12
Solarium granulatum Wamrack, 1822, An. sans. Vert., vol. 7, p. 3,
Architectonica prespectiva Tuomey and Holmes, 1857, Pleioc. Fos. S.
Ce pe i2ouple26s sige:
Solarium granulatum Dall, 1892, Trans. Wagner Free Inst. Sci., vol.
BPE Pa sZOe
Solarium Villarelloi Bose, Inst. Geol. de Mexico, Numero 22, p. 30,
pl. 3, figs. 4-11.
Solarium gatunense Toula, 1908, Jahrb, der K-K Geol. Reichsanstait,
Wien, vol. 58, p. 693, pl. 25, fig. 3.
Solarium granulatum Maury, 1917, Bull, Amer. Pal., vol. 5, p. 295,
pl. 23, fig. 3.
The Costa Rican fossils are usually quite typical and not
to be distinguished from recent examples of granulata. ‘They
vary somewhat in their height and persistence of their spiral
sculpturing. The beaded and granulated character of the spir-
als generally becomes obsolete on the later whorls so that the
spiral bands appear smooth. The umbilicus is deep but nar-
row.
This is a stable species ranging from the lower Miocene
to the recent. Its recent distribution extends from Cape Hat-
teras on the north, southward through the West Indies and on
the Pacific side from the Gulf of California to Peru.
Height 20, greater diameter 43 mm.
19, 29 mm.
12.75 20 mm.
Gatun Stage: Gatun, C. Z.
Water Cay, Panama.
Goll; 2 "RaolGocles:
Banana River.
Rio Betey.
327 Costa Rico MIOCENE——OLSSON 155
Genus NATICA Scopoli
Natica canrena Linnzeus Plato 13, figure 9
Natica canrena Vinneeus, 1758, Syst. Nat., ed. 10, p. 776.
Natica canrena Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd
series, p. 357.
Natica canrena Brown and Pilsbry, 1912, Proc. Acad. Nat. Sci. Phila.,
p. 508,
Natica canrena Maury, 1917, Bull. Amer. Pal., vol. 5, p. 298, pl. 23,
fig. Io.
This is a recent species but also common as fossil in the Mi-
ocene of the West Indies and the States. It is a warm water
species so that in the Chesapeake Miocene of the eastern United
States, it is found only in the warm, upper or Duplin stage. We
collected several specimens at Gatun but none at Costa Rica.
Gabb however has recorded it as abundant from the Limon Pen-
insula but whether his specimens were obtained ,from the Mio-
cene or Pliocene is not known.
The species is distinguished by its strong, tangential, su-
tural plice, but the main surface of the whorl is otherwise
smooth.
Gatun Stage: Gatun, C. Z.
Natica Youngi Maury, var., cocleana, n. var. Plate 13, figure 8
cf. Natica Youngi Maury, 1917, Bull. Amer. Pal., vol. 5, p. 29y, pl.
22h ISS wells 2s
Shell subglobular with 5 rounded and convex whorls; sur-
face smooth but with irregular, coarse growth lines about the su-
tures; the surface is also faintly straited with weak spiral lines
as may be seen on several otherwise smooth Naticas; aperture
semilunar with a narrow callus on the inner lip, slightly thicker
about opposite the middle of the umbilicus; umbilicus small,
with the umbilical callus small and but poorly developed.
Length or height 25, diameter 34 mm.
156 BULLETIN 39 328
A single specimen from the Gatun of Cocles Creek. It agrees
almost exactly with Dr. Maury’s type specimen of Watica
Young? from the Miocene of Santo Domingo in size and form,
but differs in its smaller umbilicus and smaller umbilical callus.
The Costa Ricam shell has the aspect of a Meverita,
Gatun Stage; Cocles Creek, near Old Harbor, C. R.
Section STIGMAULAX Moerch
Natica Guppyana Toule Plate 13, figures 13, 14, 15
Vv
Natica (Stigmaulux) Guppyana Toula, 1909, Jahrb. der K-K Geol.
Reichsanstalt, Wien, vol. 58, p. 696, pl. 25, fig. 6.
Natica Guppyana Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 360.
This species stands intermediate in its sculpture and form
to the 4. canrena Linneeus and JV. sulcata Born, both recent and
fossil Miocene species. In Guppyana, the sculpture consists of
even, wide and deep sulcations, which cross the face of the whorl
from the upper suture to the umbilicus. They are in fact the
continuation of the tangential sutural plicee of VV. canrena. With
further growth these sulcations may become obsolete on the mid-
dle of the whorls until they are lacking from the greater part of
the shell except in the immediate vicinity of the upper sutures
and on the umbilical angle. This change in sculpture is not seen
to the same extent on all shells and some spetimens may remain
strongly grooved and sulcated even when large and mature. In
other cases (in general with shells from the Banana River)
only very young shells have the sulcation continuous across
the face of the whorl, but soon become smooth, leaving the plica-
tions only about the upper suture and on the umbilical angle.
Such shells resemble closely MV. canrena, but in addition to
the grooving of the umbilical angle, have a larger and more
exanded aperture.
The WV. sulcata Born still remains to be found in Costa Rica
and Panana. It is abundant as a Miocene fossil in Santo
329 Costa Rica MiIocENE—OLSSON 157
Domingo, Jamaica and Venezuela. It differs from the Guppyana
by its closer and more irregular longitudinal sculpture crossed by
faint spirals, producing a cancellated appearance. The umbili-
cus is larger and more ample.
Gatun Stage: Gatun, C. Z.
Water Cay, Panama.
Banana River, C. R.
Genus POLINICES Montfort
Polinices subclausa Sowerby Plate 13, figures 16, 17
Natica subclausa Sowerby, 1849, Quart. Journ. Geol. Soc. London,
vol. 6, p. 51.
Natica subclausa Guppy, 1866, Quart. Journ. Geol. Soc. London, vol.
22, p. 290, pl. 18, fig. 8.
Folinices subclausa Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Philia., vol. 63, p. 560.
Folinices subclausa Maury, 1917, Bull, Amer. Pal., vol. 5, p. 300, pl.
23, fig. 14.
This is a common fossil in the Miocene of Santo Domingo
and Jamaica. The Costa Rican shells are small, the largest
specimen in our collection measuring only 17 mm in altitute.
When viewed from the dorsal side, the shell has a distinct-
ly Nerita-like aspect, due to its low spire, close coiling and rap-
idly enlarging whorls and appressed sutures. The umbilicus is
of medium size, expanding below. ‘The inner lip is provided
with a thick, heavy callus which joins above with the outer lip.
At the upper edge of the umbilicus, the callus carries a deep,
transverse groove, a most characteristic feature.
Gatun Stage: Gatun, C. Z.
flill No. 3, Banana River, C. R.
Polinices Stanislas-Meunieri Maury Plate 13, figure 7
Polinices Stantslas-meunierit Maury, 1817, Bull. Amer. Pal., vol. 5,
Pp. 300, pl. 23, figs. 15, 16.
158 BULLETIN 39 330
The Banana River beds contain a large species of Polinices
which appears to be this Dominican species. It differs from the
recent mammtllaris Lamarck, of the Costa Rican and Panana
coast by its much narrow umbilicus. Its surface is lined with
very fine spiral threads,
Young shells may be mistaken for the swbclansa which oc-
curs in the same beds, and careful study is sometimes necessary
for their separation. The present shell is less heavy, with a
higher spire and more convex spire whorls. The umbilicus is
much narrower, anda smaller callus on its inner lip which shows
but faintly the transverse groove. The surface of sudclausa is
smooth, but in Stazzslas-Meunteri very finely lined with minute
spirals.
Gabb has described from Sapote, Costa Rica, a Matica emi-
nulotdes which in form resembles young shelis of the S¢anzslas-
Meuntert. Wis figure shows a different shaped umilicus and no
transverse groove on the callus of its inner lip.
Length or aititute 44, diameter 38 mm.
Gatun) Stage: ul ra 2 > sbanang liven. Cs Re
Subgenus NEVERITA Risso
Heverita nereidis Maury
Neverita nereidis Maury, 1917, Bull. Amer, Pal., vol. 5, p. 301, pl. 23,
figs. 17, 18.
Our collection from ths Lower Gatun of Red Cliff Creek,
contains a fragment of this very interesting species. The J.
nererdts described by Dr. Maury, from the Cercado formation of
Santo Domingo, is closely allied to the recent West Coast /V.
glauca Humboldt. The Miocene \V. zeretdis is less broad, with
a narrower umbilicus and a larger umbilical callus.
Gatun Stages) Goll. ol Reds Gla Greer: | 6. fe.
33% Costa Rica MrocENE—OLSSON 159
SUB-ORDER SCUTIBRANCHIATA
A. SUPER-FAMILY RHIPIDOGLOSSA
Genus NERITINA Lamarck
Section SMARAGDIA Issel
Neritina viridemaris Maury Plates 15, figure 22
Neritina viridis Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 242,
Not NV. viridis Linneeus.
Neritina (Smaragdia) viridemaris Maury, 1917,Bull. Amer. Pal., vol.
5, p. 316, pl. 24, fig. 11.
The Costa Rican specimens are a little smaller than typical
viridemaris from the Miocene of Santo Domingo; but are identi-
cai in other respects. They still retain their original coloration of
a pale sea-green marked with straight and zigzag black lines. It
is the Miocene precursor of the common, recent West Indian and
Caribbean XV. vzrzdzs Linné.
Gatun Stage-\Coll. 5, feed Clif Creer.
Middle Creek.
Genus CREPIDULA Lamarck
Crepidula plana Say
Crepidula plana Say, 1822, Journ. Acad. Nat. Sci. Phila., vol. 2, p.
226,
Crepidula plana Wall, 1892, Trans. Wagner Free Inst. Sci., vol. 3, pt.
2, p. 358.
Crepidula plana Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila.,
vol. 63, p. 360.
This common and well-known species has been recorded by
Brown and Pilsbry from the Canal Zone. We have a single spec-
imen from Water Cay, found in the interior of the specimen of
Solenostetra Vaughani var. medioamericana. This species first
160 BULLETIN 29 3324
appears in the Lower Miocene and has continued into the re-
cent fauna.
Gatun Stage: Gatun (Brown and Pilsbry).
Genus XENOPHORA Fischer de Waldheim
Xenophora conchyliophora Born
Trochus conchyliophorus Born, 1778, Mus. Caes. Ind., p. 333.
Trochus agglutinans Lamarck, Anim. sans Vert., vol. 7, p. 14.
Fhorus agglutinans Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15,
pi24n
Xenophora conchyliophora Dall, 1892, Trans. Wagner Free Inst. Sci.,
vol. 3, pt. 2, pp. 360-362.
NXenophora conchyliophora, Maury, 1917, Bull. Amer. Pal. vol. 5, p.
297.
One immature shell was collected from the upper Gatun
beds of Old Man Sam Creek. It is one of the few species which
has continued pratically unchanged from Upper Cretaceous times
to the recent. It is rare asa recent shell along the northern
Panama and Costa Rican coast. Gatun Stage.
Genus PHASIANELLA Lamarck
Phasianelfa mollis, n. sp.
Shell small, solid, ovate with a short, blunt spire anda
small narrow umbilicus; whorls about 4, somewhat convex: su-
ture distinct; surface smooth and usually showing no features;
in some cases, color markings are preserved as brown, wavy
blotches of flammules and with very faint suggestions of fine re-
volving lines which are so common on recent small Phasianella.
Height 3.25, diameter 2.75, aperture 1.50 mm.
The Phasianella punctata Gabb from the Miocene of Santo
Domingo, has a longer spire and different color markings. It is
muuch more like the recent P. ajfinzs than the Costa Rican mollis.
P. mollis is related to P. umbilicata d’Orbigny but differs in its
sculpture.
BAZ CosraA Rica Miocknez—OLSSON 16%
The smaller forms of Phastanedla, of which there are 2 or 3
species along the northern Costa Rican coast, are often extreme-
iy abundant on the rocks and reefs exposed to the heavy, pound-
ing surf. There they are associated with a host of other gas-
tropods, principal among which are several species of /Verzta,
Fissurella, Purpura and Chitons. The fossil shells therefore like
the preceding Weritina viridemaris and several others which we
have already called attention to, have drifted from some near-by
rocky coast.
Gatun Stage: Coll. 5, Red Cliff Creek.
Middle Creek.
Genus TURBO Linnzeus
Turbo saltus, n. sp. Plate 15, figure 12
Shell perforate, ovate-conic; spire high of about 6 convex
whorls, separated by deep, canaliculate sutures; the last whorl is
large and perfectly convex in form; sculpture consists of irregu-
lar, spiral cords or liree; the earliest spire-whorls are strongly
carinated by a single, strong spiral, above'and below which the
whorl is smooth; a second spiral appears above and the succeed-
ing whorls of the spire have 2 principal spiral cords or lire, and
g smaller ones; the last whorl has three spirals that are a little
stronger and many smaller ones of different sizes; the spirals are
smooth; aperture perfectly circular, a small, spreading callus on
the body-whorl just above the perforate base.
Height 17, diameter 15.5, aperture 7.5 mm.
Among the recent species of Zurbo the saltus is related to
the large 7. Spenglerianus Gmelin,on one hand and to the small-
er J. jilosus Kiener on the other. The shell is perforate,in which
character it approaches 7. /ilosus, but the sutures are canalicu-
late or channeled as in the non-perforate 7. Spenglerianus. The
sculpture consists of irregular, smooth spiral cords or lire of
which 2 or 3 about the middle of the shell are a little larger
than the others.
162 BULLETIN 39 334
Gatun Stage: Coll. 5, Red Cliff Creek.
Genus ASTRALIUM Link
Astralium brevispinum Lamarck, var. basalis, n. var. Plate 15, figs. 4, 5
The Astralium brevispinum Lamarck is a recent species
found living along the north coast of Panama and Costa Rica.
It is most easily distinguished from the West Indian 4. long7spr-
num Lamarck by having its columellar area brilliantly stained
with scarlet.
From upper Old Man Sam Creek in eastern Costa Rica, we
collected a single specimen of an Astralium, very similiar in most
of its characters to bvevispinum. It differs most importantly in
having a more strongly sculptured base. This sculpture consists
of a wide, flat band, inside of which are 3, heavy, nearly
equal spiral cords. The whole surface is crossed by squamose
lines of growth.
Height 20, diameter 29.50 mm.
Gatun Stage: Coll. 6, Old Man Sam Creek.
Astralium caelatum Gmelin, variety
Trochus celatum Gmelin 1792, Syst. Nat., p. 3531.
Astralium celatum Pilsbry, 1888, Manual of Conch., first ser., vol. Io,
p. 224, pl. 57, figs. 45, 46.
A large Turboid shell related to the recent 4. celatum Gme-
lin was collected from the coral limestones near Port Limon. It
is so firmly imbedded in its rocky matrix that only a small! por-
tion of its surface isexposed to view. Its surface sculpture is like
that of celatum, but the radial costee on the upper part of its
whorl are closely spaced and not far apart as in the recent shell.
Its height measures 55 mm, diameter 54.
Gatun Stage: Port Limon.
335 Costa Rico M1ockENE—OLSSON 163
Genus CHLOROSTOMA Swainson
Subgenus NEOMPHALIUS Fischer
Chlorostoma costaricensis, n. sp. Plate 15, figure 16
Shell conoidal, solid, witha narrow, but deep umbilicus; su-
ture distinct; the last whorl is strongly carinate or angled below;
above this angle, the whorl is evenly convex, below, the base is
nearly fiat; the sculpture consists of beaded spiral cords, there
being on the last whorl above the angled periphery 7 or 8 cords;
the interspaces are nearly as wide as the cords, and are finely
etched by oblique lines of growth and finer spirals; on the base,
the spirals are irregular, the strongest about the middle; there
are 2 or 3 cords about the upper part of the umbilicus, but the
interior wall is smooth.
Height 10.5, diameter 16.5 mm.
The transitional beds between the Uscari and the Gatun of
Red Cliff Creek, contain in addition to the usual fauna, several
species, as certain Cerithoids, Szstrum nodulosum, Modulus mo-
dulus, Nitidella and the present species, whose normal habitat
is on the rocks and reefs of the extreme, upper limit of the lit-
toral zone. They are always much broken and worn as if they
had been rolled and washed about by the surf on a rocky or sandy
beach and then drifted out to the deeper and quiter waters off
_ shore where depostion was going on.
Although all of our specimens of C. costaricensis are very
fragmentary and much worn, we have deemed it best to place
the species on record, as this subgenus has been recorded only
very doubtefully from beds older than the Pleistocene.
Uscari Stage: Zone 6, Red Cliff Creek.
Gatun Stage: Zone 5, Red Cliff Creek.
164 BULLETIN 39 336
Genus CALLIOSTOMA Swainson
Calliostoma limonensis, n. sp. Plate 15, figures 8, 1
Shell of medium size, conic; whorls about 8, flat or but
slightly convex, so that the profile of the shell is a broad cone;
sutures indistinct; the whorls are closely sculptured with fine,
beaded spirals of primary and secondary orders; each spire-whorl
has 7 or 8 primary spirals alternating with 1, 2 or 3 secondary;
periphery sub-angular; base but slightly convex and closely spir-
alled with flat or sub-obsoletely beaded spirals (about 20), umbil-
icus narrow but deep, smooth within.
Height 19, diameter 18.5 mm.
Fairly abundant in the coral limestones along the shore west
of Port Limon. Related to the recent C. jujubinum Gmelin, it
differs by its more globose form and small, pointed attenuated
apex.
Gabb hasalso described a small Callzostoma from the Limon
Peninsular, as C. Guppyana. This shell of 8 or 9 whorls meas-
ures but 5 mm in height. It is a higher and more conic form
with the whorls sculptured with beaded spirals.
Gatun Stage: Port Limon.
Calliostoma mancinella, n. sp. Plate 15, figures 9, Io
Shell of medium size; whorls about 8, with very indistinct
sutures and a coarse sculpture of beaded spirals; the spire whorls
are flat sloping evenly from the small projecting nucleus to the
broadly rounded periphery of the last whorl; sculpture of the
spire-whorls consists of about 6, strongly beaded spirals the
largest of which boundsthe upper sutures; between these spirals
are scattered a few smaller ones, the lower half of the peri-
phery and the base have about 12 additional spirals; these
spirals are larger, more widely spaced and are smooth except
the 4 or 5 around the umbilicus; the interspaces of the basal
spirals are finely incised by minute longitudinal lines, corres-
ponding to the growth lines; umbilicus narrow, but deep,
337 Costa RrcA MroceENE—OLSSON 165
smooth within; aperture rounded, with the outer lip strongly
oblique.
Height 15, diameter 21 mm.
Several specimens of this well-marked species were obtained
from the Gatun beds of eastern Costa Rica, in the vicinity of
Manzanilla Point. In the Santo Dominican Miocene it is relat-
ed to the C. Grabaui Maury, but the Costa Rican shell is much
lower and more depressed.
The C. Grabaui and the mancinella belong to the group
which contains the large, elegant C. Sayana, described by Dall
from deep water off the Hatteras coast.
Gatun Stage: Coll. 4, East Grape Potnt Cree.
fleadwaters of Middle Creek.
Coll. 5, Red Cliff Creek.
zr mile south of shore, along Old Man Sam
Creek, (Veatch)
Calliostoma castilla, n. sp. Plate 15, figure 6, 7
Shell rather large, in form and sculpture resembling the C.
phulanthropus of the Chesapeake Miocene of eastern United States,
but umbilicated; whorls 5 plus (apex broken), flat and forming
a broadly conic spire; periphery of the last whorl, subangular,
due to a broad, cord-like ridge, which appears as if formed
form an overlapping of the base; sculpture of the spire-whorls of
fine, alternating and beaded spirals, there being about 13 on each
whorl; the base is nearly smooth the spirals being nearly obso-
lete or as broad low bands; the spirals about the umbilicus are
slightly stronger and obscurely beaded; umbilicus deep and
narrow.
Height 20, diameter 25 mm.
A large species with an angulated periphery, fine, beaded
spirals above and a nearly smooth base. Buta single specimen
166 BULLETIN 39 338
was collected from the coral limestones near Port Limon, asso-
ciated with the common /zmonensis.
Gatun Stage; Port Limon.
CLASS SCAPHOPODA
Genus DENTALIUM Linnzeus
Subgenus DENTAL!UM, s. s.
Dentalium bocasensis, n. sp. Plate 15, figures 2, 3
Shell rather larger, solid, gently curved and increasing grad-
ually in size; the tip is more curved than the rest of the shell and
strongly hexagonal in cross-section; the sculpture consists at first
of 6 strong, primary ribs and wide, flat or slightly concave in-
terspaces; the ribs are further increased by the introduction of 6
intermediate ribs and still later by 12 more; these ribs are sub-
equal in size on the later portions of the shell; the interspaces are
finely transversely lined and faintly longitudinally striated.
Length 52, diameter 6.25 mm.
44; 4.75 tam.
The Dentalium bocasensis is fairly abundant in the shale
beds of Bocas Island. It is recognized by its strongly hexagonal
tip and finely transversely lined interspaces,and strong longitudi-
nal scuipture.
Gatun Stage: Bocas del Toro.
Subgenus FISSIDENTALIUM Fischer
Dentalium uscarianum, n. sp. Plate 15, figure 1
Shell large, solid and heavy; gently curved, and the taper of
the shell is slow and gradual; the tips are all broken on our
specimens, but the later sections of the shell are circular or slight-
ly compressed dorsal-ventrally; the surface is sculptured with
about 24, low, subregular and rounded longitudinal ribs with
339 Costa RicA MrockNE—OLSSON 167
smaller intermediate ones in the interspaces; the ribs become
gradually obsolete with age and the later portions of full-grown
shells are nearly smooth; fine transverse lines of growth cross the
surface and are best seen on the earlier portions of the shell;
operture oblique.
Length 60, diameter 12 mm.
This large Dentalium is abundant and very characteristic of
certain conglomeritic zones in the Upper Uscari shales. There it
occurs associated with a small but very distinctive fauna, includ-
ing the Ptychosalpinx? dentalis, Sconsia cocleana and Scaphella
costaricana and other species. When complete, the Dentalium
uscartanune frequently reached a length of 110 mm or more.
All our specimens are unfortunately lacking in the apical
tip. In general sculpture, it recalls the Dentalium floridense
Henderson dredged from 35 to 110 fathoms of water off Florida
and in the West Indies, but the taper of the shell is more gradual
and the sculpture more irregular, becoming obsolete with age.
Bose* has described a large species as Dentalium rimosum from
the Miocene of the Tehuantepec Peninsula. In that species, the
ribs number about31, are very regular with deep interspaces, and
ribs are finely etched by transverse lines of growth.
Useari Stage: Coco Plum, Panama.
Rio Coeles.
Margarita—Old Harber trail.
Comadre Creek.
Pumbri Creek, ete.
*Boll. Inst. Geol. do Mexico, No. 22, p. 55, pl. 3, figure 1, 1906.
juL ‘7 Wee
BULLETINS
OF
AMERICAN PALEONTOLOGY
Vol. 9
No. 39
The Miocene of Northern Costa Rica
With Nores On Its GENERAL STRATIGRAPHIC RELATIONS
BY
A. A. OLSSON
Part 2
CLASS PELECYPODA
(Pp. r69-Finis)
Yun 2T,) 1922
Harris Co.
Cornell University, Ithaca, N. ¥.
Wie Se sek.
ae f i
343 Costa Rrca MroceNE—OLSSON 171
ORDER PRIONODESMACKHA
Genus NUCULA Lamarck
Nucula orbicella, n. sp. Plate 28, figures 19, 20
Shell of medium size, broadly elliptical or subovate in out-
lines; obliquely truncated at its anterior end; gently convex; the
surface is covered with close, subregular, concentric wrinkles and
fine radial threads so that the sculpture is decussated; on the
posterior dorsal slope, they are larger and pustular-like and their
course is obliquely downward across the growth-lines; they are
weak on the poorly defined lunule; the radical threads are main-
ly between the concentric wrinkles but ventrally they cut into
the wrinkles as well; interior of the shell, silvery white or faint-
ly tinged with purple; the ventral margin is finely crenulated;
chondrophore very small, oblique; anterior set containtng about 8
teeth, the posterior with about 17.
Length 9.50, height 7.50, semi-diameter 2.25 mm.
II.00 8.50 2.50 mm.
From the recent VV. crenulata A. Adams, this shell differs
by its larger size and more elliptical and depressed shell. Gabb
has described from the Pliocene of the Limon Peninsula WVzw-
cula limonensts and moenensis, but both of these species are per-
fectly smooth and sculptureless. ue Mucula orbicella is a com-
mon species in certain marly beds intercalated in the coralline
limestones of Port Limon.
Gatun Stage: Port Limon.
Nucula cahuitensis, n. sp. Plate 18, figures 21-24
Shell small, solid, obliquely subtrigonal in form; ventral
margin widely rounded, a nearly straight posterior dorsal margin
123 BULLETIN 3g 344
and a widely truncated anterior end; valves gently convex; pro-
toconch on the tip of the umbo small, smooth and white; surface
polished; the sculpture of the disk of the shell is developed to a
variable extent, but the umbos are always smooth; typically the
lower half of the shell is covered with close, regular, concentric
wrinkles which are slightly decussated by fine, even, and slight-
ly raised radial threads; in other cases, the wrinkles may be pres-
ent only on the extreme anterior or posterior ventral extremities
and the radial lines showing only as a part of the shell substance
and not as raised threads; the escutcheon is elliptical and thickly
covered with raised wrinkles which lie at right angles to the dor-
sal margin of the shell; they are similiar on the lunule; interior
of the shell tinted with brown; ventral margin crenulated; chon-
drophore small, oblique with the anterior set of teeth numbering
about 7, the posterior about 14.
Length 3.50, height 3.00, diameter 1.60 mm.
This is a small species of subtrigonal outlines, suggestive of
the Pacific VV. extgua Sowerby in general form and size. Onthe
few specimens of V. exiywa, which I have from the Bay of Pan-
ama the sculpture is finely decussated over the whole shell with
fine radial and concentric threads. The most striking features of
cahutiensts are its strongly sculpture lunule and escutcheon.
Gatun Stage: Zone G, Saury Creek.
Nucula tenuisculpta Gabb Plate 18, figure 25
Nulua tenuisculpta Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
255.
Nucula tenuisculpta Maury, 1917, Bull. Amer. Pal., vol. 5, p. 325, pl.
26, figure 8.
A species of the size and form of cahuztenszs but the valves
are entirely smooth and polished. ‘The lines of growth are fine
and irregular. The radial lines are faint and sub-microscopic.
Inner margin crenulated.
Height 3.25, length 4.00, semidiameter .75 mm.
Gatun Stage: Coll 4, Red Cliff Creek.
345 Costa RicA MrocENE—OLSSON 78
Genus LEDA Schumacher
Leda Balboae Prown and Pilsbry i Plate 18, figure 15
Leda Balboe Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila.,
vol, 63, p. 362, pl. 27, fig. 8.
This species is fairly common in Costa Rica, but generally
smaller than in the Canal Zone, where it frequently reaches a
length of 22 mmior more. Its valves are moderately convex and
regularly sculptured with even, close, concentric riblets which on
the center of the shell disk may be slightly appressed. The
escutcheon is well-defined and similiarly sculptured with the rib-
lets running parallel to the hinge margin.
Gatun Stage: Gatun, C. Z.
Coll. 1, East Grape Point Creek.
Zome G, Saury Creek.
Outtana Creek.
fiill No. 3, Banana River.
Leda Davilae, n. sp. Plate 18, figure 20
Shell small, solid, strongly convex; the beaks nearly cen-
tral; the rostrum is short, pointed and sharply defined from the
rest of the shell disk by a high, cord-like ridge; the anterior end
is bluntly rounded and carries 2, low flexures from the beaks to
the anterior ventral extremity; lunule small and narow, lacking
the concentric ribs and sculptured with small, pustular-like
ridges passing at right angles to the hinge-line; the sculpture con-
sists of about 30 concentric riblets which are somewhat larger
and with deeper interspaces on the umbos; the interspaces show
on the umbos and along the anterior slope, fine, radial lines or
punctations; interior of the shell cavity deep, with a heavy
hinge; the anterior and posterior sets of teeth number about 13
or 14, are high and V-shaped.
Length 5.75, height 4.00, semi-diameter 1.75 mm.
174 BULLETIN 39 3246
A small, plump species, coarsely sculptured with even, con-
centric riblets, separated by deep interspaces. These interspaces
may carry fine, radial lines or punctations at the base of the con-
centric ribs and are best seen on the umbones and the anterior
slope. The ZL. indigina Dall (L. dbisulcata Guppy) from the
Miocene of Jamaica is similiar but proportionately longer.
Gatun Stage: Coll. 4, Red Cliff Creek.
Bocas Island.
Subgenus ADRANA, H. and A. Adams
Leda quitanensis, n. sp. Plate 18, figure 19
Shell thin, elongate but slightly convex; beaks very low and
scarcely distinguishable from above and situated at the anterior
I-3; ventral margin an even curve from the pointed extremities;
dorsal margin straight; surface polished but covered with very
fine, concentric lines which are crossed on the lower half of the
anterior 2-3, by even, oblique lines (.Sczssuda-like); these oblique
lines are spaced about .25 mm apart; interior concealed.
Height 7.25, lenght 26.00, semidiameter .75 mm.
A much smaller species than the next (emsznoides), slightly
nlore convex and with beaks situated more anteriorly. The sur-
face is finely sculptured about the middle and the anterior por-
tion by slightly oblique lines, as seen on certain Tellinoids such
as Scissula. The Manzanilla beds of Trinidad contain Leda
(Adrana) Guppyi Dall (Cercomva ledeformis Guppy). It is less
elongate, more contracted posteriorly and has a different surface
sculpture. Dall has also recorded Leda Guppyi from the Bow-
den beds of Jamaica.
Gatun Stage: Quitana Creek.
Zone F, Saury Creek.
Coll. 1, Fast Grape Point Creek.
Leda ensingides, n. sp. Plate 18, figure 12
Shell rather large, elongate, depressed and subequilateral;
347 Costa RicA M1ocENE—OLSSON 175
beaks very small and scarcely distinguishable from above, and
situated nearly about the middle of the dorsal margin; extremi-
ties bluntly rounded, with a broad, even curve to its base or ven-
tral margin; the dorsal side is nearly straight but descending
slightly towards its extremities; surface nearly smooth, the
growth lines indistinct, but with fine, even very slightly oblique
lines on the anterior two-thirds of the shell; interior con-
cealed.
Length 54.00, height 15.00, diameter 4.50 mm.
Differs from the preceding species by its larger size, nearly
central beaks and by its finer, more concentric sculpture. ‘There
are several species of this group recent in the West Indies and
along the Pacific coast of Central America, but none appear at
all closely related to the Gatun species.
Gatun Stage: Hill No. 3, Banana River.
Leda Dalliana, n. sp. Plate 28, figure 17
Adrana sp ? Dall, 1898, Trans. Wagner Free Inst. Sci., vol. 3, pt. 1
4, P- 592.
Shell small, thin, elongate, depressed; the small, scarcely
differentiated beak is sutuated about the anterior one-fourth of
the shell; the anterior side is somewhat contracted and shortly
rounded while the posterior side is produced and 3 times the
length of the anterior and bluntly pointed at its extremity; the
rostrum is long and narrow, formed by 2 radial, scabrous threads,
separated by a wide interspace; concentric sculpture of thin,
elevated threads or Jamellae, regularly distributed on the lower
half of the shell, but widely spaced on the umbonal area; the
escutcheon is long and narrow, sculptured with threads parallel
to the hinge-margin; interior of shell cavity shallow, the long
hinge with an anterior set of 20 small teeth, the posterior with
26 or 27.
Length 10.50, height 3.50, diameter of left valve .75 mm.
176 BULLETIN 39 348
A much smaller species than the preceding and differing in
its more anterior beaks, depressed shell and heavier concentric
sculpture as well as in other characters. It is probably the shell
which Dall refers to as having fragments of from the Pliocene
of Limon. But a single perfect specimen was collected
Gatun Stage; Port Limon.
Genus YOLDIA Moller
Yoldia bocasensis, Plate 28, figures 21, 22
Shell small, thin, ovate-elliptical in form; beaks situated
slightly in front of the middle; vaives moderately convex; both
the anterior and posterior extremities are broadly rounded and
connected by the gentle curve of the basal or ventral margin; the
umbos are very small! but distinct and lie just above the greatest
convexity of the valves; surface smooth, polished with no other
sculpture than the incremental growth-lines; chondrophore dis-
tinct, scarsely oblique and bordered on each side by an equal num-
ber of V-shaped hinge-teeth; the anterior set contains about 14
teeth, which are very small next to the chondrophore becoming
progressively larger distally, the posterior set with 13 or 14; the
pallial sinus is indistinct, but rather large, broad and extends
nearly to the middle of the valve.
Length 13.00, height 7.00, semidiameter 2.25 mm.
9.50 5.50 1.50 mm.
A fairly common species in the clay beds of Bocas Island
where it is associated with a fauna of rather deep water charac-
ter. It has much the form of certain deep-water Yo/dzella but
the interior of the valves show an indistinct but deep and wide
pallial sinus.
Gabb has described from the Miocene of Santo Domingo, a
Yoldia ovalis, which has never been figured and its description is
very meager and insufficient. .
Gatun Stage: Bocas del Toro.
349 Costa Rica M10cKNE—OLSSON 177]
Genus LIMOPSIS Sassi
Limopsis menilis, n. sp. Plate 26, figures 23, 24
Shell small, solid, moderately convex and subcircular in form
but slightly oblique; the hinge-line is straight, the anterior dor-
salsubmargin very small, the posterior large undifferentiated
from the rest of the shell ; the umbos are high and prominent,
lying a little in front of the middle; exterior sculptured with
strong radial ribs which are largest and strongest on the middle
of the shell disk, finer on the dorsal submargins; these ribs are
crossed by high widely spaced concentric lamellae, more closely
spaced on the umbos; the lamellce on crossing the ribs are raised
into short, spine-like fringes which are directed ventrally; inter-
ior of shell deep; the hinge-line with a small cartilage pit at the
anterior 1-3 of the shell; it is bordered on the anterior side by a
set of 3 high, projecting teeth and 4 onthe posterior side, the
ventral and a part of the anterior margin is crenulated, most fine-
ly in the middle and quite coarsely at the posterior portion.
Length 1.75, height 2.00, semidiameter .60 mm.
This is a small, subcircular species with a crenulated inner
margin and a strong surface sculpture of ribs crossed by distant
concentric fringes. It is fairly common at its type locality, giv-
en below.
Gatun Stage: Old Man Sam Creek, 1 mile south of beach
(A. C. Veatch, collector).
Genus GLYCYMERiS, Da Costa
Glycymeris canalis Brown and Pilsbry Plate 18, figures 2-7
Glycymeris canalis Brown and Pilsbry, Igt1, Proc. Acad, Nat. Sci.
Phila., vol. 63, p. 364, pl. 28, fig. Io.
Glycymeris trilobicostata Pilsbry and Brown, 1917, Proc. Acad, Nat.
Sci. Phila.
Very abundant locally in the Gatun beds of Costa Rica, are
small and medium sized Pectunculids belonging to the group of
178 BULLETIN 39 350
G. acuticostata Sowerby of the Santo Dominican Miocene. They
vary somewhat in size, (20 to 32 mm in height), are usually de-
pressed and with an outline suggestive of the recent East Coast
and Upper Miocene G. pectinata Gmelin. Typical acuticostata
does not seem to occur in Costa Rica and I have adopted Brown
and Pilsbry name of canals.
The G. canalis Brown and Pilsbry from the Canal Zone, and
the frilobtcostata Pilsbry and Brown from Colombia, are both bas-
ed on young shells 15 mm or less in height. Both forms can be
duplicated amongst the Costa Rican shells and are but varieties
of the same species.
In its usual and more abundant form, the shell is depressed,
with numerous (about 26), closely spaced rounded or angled ribs.
These ribs are overrun by even, close, concentric threads. They
represent the usual caza/zs or its variety frzlobicostata, and aver-
age in height about 25 mm. More rarely, the shells are larger
and as illustrated by figure 4 and 5 the ribs may become divided
into 2 or 3, strong, radial cords. The following measurements
will show the usual range in size.
Height 22, diameter 21, thickness 5.5
24 23 5-5
22 32 8
29 29 8
Gatun Stage: Gatun, C. Z.
Comadre; Banana River; Coll. 4 East Grape
Point Creek.
Old Man Sam Creek, C. R.
Glycymeris jamaicensis Dall Plate 19, figures 3, 4
Pectunculus pennaceus Guppy, 1866, Quart. Journ. Geol. Soc. London,
vol. 22, p, 293. Not of Lamarck.
Axinea pennacea Gabb, 1873, Trans. Amer, Phil. Soc., vol. 15, p. 255.
Pectunculus decussatus Guppy, 1876, Qart. Journ. Geol, Soc. London,
vol. 32, p. 532. Not of Linné.
351 Costa Rica M1ocENE—OLSSON 179
Glycymeris jamaicensis Dall, 1898, Trans. Wagner Free Inst., Sci.,
vol. 3, pt. 4, p. 608.
Glycymeris jamaicensis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 345,
pl 26; fig! 13)
This large Pectunculid, externally resembles in its angular
posterior side the Glycymerts pennacea with which it was confus-
ed by Guppy and by Gabb, It differs in being somewhat less con-
vex, less strongly sculptured and most importantly in its central
beaks, located about the middle of the ligamental area. It is, on
the other hand, much more closely allied with the recent West
Indian G. undatus Linné (G. lineatus Reeves) .of which it is
doubteless the Miocene ancestral form, the main difference being
in details of its finer sculpture.
The Glycymerts jamaicensis occurs in the Miocene of Jamaica
and Santo Domingo but there never seems to reach the large size
that the species sometimes attains in Costa Rican. A large shell
in our collection from Old Man Sam Creek measures as fol-
lows:
Height 60, lenght 60, diameter of both valves 37 mm.
Gatun Stage: Water Cay.
Collis Red (Clip) Oreck.
Coll. ga, East Grape Point Creek.
Colls. 5, 6, Old Man Sam Creek.
Soust Creek of Upper Hone Creek.
Glycymeris carbasina Brown and Pilsbry Plate 19, figures I, 2, 5
Glycymeris carbasina Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 363, pl. 28, fig. 9.
The Glycymeris carbasina was described by Brown and Pils-
bry from a small individual from Gatun, measuring only 16 mm
in height. It is closely related to the G. jamatcensis Dall and
probably represents but a varietal form. Its main differences is
its more circular and less convex shell, and less number of hinge-
teeth which number about 1o to 12 on the anterior and posterior
sides respectively.
180 BULLETIN 39 352
All our specimens were collected in the lower Gatun of E.
Grape Pt., Cliff Creek in Eastern Costa Rica, the largest shell
measuring as follows:
Height 56, length 55, diameter of both valves 32 mm.
Gatun Stage: Coll. 4, East Grape Point.
Glycymeris castaneus Lamarck Plate 18, figures 13, 14
Fectunculus castaneus Vamarck, 1819, Anim. sans Vert., vol. 6, p
53.
Fectunculus castaneus Reeve, 1843, Conch. Icon., Pectunculus pl. 6,
fig \/32)
Our collection from the Banana River contains several small
Pectunculids which differ from the jamazcensis and its allies by
their less circular shell, greater convexity and coarser radial
sculpture. These shells agree however very closely with a recent
species which is abundant on the beaches of northern Costa
Rica, which appears to be the Glycymeris castaneus of Lamarck.
Glycymerts castaneus is distinct from undutus Linné (G. lineatus
Reeve) with which it is sometimes united, differing by its form
which is more produced posteriorly but not angulated and by its
more pronounced radial costeze. Both species are similiarly col-
ored with chestnut markings.
Gatun Stage: Hill No. 3, Banana River.
Quitana creek.
Glycymeris decussatus Linnzeus Plate 19, figures 6, 7
Arca decussatus Linneus, 1758, Syst. Nat., ed. 10, p. 694.
Fectunculus pennacea Vamarck, 1819, An. s. Vert., vol. 6, p. 51.
Fectunculus pennacea Reeve, 1843, Conch. Icon., /ectunculus, pl.
5s ten 248
Fectunculus decussatus Linné. P. pennaceus Wamarck, (Lamy) 1911,
Journ. de Conchy., vol. 59, p. 119, pl. 3, fig. 7.
Of this rare shell, more commonly known as the G. penna-
cea Lamarck we have several excellent specimens collected by
353 Costa RicA M1ockENE—OLSSON 181
Dr. A. C. Veatch from the Upper Gatun of Old Man Sam Creek
in eastern Costa Rica and a few specimens from the coral-reef
limestones of Port Limon. Although a very distinct species and
not to be confused with any other Pectunculid recent or fossil,
its characters have been generally misunderstood and its name
has often been applied to shells of the G. /éneatus group.
The shell is of moderate size, strongly convex and inequila-
teral. The beaks are slightly posterior of the middle, with the
ligamental area entirely anterior to the beaks as seen in figure 7.
The surface is sculptured with numerous, moderately coarse,
radiating threads which are simple on the umbonal area but be-
come divided by 3 or more finer threads ventrally.
Gatun Stage: Old Man Sam creek.
Port Limon.
Glycymeris Lloydsmithi Pilsbry and Brown Plate 25, figures 8. 9, I0
Glycymeris Lloydsmithi Pilsbry and Brown, 1917, Proc. Acad. Nat.
Sci. Phila., vol. 59, p. 39, pl. 6, fig. 6.
This is an abundant fossil in eastern Costa Rica, where it
frequently forms zones in the lower and middle Gatun. Itstype
of sculpture of broad, smooth, rounded ribs, is strikingly like
that of G. swbovata Say of the Miocene of eastern United States.
It differs in its higher and narrower umbos and by its nearly
smooth cardinal area.
The G.Lloydsmithi was described by Pilsbry and Brown,
from beds equivalent to the Gatun, near Cartagina, Colombia.
Gatun Stage: Zone 3, East Grape Point Creek, C. R.
Coll a Red: \Cligh Greeks (Guipie
Headwater of Middle Creek, C. R.
Genus ARCA Linnzeus
Arca occidentalis Philippi Plate 22, figure I
Arca occidentalis Philippi, 1847, Abbild. u. Beschr., 3, p. 14, pl. 17b,
fig. 4a-c.
182 BULLETIN 39 354
Arca noe Guppy, Journ. Geol Soc. London., vol. 22, p. 293.
Arca occidentalis Sheldon, 1916, Paleont. Amer., vol. 1, p. 8, pl. 1,
figs. 8-11.
Arca occidentalis Maury, 1917, Bull. Amer. Pal. vol. 5, p.327, pl. 29,
Tey
This is a common recent species of the West Indian and
Caribbean fauna. Asa Miocene fossil it is found in Jamaica
and Santo Domingo and it is here recorded from the Miocene of
Costa Rica.
Gatun Stage: Port Limon.
Arca umbonata Lamarck Plate 22, figure 2
Arca umbonata Lamarck, 1819, An. s. Vert., vol. 6, p. 37.
Arca imbricata Gabb, 1873, Trans. Amer. Phila. Soc., vol. 15, p. 254,
In part.
Arca umbonata Sheldon, 1916, Paleeont. Amer., vol. 1, p. 8. pl. 1,
figs. 12-17.
Arca umbonata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 327, pl. 30,
Ag.) TN.
Arca mubonata Maury, 1920, New York Academy of Science, Sci. Sur.
Of PAR vols) 3\ipt.) 1s) pii6:
The Arca umbonata and the preceding occidentalis area com-
mon as recent shells along the entire northern coast of Panama
and Costa Rica. The A. umbonata is recognized by its more
convex and angular shell and finer sculpture of the middle of
the shell disk. Asa Gatun fossil we have but a single small
shell from the Hast Grape Point Creek. Dr. Maury records
the species from the Miocene of Santo Domingo and from
Porto Rico.
Gatun Stage: Coll. 1, East Grape Point Creek.
Subgenus BARBATIA (Gray) Adams
Arca Mauryae, n. sp. Plate 22, figure 4, 7
Barbatia cf. Bonaczyi Maury, 1917, Bull. Amer. Pal., vol. 5, p. 329,
pl. 30, fig. 15. Not of Gabb.
355 Costa RicA MIocENE—OLSSON 183
Shell small, thin, convex, subrhomboidal in form; umbos
wide and full; with the beaks at the anterior 1-3 of the shell; a
rounded posterior umbonal slope; the anterior extremity is brok-
en inthe type specimen but from the earlier growth lines is
simply rounded in form; the posterior side is obliquely truncate
and meets the dorsal margin in a well-marked angle; the exteri-
or is sculptured like that of Arca barbata, with the riblets group-
ed in wide, regular bands which are defined by slightly deeper
interspaces; the radial riblets are crossed and crudely granulat-
ed by concentric threads; the sculpture on the posterior submar-
gins is much heavier and predominantly radial that of the an-
terior submargin is but slightly heavier than on the middle of
the shell disk; interior of shell cavity deep, with a straight hinge-
margin, the teeth arranged asin darbata; cardinal area rather
high, longitudinally striated posterior of the beaks, plain in
front.
Length 24? height 11.50, semidiameter 5.50 mm.
This species is closely related to the recent Avca barbaia
Linné. It differs from daréaza in its proportional shorter shell,
greater convexity, less anterior beak and higher cardinal area.
It is equivalent to the Dominican shell figured by Dr. Maury as
Barbatia cf. Bonaczyt Gabb. Gabb in his remarks following his
his description of Bonaczyi, compares his shell with éarbata, and
states that in Bonaczyz, the beaks are more anterior and the pos-
terior end more produced and sloping. ‘This is the reverse of
the conditions as seen in the present shell. Dall on the other
hand, considers Gabb’s shell as synonymous with Arca umbonta
Lamarck.
Gatun Stage: Port Limon.
Section CALLOARCA Gray
Arca cf. candida Gmelin Plate 22. figures 5, 6
Arca candida Gmelin, 1792, Syst. Nat., p. 3311.
Barbatia (Calloarca) candida Dall, 1898, Trans. Wagner Free Inst.,
SCi., Vols) 34) pt 4). p. 626.
184 BULLETIN 39 356
Arca cadida Shelton, 1916, Palaeont. Amer., No. 1, p. 16, pl. 3, figs.
The 02)
We have several small valves of a Calloarca from the Mio-
cene limestones of Port Limon, the largest of which has a length
of 1 mm. ‘They belong to the Arca candida group and may
possibly represent the young of that species. They seem to dif-
fer in being more finely sculptured, especially about the middle
of the shell disk. ‘This sculpture consists of fine, beaded or
granulated radial threads, which become coarser on the anterior
and posterior submargins.
The Arca candida is recorded by Dall from the Bowden beds
of Jamaica, the equivalent of the coral limestones of Limon. Dall
also mentions from the same locality, the occurrence of a smaller
and possible distinct species. The Arca candida is a common re-
cent species of the West Indies and the Caribbean, and is abund-
ant on the north coast of Panama and Costa Rica.
Length 18, height 11?, semidiameter 3.75 mm.
Gatun Stage: Port Limon.
Subgenus SCAPHARCA Gray
Arca dariensis Brown and Pilsbry Plate 22, figures 10-13
Arca oronlensis Dall, 1898, Trans. Wagner Free Inst. Sci., vol., 3, pt.
4, p. 658. Not of Gabb.
Arca dariensis Brown and Pilsbry, 1911, Proc. Acad, Nat. Sci. Phila.,
vol. 63, p. 362, pl. 22, fig. 10.
Arca gatunensis Toula, Dec. 1911, Jahrb. der K-K Geol. Reichsanstalt,
vol, 61, p. 493, pl. 30, fig. 4. !
The common Ark of the Canal Zone. The shell is elongate
in form with about 30 ribs. The ribs of the left valve are usually
granulated over the whole shell, while those of the right valve
are narrow and smooth on the middle of the shell disk. The ribs
are characteristically divided on the anterior and posterior ends
of the shell.
Gabb has described an Avca oronlensts from the black shales
357 Costa Rica M1ocENE—OLSSON 185
of Oronli Creek in the Talamanca Valley (probably the Uscari
shell). Itis like A. dariensis, in form, but with more central
beaks and umbones and simple ribs. We have not seen this
species.
Gatun Stage: Gatun and Mt. Hope, C. Z.
Woter Cay.
Arca actinophora Dall Plate 23, figs. 7, 8; Plate 25, fig. 3
Scapharca (.Scapharca) acitnophora Dall, 1898, Trans. Wagner Free
Inst. Sci., vol. 3, pt. 4 p. 647, pl. 33, fig. 26.
Scapharca (Scapharca) actinophora Sheldon, 1916, Paleont. Amer.,
VOl tA Ps) 50) pl. Tn, fees:
A larger species than the davzenszs, with broad, wide umbos.
The ribs number 36 to 40, are narrow and separated by inter-
spaces of a little more than their own width. ‘These interspaces
are generally finely ornamented by regular, even, concentric
threads. The ribs are simple and on the left valve, usually strong-
ly beaded or granulated throughout, becoming wider and smooth-
er on the posterior portion, or with the granulations only on
their edges, so that the ribs appear as if mesially depressed. The
ribs of the right valve are less strongly beaded and may become
quite smooth, especially on the center of the shell disk.
Its ventral margin is gently curved to the rounded and not
pointed posterior extremity.
It is a common species at Mt. Hepe in the Canal Zone, but
it also occurs but less abundant in Western Panama and in
Costa Rica.
Catun Stage. Mi. ope, C. Z.
Water Cay, Panama.
Zoned, hea Clif, Greer iGy ie
Coll. 3, Hone Walk Creek, C. R.
Hill t Banana River, C. R.
186 BULLETIN 39 358
Arca Henekeni Maury Plate 24s aman ts
Arca consobrina Sowerby, 1849, Quart. Journ. Geol. Soc. London, vol.
6, p. 52, pl. ro, fig. 12. Not of d’Orbigny, 1844.
Arca consobrina Guppy, 1876, Quart. Journ. Geol. Soc. London, vol.
32, P. 531.
Arca consobrina Dall, 1888, Trans. Wagner Free Inst. Sci., vol. 3, pt.
4, p. 646.
Arca consobrina Sheldon, 1916, Paleont. Amer., No. I, p. 49.
Arca Henekeni Maury, 1917, Bull. Amer. Pal.,vol. 5, p. 331,pl. 29,fig. 2.
Arca (Scapharca) sobrina Pilsbry and Johnson, 1917, Proc, Acad. Nat.
Sci. Phil., vol. 64, p. 186.
Our shells from Water Cay are similiar in form and general
sculpture to the 4. Heneken? Maury (A. consobrina Sowerby),
from.the Miocene of Santo Domingo, the only noticeable differ-
ence being the greater number of ribs on the Panama shell. The
ribs of Hlenekeni from Water Cay average about 39 in number,
and 35 to 36 on the Dominican shells. The ribs of the left
valve are evenly beaded or granulated by concentric threads
which pass across the interspaces as elevated threads. The
ribs of the posterior-dorsal submargins are nearly smooth.
They are more or less divided at the anterior extremity and
very finely beaded.
Length 29, height 18.25, diameter 14.15 mm.
Bile 18, semidiameter of right value
7.25 mm.
Gatun Stage: Water Cay.
Arca costaricensis, n. sp. Plate 25, figures I, 2
Shell moderately large, elongate, convex, with the beaks sit-
uated at the anterior 1-3;ventral margin but gently curved to the
attenuated and pointed posterior extremity; right valve with
about 30 ribs, of which the most anterior and posterior ones are
rather wide, the others narrow; interspaces on the center or the
disk 3 times or more the width of the ribs; the ribs of the right
valve are obscurely granulated on the anterior extremity, more
or less smooth on the center but on the posterior extrem-
ity, they are double and peculiarly granulated along their sides;
359 Costa RrcA MiocENE—OLSSON 187
this granulation begins on the sides and on the more anterior
ribs, the center is smooth; this granulation is in the form of in-
verted V’s; their acute apices lying along the center of the ribs;
the ribs of the left valve are more or less granulated or beaded
throughout and with the ribs on the posterior extremity, double
and similiar to those of the right valve; hinge-line straight,
with very numerous small, uninterrupted teeth; a rather wide
cardinal area with 5 or 6 ligamental grooves; ventral margin in-
ternally fluted in harmony with the external ribs.
Length 52, height 28, diameter 34 mm.
52, Bn 17 mm (right valve).
The form of this shell with its gently curved ventral mar-
gin and attenuated posterior extremity is like the Avca actino-
phora Dall, but is longer and with fewer and heavier ribs. On
the other hand, this species may be compared with darzenszs
Brown and Pilsbry, from which it differs most strikingly by its
larger size, wider, higher and more central umbos and more
pointed attenuated posterior extremity.
Gatun Stage: Hill ra, Banana River.
Fill 2, Banana River.
Zone 7, Pumbri Creek.
Arca honensis, n. sp. Plate 22, figures 8, 9
Shell rectangular, elongote, moderately convex; beaks situ-
ated at about the anterior 1-4; right valve with about 26, nar-
row, smooth or irregularly granulated ribs, separated by inter-
spaces of twice their own width; the left valve with the same
number of ribs and similiar to those of the right valve; the ribs
of both valves are simple and undivided and if granulated, most
heavily on the anterior half of the left valve; interspaces smooth
or with irregular, raised concentric threads; cardinal area long
and rather wide and with 5 or 6 linear grooves; hinge-lines
straight with very numerous, small, uninterrupted teeth; ven-
tral margin fluted in harmony with the external ribs.
188 BULLETIN 39 260
Length 39, height 23.5, semi-diameter 11 mim (lest valve)
36.5 23 12 (left valve)
34 23 ie) (Galea wade)
In general form, quite similiar to the Avca dariensis Brown
and Pilsbry, but distinguished mainly by its simple, undivided
ribs. Itis also amore convex shell, with higher and more
prominent umbos. It differs from Gabb’s figure of A. ovonlen-
sts in its more anterior umbos.
It isa common species, occurring in zones in the Gatun
sandstone of Hone Walk Creek of Eastern Costa Rica.
Gatun Stage: Hone Walk Creek.
Arca golfoyaquensis Maury, var. medioamericana, n. var.
Plate 23, figures 4-6
ef. drca golfoyaquensis Maury,1917, Amer. Pal., vol. 5, p. 332, pl. 28,
fig. 5.
? drca (Andara) consobrina Gabb, 1881, Journ. Acad. Nat. Sci. Phila.,
and series, vol. 8, p. 378.
The Arca golfoyaquensis was described by Dr. Maury from
the Gurabo and Cercado formation of Santo Domingo. It is
closely related to the Arca Henekent Maury, (the Arca consob-
yina) differing mainly in form.
The Costa Rican shells are very similiar to Dr. Maury’s
type specimens of golfoyaquensis, but differ in having 35 instead
of 38 ribs, a higher shell, fuller umbos and sharper ribs on the
posterior-dorsal area. The central ribs of the left valve are ele-
gantly beaded with their interspaces evenly sculptured with
raised concentric threads. On the right valve, the ribs are more
smooth but their interspaces are similiarly sculptured to those of
the left. The ribs of the posterior-dorsal angle of both valves
become divided ventrally by 2 or 3 fine threads. ‘The posterior
extremity is straight, meeting the hinge-line at an angle of
nearly 120 degrees.
Length 54, height 34, diameter 34 mm.
361 Costa RicA MiIocENE—-OLSSON 189
Gatun Stage: Hill 1, Banana River.
Coll ss ied. Glan Greek
Rio Blanco.
Old Man Sam Creek, r mile south of beach.
Arca Veaichi, nu. sp. Plate 23, figures 1-3
Shell large, moderately convex; in outline, the shell is near-
ly square with the height about equal to its length; the anterior
side is widely rounded to the ventral margin, the posterior side
_ nearly straight, nearly at right angles to the hinge-line; umbos
wide, with the unbonal angle widely rounded and with the ex-
treme tip of the beaks situated at the anterior 1-3 of the total
length of the shell; right valve with about 45 ribs, of which the
posterior 14 are on the posterior-dorsal slope; the ribs are rec-
tangular in section with the interspaces on the center of the
disk about 3-4 of the width of the ribs; the ribs are finely scul-
ptured but less strongly on the right valve; on the center of the
shell disk, the ribs are finely nodulated or granulated but on the
posterior-dorsal slope and on the anterior extremity, the granu-
lations of the ribs is gradually overshadowed by the introduc-
tion of 4 or 5 fine, threads on the top of each rib the interspaces
on the center of the disk of the left valve are finely sculptured
with even, raised concentric threads; the left valve is slightly
larger and overlaps to a small extent the right; hinge-line
straight with numerous small teeth, larger and higher at the ex-
tremities; cardinal area that of typical Scapharca, and grooved
with 3 lines; interior of the shell deep, with the posterior ad-
ductor scar about twice the size of the anterior; basal margin
fluted in harmony with the external sculpture.
Length 51, height 51, diameter 41.5 mm.
This isa rather large, Cardiwm-like Arca of nearly square
outlines and numerous finely sculptured ribs. Its form is also
suggestive of Avgzna, but its cardinal area is that of a true
Scapharca. Thetype specimen, a finely preserved individual
1gO BULLETIN 39 362
with both valves comes from the Gatun of Water Cay. It is
quite unlike any other known American species.
It is named for Dr. A. C. Veatch, Director of the Explora-
tion Department of the Sinclair Consolidated Oil Corporation.
Gatun Stage: Water Cay.
Mit TLopen Guz
Arca auriculata Lamarck Plate 22, figure 3
Arca auriculata Lamarck, 1819, n. s. Vert., vol. 6, p. 43.
Arca (Scapharca) auriculata Dall, 1898, Trans. Wagner Free Inst.,
Sci., vol. 3, pt. 4, p. 649.
Arca auriculata, Sheldon, 1916, Paleont, Amer., vol. 1, p. 50, pl. I1.
fig. 19,
Scapharca auriculata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 339, pl.
28.) fi) 3)
A common, recent species of the West Indies. Its hinge-
line is generally produced or auriculated at its posterior extrem-
ity.
It occurs in the Miocene of Jamaica and Santo Domingo.
Dall also records it from near Limon.
The Costa Rican specimens in our collection are all small
but otherwise typical.
Gatun Stage: Coll. 3, Hone Walk Creek.
Old Man Sam Creek, 1 mile south of the beach,
Section CUNEARCA Dall
Arca cacica, n. sp. Plate 24, figure 1
Shell of moderate size, strongly convex and with high, in-
flated umbos; beaks and umbos about the middle of the shell;
the greatest convexity of the shell lying about the anterior 1-3,
and with a narrow but deep, radial depressed zone extending
from the beaks to the ventral margin, just anterior to the um-
bonal angle; the anterior end is slightly rounded, the posterior
363 Costa Rica MiocENE—OLSSON 19]
submargins strongly depressed or flattened with a straight post-
erior margin; the left valve with 30 smooth and strongly nodul-
ous radial ribs of which 9 are situated on the posterior sub-
margin; the ribs of the posterior submargin are low, wide and
nearly smooth; on the rest of the shell, the ribs are nodulose,
very regularly in the middle of the valves more distantly on the
anterior ribs; the interspaces between the ribs are narrow on the
middle of the valves, become wider anteriorly and on the ex-
treme anterior extremity equal the ribs in width; the cardinal
area is high, as nearly % of the length.
Length 39, height 33, diameter of the left valve 15 mm.
The above description is based on a single left valve from
the Upper Gatun beds near Cahuita. Distantly related to the
Arca tncongrua Say and its southern form 6vaszliana Lamarck,
the fossil species differs in its higher and more convex shell, and
in its high cardinal area. In this latter feature, this species is
like the recent West Coast Arca labiata Sowerby, but has a
greater number of ribs and different form.
Gatun Stage: Across the divide trom Comadre Creek.
Groups of Arca Pittiert Dall
This is a group of small and medium-sized Arks, containing
3 species, whose general appearance is that of the recent Avca
Chemnitzt Phil. The Costa Rican shells are usually abundant
wherever they occur in the Gatun formation. Their relations
and differences may be summarized as follows:
A. Both valves more or less similiarly sculptured, that is
the ribs of the posterior half of the right valve, at least obsolete-
ly beaded.
B. Ribs 25 to 28; umbos high and full; posterior-dor-
sal slope, angular; posterior extremity somewhat produced,
giving an elongate outline to the shell.
Area Pittiert Dall
AA. Valves not similiarly sculptured; posterior half of
192 BULLETIN 39 264
the right valve with narrow ribs, separated by wide inter-
spaces.
B. The intervals between the ribs on the posterior
half of the right valve with small (sometimes faint) interstitial
threads. Ribs about 29.
Arca Lloydi, n. sp.
BB. The intervals between the ribs on the posterior
half of the right valve smooth; ribs 25, shell usually
larger.
Arca Flindsiz, n. sp.
Arca Piitieri Dall Plate 24, figures 2-6
Arca (Scapharca) Pittieri Dall 1912,Smith. Misc. Coll., vol. 59, No. 2,
P. 9.
This isa common species in the Gatun beds of the Banana
River. The ribs of the 2 valvesare similarly sculptured but
somewhat less strongly on the right. The umbos are high.
The posterior extremity is pointed and produced so that the out-
line of the shell is more elongate than in the following species:
Length 25, height 25, diameter 26 mm.
20 28 semi-diameter 14.5 mm.
Gatun Stage: FHiill ra, Banana River, 3.
Zone Ei, Saury Creek.
Rio Betey.
Goll. 7, Pumbre Creek.
Arca Llioydi, n. sp, Plate 24, figures 10-12
Shell small, cordiform, covex, subrectangular in outline but
with the left valve somewhat more pointed at its posterior-ven-
tral extremity; anterior side well rounded; posterior side straight
and meeting the basal margin at an angle of about 75 degrees;
umbonal angle rounded; the left valve has 29 ribs of which g
are found on the posterior- dorsal slope; the ribs of the left
valve are strongly beaded or granulated, and separated by inter-
spaces of about their own width; on the right valve, the first 10
anterior ribs are strongly beaded; the next set as far as the um-
365 Costa Rica MrocENE—OLSSON 193
bonal angle are smooth, beyond which on the posterior-dorsal
slope the ribs again become beaded as they approach the dorsal
margin; the interspaces on the right valve are wide, and on the
shell disk anterior to the umbonal angle they carry a fine, but
often faint interstitial thread; the intervals are otherwise smooth
or finely etched with concentric lines; hinge-lines straight with
numerous small, vertical teeth and a wide, smooth cardinal
area.
Length 18, height 18, semi-diameter 9.5 mm.
A smaller species than the following (//zzdsz) and easily
distinguished by the interstitial thread of its right valve. It is
much less common than the other two species.
Named for Dr. E. R. Lloyd, formerly of the United States
Geological Survey, and who was a member of the first Hxplora-
tion Party of the Sinclair Oil Company in Panama and Costa
Rica.
Gatun Stage: Hill 3, Banana River.
Zone 5, ked Cliff Creek.
Coll. 7, Pumbri Creek.
Arca Hindsi, n. sp. Plate 24, figures 7-9
Shell small or of medium size, cordiform and similiar in out-
line to the preceding species but larger; umbos high and wide,
and but slightly anterior to the middle; the left valve has about
26 strongly nodulated ribs, separated by interspaces of slightly
greater width; the first 9 or 10 ribs ef the right valve are nodu-
lated, followed by smooth ones to the umbonal angle, beyond
which the remaining 7 become nodulated as they approach the
dorsal margin; interspaces wider than the ribs, smooth and
without any interstitial thread; hinge-line straight with numer-
ous, small vertical teeth and a wide, smooth cardinal area; on
some shells the ligamental area may carry one or two, small
diamond-shaped grooves.
Length 24, height 25, semi-diameter 14 mim.
A much larger species than the Z/oyd7 and without any in-
194 BULLETIN 3G 266
terstitial thread in the interspaces of its right valve. It is nam-
ed ror Dr. Henry Hinds, formerly of the United States Geologi-
cal Survey and the Exploration Department of the Sinclair Con-
solidated Oil Corporation.
Gatun Stage: Hill No. 3, Banana River.
Zone f, Saury Creek.
Coll, 2, East Grape Point Creek.
Coll. 3, East Grape Point Creek.
Rio Betey.
Subgenus NOETIA, Gray
Arca MacDonaidi Dall Plate 25, figures 4-7
Area (Noetia) MacDonaldi Dall, 1912, Smith, Misc. Coll., vol. 59, No.
PISO (0)
This large Voétia, is very common and characteristic of the
Gatun formation in many parts of Northern Costa Rica, and
large specimens may reach a length and height of 55 mm or
more. It was originally described by Dall from specimens col-
lected by Pittier and MacDonald from the Gatun beds of the
Banana River, where it is associated with Avca Pittieri and is
very abundant.
Two forms or varieties may be recognized. In typical J7Zac-
Donaldi, the shell is high, trigonal, with elevated, narrow umbos
and a producted anterior extremity. ‘The beaks are separated from
the hinge-line by a wide ligamental area. ‘This is the conimon
form along the Banana River. Frequently associated with typi-
cal MacDonaldi are shells'with a more rectangular outline, lower
but wzde umbos. ‘The beaks are close to the hinge line so that
the ligmental area ts narrow. These two types have a very dif-
ferent appearance but seem to be connected by transitional forms.
To the last variety, I propose the name of subreversa.
Among recent shells, the 47ca MacDonald? finds its closest
relation with the common Avca reversa Sowerby of the Pacific
coast of Central America.
367 Costa RicA MrockNE—OLSSON 195
Gatun Stage: Hill 1, ra, Banana River, typical and var.
suberversa
Coll. 7, Pumbric Creck (typical)
Coll. 6, Red Cliff Creek (typical)
Betey (subreversa)
Zone G, Saury (subreversa)
Genus PTERIA, Scopoli
Pteria inornata Gabb Plate 28, figure 9
Avicula inornata Gabb, 1873, Trans. Amer. Phil. Soc., vol., 15, p.
253.
Pteria inornata Dall, 1898, Trans. Wagner Free Inst. Sci., vol. 3, pt.
4, p. 669.
Pteria tnornata Maury, 1817, Bull. Amer. Pal., vol. 5, p. 345, pl. 26,
fig. 14.
This species is locally common in Costa Rica, but generally
difheult to collect because of its thin and fragile shell. Its
valves are as high as long, strongly convex and smooth. The
anterior ear is of medium size, the posterior much larger and
produced.
It was described as a Miocene fossil from Santo Domingo,
but has also been collected at Bowden.
Uscari formation. Old Harbor.
Gatun Stage: Zone 3; Red Cliff Creek.
Genus OSTREA, Linnzus
Ostrea megodon Hanley Plate 18, figure 1
Ostrea megodon Hanley, 1845, Proc. Zool. Soc. London, p. 106.
cf. Ostrea cerrosensis Gabb, 1869, Geol. Surv. Calif., vol. 2, p. 35, pl.
II, fig. 61.
Ostrea megodon Dall, 1897, Trans. Wagner Free Inst., vol. 3, pt. 4, p.
685.
Ostrea megodon Maury, 1917, Bull. Amer. Pal., vol. 5, p. 347, pl. 34,
LGR Be
196 BULLETIN 39 363
This is a medium sized and sickle-shaped oyster with 4 or 5,
large, tooth-like folds on the anterior and ventral margins. The
beak and ligamental area are directed upwards and sometimes
strongly to the posterior side. Its external surface, although
strongly and deeply folded, is plain and nearly smooth.
The Ostrea megodon isa recent species along the west or
Pacific coast of Central America. It occurs also as a Miocene
fossil in Santo Domingo and in the Bowden beds of Jamaica.
Gatun Stage: Water Cay.
Coll. ga, East Grape Point Creek.
fiill ra, Banana River.
Ostrea costaricensis, n. sp. Plate 21, figure 2
Ostrea sculpturata Bose, 1906, Boll. de Inst. Geol. de Mexico, numero
22, p. 76, pl. 9, ig. 4. Not of Conrad, 184o.
Shell of medium size, subcircular in form but somewhat ob-
lique; the shell is somewhat flexed or slightly saddle-shaped, the
concave side of which corresponds to the left valve; both valves
are strongly sculptured, the right with narrow, rib-like costz
and wider interspaces, the left valve with the reverse of the
sculpture of the right; the ribs are more or less nodose or knob-
bed and are crossed by irregular concentric lines, which in some
cases may become raised or lamellar-like; the hinge margin is ir-
regularly crenulated:
Height 50, length 47, diameter I9 mm.
The type is aspecimen from the Gatun of the Rio Blanco
with both valve tightly closed so that the interior cannot be
seen. What appears to be the same species, is represented by
several sessile and free valves from East Grape Point Creek, and
which are associated with O. megodon.
It differs from the Ostrea gatunensts - haitensis group by its
smaller size; less heavy shell and in that the ribs of the valves
are narrow and coste-like and not long, strong plicee. Bose
has recorded and figured this species as Ostrea sculpturata Con-
369 Costa Rica MIocENE—-OLSSON 197
rad from the Miocene (as Pliocene) of the State of Oaxaca,
Mexico. Itis very distinct from the East Coast, Chesapeake
Miocene Os/rvea sculpturata both by its form and sculpture.
Gatun Stage: Rio Blanco.
Coll. ga, East Grape Point Creek.
The collection also contains fragments or much decayed
specimens of two or more large oysters, one of which is probably
the gatunensis Brown and Pilsbry of the Canal Zone, and a
large, heavy selleformis-like species, possibly the Galbertharrist
Maury of the Dominican Miocene. The specimens are however
too fragmentary for positive determination.
Genus PECTEN, Muller
Pecten gatunensis Toula Plate 16, figures 3, 4
Pecten (Flabellipectcn) gatunensis Toula, 1908, Jahrb. der K-K Geol.
Reichsanstalt Wien, vol. 58, p. 711, pl. 29, fig. 2.
Pecten gatunensis Brown and Pilsbry, 1911, Proe. Acad, Nat. Sci.
Phila., vol. 63, p. 365.
Shell rather large, subcircular; the left valve is nearly flat
or slightly vaulted transversely across the middle of the shell;
the right valve is moderately convex; the ears are small and
nearly equal,those of the right valve slightly flaring at the sides;
the right valve has 23 or 24 fairly strong ribs separated by inter-
spaces a little more than % of their width; in large right valves,
some of the central ribs become dichotomously or trichotomously
divided but this feature is not always developed to the same ex-
tent on all shells; the left valve has 18 or 19 ribs which fade out
on the wide dorsal submargins; the ribs of the left valve are
narrow with the interspaces nearly twice as wide; the surface is
covered with fine, even, raised concentric threads best preserved
on the left valves; interior fluted.
Length 62, height 60? diameter 12.5 mm.
62 60? 4 mm (left valve)
198 BULLETIN 39 37°
We have redescribed this species as Toula had but a speci-
men of the right valve. Itisa true Pecten, distantly related to
the Pecten hemicyclicus Ravenel from the Chesapeake Miocene of
Florida and the Carolinas, but is smaller and less broad. In both
species, the ribs on the right valves of large specimens become
divided in the same manner.
Gain Stage Gatun GC. Z:
Water Cay,
Pecten MacDonaldi, n. sp. Plate 16, figures 1, 2
Shell large, subcircular; ears of medium size and equal; left
valve is slightly convex due tothe middle of the shell being
transversely humped or vaulted and a depressed zone follows on
the inner slope of each of the raised dorsal submargins; the
right valve is slightly but evenly convex; sculpture of the right
valve consists of about 26, low ribs which widen out as they
approach the ventral margins; their interspaces are at first nearly
as wide as the ribs themselves but become only % or 4% as wide
ventrally; the left valve has 21 or 22 narrow ribs and wider inter-
spaces; on the dorsal submargins, the ribs are small and fade
away; surface with fine, even, raised lines best seen on the left
valve.
Length 106, height 97, semi-diameter 12 mm.
83 80 mim.
The fragmentary Toro limestone at Gatunand Toro Point
contains very few good fossils,although the rock itself is compos-
ed almost entirely of broken and ground fragments of shells and
barnacles. Dall has described an Afztonzum toroensis from the
Toro Point limestone where it is fairly common. ‘The Pecten
MacDonald is found in the Toro limestone which caps the hills
just west of the locks at Gatun.
This species has much the same contour as the recent Pecten
maximus Linné, from Europe. It isa larger species than the
Pecten gatunensts Toula.
Toro Limestone. Gatun, west of the locks.
Sur Costa Rrcsa MrockNE—OLSSON 199
Pecten coralliphila, n. sp, Plate 17; figures 5, 6
Shell small, very thin and generally translucent; broader
than high; right valve moderately convex, the left concave with
raised submargins; ears small and subequal; sculpture of the
right valve of about 23 low, flat ribs, with flat interspaces a
little more than ¥%4 the width of the ribs; some of the ribs may
become divided medially; the ribs of the left valve, about 20 in
number, are narrow, usually somewhat irregularly spaced so
that their interspaces are of variable width and new ribs may
occasionally appear inthe wider interspaces near the ventral
margin; surface in addition finely sculptured with even, raised
concentric threads; interior of the shell fluted well into the cav-
ity of the shell, in harmony with the external ribs.
Length 34, height 29, semi-diameter 4 mm (right valve)
This Pecten is fairly common in the coralline phases of the
Miocene and its associated marls and sands. It is extremely thin
and fragile in texture. Its sculpture resembles that of /Pecten
gatunensts Toula but is much more irregular. The interspaces
of the left valve are of very variable width.
Gatun Stage: Port Limon.
Subgenus CHLAMYS, Bolton
Section PLAGIOCTENIUM Dall
Pecten levicostatus Toula Plate 16, figures 5-7
Fecten tevicostatus Toula, 1908, Jahrb. der K-K Geol. Reichsanstalt,
Wien, vol. 58, p. 713, pl. 26, figs. 4, 5, 6.
FPecten (Chlamys) santarosanus Bose, 1906, Boll. de Inst. Geol. de
Mexico, numero 22, p. 73, pl. 6, figs. 3,4, ? FPecten (Chlamys)
santarosanus Bose, op. cit. p. 23, pl. 1, figs. 1 and 4, 2 and 5.
A scallop of the general type of Pecten ventricosus Sowerby;
both valves equally and strongly convex; ribs about 21, which
are square or rectangular in section on the umbos and with deep
interspaces; the ribs generally flatten or become more rounded in
206 BULLETIN 39 372
large shell near the ventral margins; the interspaces are covered
with fine, raised, concentric lines, best seen on the umbos or on
young shells; they become more crowded towards the ventral
margins and may pass over the tops of the ribs as well; the dor-
sal and posterior submargins are strongly flattened and sculp-
tureless; the anterior ear is a little larger, projects forward and
carries, in the right valve, a byssal notch; it is sculptured with
several radial riblets; the posterior ear has its upper half nearly
smooth or but faintly sculptured, but its lower half carries a
band of 5 or 6 strong, radial cords.
Height 29, breadth 34, thickness 10.50 mm (right valve)
34 38 23 (both valves)
39 43 T4 (left valve)
This is the common /Pecten in Costa Rica and frequently
forms zones in the lower part of the Gatun formation. All our
specimens have 21 ribs, which vary from strongly rectangular
in section with deep interspaces to more flattened and rounded
on the larger individuals.
Bose has described as Fecten santarosanus, a small scallop
shell from the Miocene of the State of Oaxaca, Mexico. The
ribs number 23 to 25. Later in the same volume, he figured a
larger Pecten from the Miocene of La Barranca de Santa Marie
Tatella, as Pecten santarosanus. ‘This shell has but 21 ribs and
is equivalent to Toula’s Jevicostata. If this larger specimen of
Bose’s is equivalent to his typical saztarosanus, this name must
of course take priority over Toula’s levicostatus. As Bose’s typi-
cal santarosanus has 23 to 25 ribs, whilein /evicostatus the ribs
appear to beconstantly but 21 in number, they probably repre-
sent two different species.
In the Dominican Miocene J/evzcostafius is represented by the
Pecten excentricus Gabb, which appears to differ constantly by its
lower and more rounded ribs and smaller ears. The recent
Pacific Pecten ventricosus Sowerby is closely related and probably
the direct descendent of /evicostatus. In that species, the ribs
are low and rounded as in excenfricus and as occasionally seen in
373 Costa Rica MiocENE—OLSSON 201
large examples of /evicostatus. ‘The sculpture of the posterior
ear is uniform. Inthe Chasapeake Miocene fauna the /ecten
eboreus Conrad, variety comparilis Tuomey and Holmes is dis-
tantly related, but is larger, less convex and generally with
more numerous ribs.
Gatun Stage: Gatun, C. Z. Quitana Creek.
Water Cay.
Flone Creek.
Banana River.
ftstrella River.
Pecten costaricensis, n. sp. Plate 18, figures 8-11
Shell small, inequivalve, oblique; the right valve is moder-
ately convex, the left but slightly; right valve with 17 flat-top-
ped ribs, square in section and with deep U-shaped interspaces
of a width equal to the ribs; these interspaces are sculptured
with fine, concentric, raised lines which are quite far apart on
the umbos, but become crowded ventrally or disappear entirely;
the left valve has 17 or 18 ribs which are lower and more round-
ed in section; the fine concentric lines on the left valve are con-
fined to the umbos, the interspaces over the greater part of the
shell being smooth; the anterior and posterior dorsal slopes are
flattened and smooth, but sculptured below in each valve by
strong radial threads on the lower side of the ears; ears subequal;
the right valve with a deep, byssal notch in its anterior ear; the
left valve has the lower half of each ear with 3 strong radial
threads, while on the upper half, the radial threads are largely
obsolete; valves internally grooved in harmony with the exter-
nal ribbing to about the center of the shell.
Height 26.50, breadth 28, thickness 7.50 mm (right valve)
24 24 4.00 mm (left valve)
2g 28 5.50 mm (left valve)
Differs from the Pecten levicostatus Toula, by its unequal
valves, less convexity and fewer ribs. It is somewhat like the
Pecten ineqgualis Sowerby, from which it is distinguished by its
202 BULLETIN 39 374
more flattened left valve and lower, smaller ribs of its right. In
inequalis, the fine, lamellose, concentric lines cover the entire
interspaces between the ribs on both valves, while in the present
species the interspaces of the left valve are largely smooth with
the concentric lines confined to the extreme umbonal area.
Gatun Stage: Coll. 4, Red Cliff Creek.
ZI mile south of the beach, along Old Man Sam
Creek.
Coll. 4, East Grape Point Creck.
Rio Blanco.
Section AEQUIPECTEN, Fischer
Pecten (Aequipecten) preglyptus, n. sp. Plate 17, figures 2, 7
Shell of moderate size, subcircular in form and convex; ears
of usual size, subequal and with a straight hinge-line; sculpture
of about 16 or 17 wide, little elevated ribs and narrower, scarce-
ly distinguishable interspaces; the surface is otherwise smooth
or with fine, concentric lines; interior of the shell with 29 or 30
Amustum-like internal liree which are evenly spaced and not in
pairs; these liree are very faint in the middle of the shell cavity
but strong distally.
Length 50, height 49, semi-diameter 6.5 mm.
An Amustum-like species found in the lower sandstones and
conglomerates of the Gatun formation in upper Cocles Creek.
It is closely related to the recent deep-water Pecten glyptus Ver-
rill from the cost of Hatteras and Marthas Vineyard (see Dall
Brocy US Nat-iMiusiivol Mas pe 2asiuple) Ss. . fgs) 22) bot
species having about the same number of low, wide ribs and in-
ternal liree. ‘The fossil shell seem to have been nearly circular
_ while glypfus is decidely oblique in outline.
Gatun Stage: Upper Cocles Creek.
375 Costa R1cA MIOCENE
OLSSON 203
Subgenus PSEUDAMUSIUM, H. and A. Adams
Section PSEUDAMUSIUM, s. s.
Pecten almirantensis, n. sp. Plate 18, figure 16
Shell small, thin, compressed or but slightly convex; the
posterior and anterior dorsal slopes are straight, meeting the
beaks at an acute angle; the right valve is finely sculptured with
close, regular, concentric lamella and wider interspaces; the
posterior and anterior sides show a few, irregular, radial threads
which cross the concentric lamellze; the ears are unfortunately
broken; the interior of the shell cavity is filled with a thin de-
posit of callus; ventral margin plain; hinge line minutely groov-
ed with vertical lines.
Length 4.75, height 4.75, diameter of right valve 60 mm.
This small, elegant species is represented solely by an im-
perfect right valve and represent the first true Pseudamusium to
be discovered in the Miocene beds of Panama. Its sculpure is
like the recent deep water Pseudamustum strigillatum Dall from
the West Indies, but differs in having its anterior and posterior
dorsal submargins meeting at the beaks at a much more acute
angle.
Gatun Stage: Bocas del Toro.
Section CYCLOPECTEN, Verrill
Pecien oligolepis Brown and Pilsbry
Fecten aff. subhyalinus Smith, Toula, 1911, Jahrb. der K-K Geol.
Reichsanstalt, Wien, vol. 61, p. 492, pl. 31, figs. Ia, b, c.
Fecten (Cyclopecten) oligolepis Brown and Pilsbry, 1912, Proc. Acad.
Nat. Sci. Phila., vol. 64, p. 512, text-figure 5.
A very small species with broad, nearly smooth valves and
large ears. The right anterior ear has a deep byssal notch and
its surface is sculptured with fine, raised, concentric, lamellose
lines. The ears of the left valve are similiarly sculptured but fin-
204 BULLETIN 39 376
er. The surface of the valves appear nearly smooth but in well-
preserved specimens very faint concentric lines and fine radial
strisee may be seen. The left valve is more coarsely sculptured.
The hinge is finely grooved with vertical lines. The measure-
ments given by Pilsbry for his Gatun specimen, length 2.8,
height 2.7, diameter of right valve .8 mm, are the usual dimen-
sions for the Costa Rican specimens.
Gatun Stage: Gatun, C. Z.
Coll. 4, East Grape Point Creek.
Middle Creek.
Pecten aotus, n. sp. Plate 18, figures 17, 18
Shell small, thin, depressed or but slightly convex and
Amusium-like; both valves except on magnification appearing
smooth; ears of the left valve large and subequal; the anterior
ear of the right valve about twice that of the posterior and car-
rying a deep, byssal notch; the surface of the left valve on mag-
nification is seen to be sculptured with fine, radial lines which
often occur in irregular streaks or blotches, imparting a peculiar
appearance to the shell; the right valve is perfectly smooth and
featureless, except for a band of rough scales on its anterior ear
above the byssal notch; interior of shell cavity shallow, a simple
ventral margin and the hinge-line vertically grooved with fine
lines.
Length 4.5, height 4.5, diameter .75 mm. left valve.
Differs from Pecten oligolepis Brown and Pilsbry in its larg-
er size, thinner and more Amuszum-like shell, and strongly sculp-
tured left valve. This sculpture consists of fine lines or camp-
tonectes striations, which may be uniformly distributed but more
usually occur in irregular streaks or blotches, imparting a dis-
tinctive appearance to the surface. This sculpture is seen only
on a slight magnification, the shell otherwise appearing smooth
and featureless. It is quite common in the coralline limestones
of Port Limon.
377 Costa RicA MrocENE—OLSSON 205
Gatun Stage: Port Limon.
Coll. 5, Red Cliff Creek
Genus AMUSIUM, Bolton
Amusium luna Brown and Pilsbry Plate 17, figure I
Amusium Mortont Bose, 1906, Boll. de Inst. Geol. de Mexico, Numero
22, p. 24, pl. 1, figs. 3,6, 7,9. Not Amusium Mortoni Ravenal,
1844,
Amusium Mortoni Bose, op. cit. p. 74, pl. 8, figs. 1, 2; pl. 3, fig. 3.
Pecten (Amusium) cf. Mortoni Toula, 1908, Jahrb. der K-K Geol.
Reichsanstalt, Wien, vol. 58, p. 714, pl. 26, figs. 8, 9.
Pecten (Amustum) luna Brown and Pilsbry, 1912, Proc. Acad. Nat.
Sci. Phila., vol. 64, p. 514, pl. 23. fig. 1.
This is the common Gatun Amusium in Panama and Costa
Rica. Its valves are nearly equal and but slightly convex; ears
equal and small, and defined from the rest of the shell by a
sharp ledge; the umbos, as well as the rest of the shell disk are
smooth, except for the faint lines of growth; the internal ribs
are in pairs, numbering about 23 and spaced at intervals nearly
twice the width of the pairs of ribs themselves.
Bose and Toula both referred this species to the Upper
Chesapeake Miocene and recent Amusium Mortont Ravenel,
from which it is very distinct. Amuszum Mortoni is not only
much larger (height 170 mm), but is porportionately much
broader, and with larger ears which are defined simply by a line
from the rest of the shell.
The Dominican A. papyraceum Gabb has larger ears, the in-
ternal ribs are more crowded and the left valve is generally
strongly flexed along the anterior and posterior submargins.
Amustum Toxule Brown and Pilsbry, is found in the Gatun of
the Canal Zone. Its surface is marked with dark-colored rays
and is said to contain no internal ribs.
206 BULLETIN 39 378
Gatun Stage: Gatun, C. Z.
Flotel Creek.
Zone 6, Old Man Sam Creek.
Coll. 4, East Grape Point Creek.
Amusium bocasensis, n. sp. Plate 17, figures 3, 4
Shell rather small, thin and subcircular in outline; moderate-
ly convex; ears proportionately larger than in 4. /una, and with
their dorsal margins slightly inclined upwards from the beaks to
their outer edges; the surface is smooth and polished, finely
covered with concentric growth-lines and faintly showing the
position of the internal lire; interior of shell with about 34
lire, which are usually nearly evenly spaced and not in pairs;
the internal lire commence well in the interior of the shell, but
are strongest distally.
Height 4o, breadth 43.
25 38, thickness 6 mm. (right valve)
A much smaller and more delicate species than the Amuszum
luna Brown and Pilsbry. The internal liree are more or less
equally spaced and not in pairs. It differs from Amuszum Lyonit
_Gabb from the lower Miocene of Sapota, Costa Rica by its much
smaller ears.
Gatun Stage: Bocas del Toro.
Genus SPONDYLUS, Linnzeus
Spondylus chiriquiensis, n. sp. Plate 20, figures 1, 2, 5, 6
Shell large, ponderous, equivalve and strongly convex; both
valves of the same convexity, but with the beak of the right,
high and projecting above the strongly incurved beak of the left;
in large shells both valves are provided with a cardinal area,
that of the right being both high and wide and for the greater
part straight but the beak curving over it toa small extent at
its upper end; the cardinal area of the left vale is much smaller,
379 Costa RicA MIocENE—OLSSON 207
narrow and appears only with maturity and is more or less hid-
den by the strongly incurved beak; the sculpture consists of
strong radial cords and small ribs divisible into 3 sets; a primary
set of 7, which in perfect specimens and in the young carry large,
strong spines; a secondary thread in each of the wide interspaces
and when perfect bearing smaller spines, and tertiary threads of
a variable number between the secondaries and primaries; the
whole surface is in addition, finely, longitudinally starited with
minute, scale-bearing threads; young shells have a small foliace-
ous area on the umbo of the sessile or right valve and strong
spines on both valves.
Height 151, length 120, diameter 115 mm.
A large, ponderous species with nearly equal and strongly
convex valves which show no distortion due to attachment |
Young shells associated with the type have a small, somewhat
foliaceous attachment area on the right valve, and each valve
bears large, strong spines on its primary set of radial cords and
finer spines on its secondary and even on some of the tertiary
threads.
Of described species, the chzrigutensis seems nearest related
to the dostrychttes Guppy from the Miocene of Santo Domingo
and Jamaica but differs by its much larger size, proportionately
more convex whorls and by its cardinal area which is more up-
right and with a more incurved beak at itsupper end. The
sculpture of chtviqutenszs is more spiny,the spines being borne by
the primary radials and to a less degree by the secondaries, and
even by the tertiaries. In dostrychites only the primaries bear
spines, the others appearing as large, nearly smooth, intermedi-
ate threads and cords. We have specimens ofa large, closely
allied Spondylus from the Chesapeake Miocene of Florida, dif-
fering mainly in its cardinal area and in details of sculpture.
Gatun Stage: Water Cay.
Spondylus gumanomocon Brown and Pilsbry Plate 21 figure I
Spondylus americanus Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15,
p. 257. Not of Lamarck.
208 BULLETIN 39 380
Spondylns gumanomocon Brown and Pilsbry, 1912, Proc. Acad. Nat.
Sci. Phila., vol. 14, p. 514, footnote.
Spondylus gumanomocon Maury, 1917, Bull. Amer. Pal., vol. 5, p.
355-
This is a species with very unequal valves, that of the left
or upper, being small, pectiniform and lacking ina cardinal
area, while the right or lower valve is strongly convex, with a
high produced beak and a high cardinal area. The sculpture of
the two valves is similiar, except that the umbo of the right is
strongly foliaceous. It occurs in the Miocene of Santo Domin-
go, where it was referred to the recent S. americanus by Gabb.
Gatun Stage: Coll. 2, Hone Walk Creek.
Port Limon.
Genus PLICATULA, Lamarck
Plicatula marginata Say Plate 28, figures 6, 7
Flicatula marginata Say, 1824, Journ. Acad. Nat. Sci. Phila., vol. 4,
pp. 136-137, pl. 9, fig. 4.
Plicatula marginata Dall, 1898, Trans. Wagner Free Inst. Sci., vol. 3,
pt. 4, p. 764.
The Panama and Costa Rican examples are not typical but
they approach more closely the Plicatula marginata Say than
they do the densata which Dall has recorded from the Bowden
beds of Jamica.
The shells vary from subcircular to elongate and in some
cases carry the dark, marginal band so frequently seen in typical
marginata, ‘The ribs number five to seven; the shells with more
rounded form and more numerous ribs suggest the dexsata, but
the ribs are higher and more foliaceous, and can be exactly du-
plicated by scores of true marginata in the Cornell collection.
The above determination must however be considered as purely
provisional at the present time.
The typical P. densata was described by Conrad from the
lower Chesapeake Miocene of New Jersey, but it also occurs in
381 Costa Rica Mi0cENE
OLSSON 209
the synchronous Calvert formation of Maryland. It is also found
as a lower Miocene fossil in Florida and Dall recorded it from
the Bowden beds of Jamaica and from the Oligocene of Gual-
lava, Costa Rica. The ?. marginata Conrad is an Upper Chea-
peake Miocene and Pliocene fossil of the eastern United States.
Gatun Stage: Water Cay,
Old Man SamCreek, 1 mile south of the beach.
Genus LIMAEA, Born
Limaea solida Dall Plate 28, figure 10
Limea solida Dall, 1898, Trans, Wagner Free Inst. Sci., vol. 3, pt. 4,
p. 769, pl. 35, figs. 4, 5.
A small, solid shell, externally sculptured like a small Gly-
cymeris. It has 12 rounded ribs and a coarsely crenulated basal
margin. Our single specimen from Costa Rica isa little smaller
than Dall’s specimens from Bowden.
Gatun Stage: Coll. 4, Red Cliff Creek.
Genus ANOMIA, (Linné) Muller
Anomia simplex d’Orbigny Plate 21, figure 6
Anomia ephippium Conrad, 1845, Fossils of the Medial Tertiary, p.
75, pl. 43, fig. 4. Not of Linné.
Anomia simplex d’Orbigny, 1845, Hist. Pol. y Nat. Isla de Cuba, p.
367.
Anomia ephippium Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
257:
Anomia simplex Dall, 1898, Trans. Wagner Free Inst., Sci., vol. 3,
pt. 4, p. 784.
Anomia simplex Bose, 1906, Boll. de Inst. Geol. de Mexico, numero
22, p. 25, pl. 2, figs. 18-23.
Anomia simplex Maury, 1917, Bull. Amer. Pal., vol. 5, p. 355, pl. 26,
fig. 15.
The Anomia simplex is both common and characteristic in
the Gatun of Costa Rica as well as in certain parts of the Prov-
210 BULLETIN 39 382
ince of Colton, Panama, near the Canal Zone. It frequently
gives rise to important zones in the lower part of the Gatun for-
mation, where it occurs in large numbers associated with Pecten
levicostatus Toula.
The specimens are generally large, thin-shelled, smooth and
irregular in form, due to the object on which they were attach-
ed. They average for the upper valves about 35 mm in
height.
Anomia simplex appears as a Miocene fossil in the eastern
United States for the first time in the St. Mary’s formation of
Maryland in the upper part of the Lower Chesapeake. It occurs
throughout the Upper Chesapeake Miocene and Pliocene and
passes into the recent fauna. Locally asa fossil it may become
very abundant, as at the base of the Pliocene overlying Upper
Chesapeake Miocene along the shores of Lake Waccamaw,
North Carolina. It occurs rarely as a Miocene fossil in Santo
Domingo, where it has been recorded by Gabb and Maury. In
Mexico, it occurs in the Miocene of Santa Rosa, Vera Cruz as
figured by Bose.
Gatun Stage: Several localities, Prov. of Colon, Panama.
Middle Creek. Comadre Creek.
Fone and Hotel Creeks.
Hill No. 2, Banana River, Soury Creek, ete.
Genus PLACUNANOMIA, Broderip
Placunanomia lithobleta Dall Plate 21, figures 3, 4, 5
Placunanomtia lithobleta Dall, 1898,Trans. Wagner Free Inst. Sci.,vol.
3; Ptr 4p. 778:
Placunanomia lithobleta Maury, 1917, Bull. Amer. Pal., vol. 5, p. 356,
Di s4A igs ine:
The Costa Rican specimens are frequently widely plicate,
nearly as much asin the Duplin Miocene P. plicata Tuomey and
Holmes, but the surface is marked with fine, wavy, radial threads
so that even very small fragments of this species are very easily
recognized. It occurs in the Miocene of Jamaica, its type local-
ity, and in the Miocene of Santo Domingo.
Gatun Stage: Hill 3, Banana River.
383 Costa RrcA MrockENE—OLSSON 211
: ORDER ANOMALODESMACEA
Genus THRACIA, Blainville
Subgenus CYATHODONTA, Conrad
Thracia Tristani, n. sp. Plate 20, fig. 3
Shell thin, subquadrate, the left valve slightly convex; the
beaks situated at the posterior 1-3 of the shell, are low and in-
conspicuous; anterior end, wide and broadly rounded, the poster-
ior contracted, depressed and truncated at its extremity; the sur-
face is sculptured with regular wave-like folds, which on the
middle of the shell are concentric, but are oblique on the anter-
ior portion; they are lacking from the posterior area; in addi-
tion the whole surface is very finely pustulose and crossed by
very indistinct growth-lines.
Length 42, height 30, diameter of the left valve 7.00 mm-
The Cyathodonta gatunensis Toula of the Canal Zone isa
similiar but larger species, reaching a length of 52 mm. It dif-
fers from the Costa Rican species in its greater proportional
length, the more central beaks and in its sculpture, which ac-
cording to Toula’s figure is strictly concentric.
The C. undulata Conrad, a recent species from the Gulf of
California, is sculptured like 77zsfanz with oblique riblets but
the left valve is depressed and strongly flattened.
This species is named for Professor J. Fidel Tristan, Director
of the National Museum of Costa Rica.
Gatun Stage: Rio Betey.
212 BULLETIN 39 384
ORDER TELEODESMACEA
Genus CRASSATELLITES, Kruger
Crassateilites Reevei Gabb Plate 20, figure 4
Crassatella antillarum Reeve? C. Reevet Gabb 1873, Trans. Amer.
Phil. Soc. vol. 15, p. 252.
Crassatellites reevei Brown and Pilsbry, 1913, Proc. Acad. Nat. Sci.
Phila., vol. 64, p. 515, pl. 23, fig. 5.
The Crassatellites Reevei Gabb is not an uncommon fossil
in the Gatun beds of western Panama and Costa Rica. Our
specimens agree quite closely with Brown and Pilsbry’s figure of
the single right valve collected by Gabb in the Miocene of Santo
Domingo. Young and full-grown specimens show a strong
posterior angulation which extends from the beak to the poster-
ior ventral extremity. “The posterior end is obliquely truncated
but less so in young shells. ‘The surface sculpture consists of
even, concentric costze which on the slightly flattened umbos are
a little wider apart asin the Cvassatedlites of the section Scam-
bula Conrad. In this later character, our shells disagree with
Brown and Pilsbry’s redescription of Gabb’s type specimen, in
which the sculpture is said to extend without any irregularity
upon the rather flattened umbos.
The C. mediamericanus Brown and Pilsbry from Sapote,
Costa Rica is an earlier shell and is higher, less obliquely trun-
cated behind, more coarsely sculptured and lacks the posterior
angulation. In these characters except the lack of the posterior-
umbonal angulation it is like the young shells of Reevez.
Length 55, height 36.5, semi-diameter 10.5 mm.
50 35 diameter Pe iGaabonl,
58 42 seni-diameter 14 mm.
385 Costa RicA MI0ockENE—OLSSON 202
Gatun Stage: Water Cay.
Old Man Sam Creek, Coll. No. 6:
Banana River.
Subgenus CRASSINELLA, Guppy
Crassatellites bowdenensis Dall, variety costaricensis, n. var.
Plate 29, figure 12
Cf. Crassatellites (Crassinella) bowdenensis Dall, 1903, Trans. Wagner
Hreevinsty sci Vol) 3 pt. GN p.T4 76s" pla) 50st eSy Qua.
Shell small, triangular, slightly convex, subequilateral, the
anterior end generally a little longer and more pointed; beaks
triangular, slightly pointing forwards; lunule large, narrowly
lanceolate, smooth; escutcheon small; surface sculptured with
small, even, concentric riblets which on some shells may be
strong and persistent over the greater part of the shell, or on
some specimen sub-obsolete; interior of shell cavity shallow, with
a moderately heavy hinge.
Wength!)2o0, height 2.75, diameter }75 mm:
Ba5O) 250) 1715) Mal
The typical dowdenenszs Dall from the Bowden Miocene of
Jamaica is a more solid, more convex and triangular shell. The
variety costaricensis is very abundant at several localities in Costa
Rica in certain horizons. The sculpture varies from strong,
even, concentric riblets covering the greater part of the shell, to
forms which are nearly smooth.
Gatun Stage: Hiill ra, Banana River.
Middle Creek.
Zone 3, Quttana Creek.
Crassatellites midiensis, n. sp. Plate 20, figure 11
Shell small, trigonal, slightly convex and equilateral; the
beaks are central and erect; the dorsal margins straight and
meeting at the beaks nearly at right angles; the surface is sculp-
tured with subregular, distant, concentric lamelle-like riblets,
214 BULLETIN 39 386
numbering about 10; the interspaces carry irregular, finer, con-
centric lines; lunule narrowly lanceolate, with the escutcheon
narrow and smaller; interior of shell cavity shallow, with a mod-
erately heavy hinge.
Length 3.25, height 3.25, diameter of one valve 1.00 mm.
A small, rare species, somewhat like the C. Guppyz Dall of
the Miocene of Jamaica, but more trigonal, with more central
beaks and somewhat different sculpture. The sculpture and
form are very similar to specimens of C. galvestonensis Harris in
the Cornell Collection from the Chesapeake Miocene of York-
town, Va. ‘This latter species is generally much worn but per-
fect specimens have a coarse concentric sculpture like that of
lunulatus Conrad. ‘The Costa Rican shells are however very
much smaller.
Gatun Stage: Middle Creek, C. R.
Coll. 4, East Grape Point Creek.
Water Cay, Panama.
Genus CARDITA, (Bruguiére) Lamarck
Cardita caribbeanensis, n. sp. Plate 26, figure 22
Shell small, moderately convex, elongate, its length a little
less than twice its height; beaks very near the anterior end are
small but distinct; an umbonal ridge extends from the beaks to
the posterior-ventral extremity and along which and bordering
the ribs are largest; the right valve with about 20, hight, narrow
and sharply nodulose ribs, separated by wide, triangular, smooth
interspaces; 6 of the ribs are situated on the posterior-dorsal
submargins, and of which the 4th and 5th from the dorsal mar-
gin are much smaller than the others; of the remaining ribs, the
largest are found on the umbonal ridge, and become progressive-
ly smaller anteriorly; lunule small but distinct; hinge of the
right valve that of typical Cardita, with 3 cardinal teeth of which
the middle one is much the largest, no laterals; ventral margin,
internally fluted in harmony with the external ribs.
387 Costa RicA MIocENE—OLSSON 215
Length 18, height 12.50, diameter of the right valve
5.50 mim.
This small species represented by a single right valve in our
collection appears to be a true Cavdzfa, its hinge being essential-
ly that of the recent West Coast Cardzta laticostata Sowerby.
With the exception of a few doubtful species in the Eocene of
United States, true Cardita has heretofore not been found as a
fossil in the Americas and its recent American species are con-
fined to the Pacific coast.
Compared with the recent Cardzta laticostata Sowerby, which
is abundant along the Pacific Coast of Panama, the fossil shell
differs in its smaller size, smaller umbos and in form. Its lunule
is like that of /aticostata.
Gatun Stage: Banana River.
Section CARDITAMERA, Conrad
Cardita matima, n. sp. Plate 32, figure 8
Shell robust, solid, strongly inequilateral with the beaks
situated at the anterior extremity of the shell; umbos
large and full with strongly curved beaks above the small,
sunken lunule; outline of the shell from within is subrectangu-
lar with straight ventral and posterior sides which meet at
nearly right angles; sculpture of about 18 strong, heavy cord-
like ribs, separated by interspaces 1-3 of their width; the ribs
are strongly nodulated on the disk and umbonal portion of the
shell, but below they are simply irregularly wrinkled; a deep
furrow extends from the beaks to the posterior extremity, just
above the umbonal angle; the rib forming the summit of the
ridge between the umbonal furrow and the hinge-line is larger
than those on each side; shell cavity deep with a crudely den-
ticulated ventral and posterior margin, hinge large (broken in
our specimen), with a well-developed anterior lateral tooth in its
left valve.
Length 30, height 25, semi-diameter 12 mm.
216 BULLETIN 39 288
But a single specimen of this very distinet spceies was col-
lected by Dr. A. C. Veatch from the Upper Gatun of Old Man
Sam Creek in eastern Costa Rica. It differs from the other
known American Carditameras by its convex shell and strongly
curved beaks. The ribs are large and strongly nodulated at
first, becoming irregularly wrinkled near the ventral margin.
The hinge is unfortunately broken on our specimen, a left valve,
but the small anterior lateral still remains which shows this shell
to belong to the section Carditamera
Gatun Stage: Old Man Sam Creek, 1 mile south from
shore.
Genus VENERICARDIA, Lamarck
Venericardia scabricostata Guppy Plate 32, figures 14, 17
Cardita scabricostata Guppy, 1866, Quart. Journ. Geol. Soc. London,
Voll22h pi 2oB4 ple ce yon nos
Cardita scabricostata Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
252.
Venericardia scabricostata Maury, 1917, Bull. Amer. Pal., vol. 5, p.
ZBO2s plus Milo:
This species originally described by Guppy from the Bowden
beds of Jamaica is common in the Miocene of Costa Rica and
frequently forms small zones in the Gatun beds. It occurs also
in the Miocene of Santo Domingo.
The ribs numbering about 18 are high, narrow and strongly
granulated. The interspaces are wide and generally smooth.
Gatun Stage: Zone E, G, Saury creek.
Zone l, 3, East Grape Point creek.
Coll. 5, Red Cliff creek.
fleadwaters of Middle creek.
Coll. C, Comadre creek.
Venericardia Terryi, n. sp. Plate 32, figures 12, 13
Shell of medium size, solid, moderately convex and sub-cir-
G3
e=2)
Ns)
Costa Rica M1ockNE—OLSSON Zi
cular in form; beaks situated well forward and but slightly
back of the extreme anterior end of the shell; the beak itself is
small and loosely coiled over the small sunken lunule; sculpture
consists of about 13, low, wide, smooth ribs which fit so closely
together that there are no interspaces between them; the ribs are
rounded in section on the umbos, wide and low ventally; the ribs
are largest about the middle, small and fading away on the poster-
ior and anterior dorsal submargins; interior concealed in a hard
sandstone matrix.
Length 22, height 21, semi-diameter 6 mm.
The first specimens of this interesting species was collected
by Mr. R. Terry from some Gatun sandstones in Quitana Creek,
a small tributary streams of the Rio Betey. In that locality, it
is an abundant species and occurs in small zones.
It is related to the recent Pacific, V. crassicostata Sowerby
but is a much smaller species.
Gatun Stage: Outtana Creek.
fiili No. 3, Banana River.
Subgenus PLEUROMERIS, Conrad
Venericardia Conradiana Gabb, var. limonensis, n. var. Plate 26, figs. 19, 20
Cardita Conradiana Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8,
2nd series, p. 377, pl. 47, fig. 79.
Shell small, subcircular, moderately convex; umbos high
and prominent, with small, inconspicuous beaks at the anterior
1-3 of the shell; the sculpture consists of about 18 heavy, gran-
ulated ribs, separated by somewhat narrower interspaces; interi-
or of the shell deep, the hinge with 2 cardinal teeth and a small
posterior and anterior lateral.
Length 7.00, height 7.00, diameter of the left valve
3.00 mm.
Gabb specimen described from the Pliocene beds of the
Limon Peninsula, measures 1% mm in length. His enlarged
figure is very similiar to the present shell, in its subcircular fori
218 BULLETIN 39 390
and in number of ribs. The figure of the hinge shows the two
cardinal teeth but no laterals.
The common ¢ridentata Say of the Upper Chesapeake Mio-
cene and recent fauna of the West Indies differs mainly in its
more triangular form.
Gatun Stage: Port Limon.
Genus CHAMA, (Linné) Bruguiére
Chama congregata Conrad Plate 28, figure 11
Chama congregata Conrad, 1833, American Journal of Science, vol. 23,
Pp. 341.
Chama congregata Conrad, 1838, Fossils of the Medial Tertiary, p. 32,
plen7 A tiga:
Chama congregata Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3,
pt. 6, p. 1400.
Chama congregatoides Maury, 1917, Bull. Amer. Pal., vol. 5, p. 200,
pl. 33, fig. 8.
Our specimens from Costa Rica are all small and belong to
the upper or free valve, which in this species is the right. These
specimens cannot be distinguished from true congregaza of the
same size from the eastern United States, where it is an abund-
ant fossil throughout most of the Chesapeake Miocene. The
upper or right valve is finely, radiately frilled or fluted and these
frills are cut by the concentric lamelle. The attached or left
valve is more coarsely sculptured,both radially and concentrical-
ly.
Gatun Stage: Water Cay.
Coll. Va Red i Glo Cxeek:
Genus ECHINOCHAMA, Fischer
Echinochama antiquata Dall Plate 28, figure 8
Chama antiquata Guppy, 1866, Quart. Journ. Geol. Soc. London, vol.
22, p. 294. Not of Linné.
Echinochama antiquata Dall, 1903, Trans. Wagner Free Inst. Sci., vol.
3, pt. 6, p. 1404, pl. 54, fig. 9.
391 Costa RicA MIocCENE—OLSSON 219
Not “chinochana antiquata Maury, 1917, Buil. Amer. Pal., vol. 5, p.
365, pl. 33, fig. 10. F. vaguensis Maury.
This species described by Dall from the Bowden beds of
Jamaica differs from the common, recent avcinella Linné of the
West Indies, by its more numerous, irregular and lower ribs
which bear only small, short spines. In arcenella the middle
ribs are heavy and bear large, stump-like spines and the wide
interspaces are reticulated with a coarse, mesh-like sculpture.
Gatun Stage: Middle Creek,
Coll. 6, Old Man Sam Creek.
Flill No. 1, Banana River.
Rio Blanco.
Echinocama yaquensis Maury Plate 28, figure 5
Chama arcinella Moore, 1853, Quart. Journ. Geol. Soc. London, vol.
9, p. 130. Not of Linné.
Chama arcinella Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
251:
Chama arcinella Guppy, 1874, Quart. Journ. Geol. Soc. London, vol.
Behn Ou eee
E-chinochama antiquata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 365,
pliiz3, fg. ro!) Not of Dall.
Echinochama antiquata var. yaquensis Maury, 1917, op. cit. p. 365, pl.
Bos SIT L2.
E-chinochama trachyderma Pilsbry and Johnson, 1917, Proc. Acad.
Nat. Sci. Phila., p. 197.
The £Echinochama yaguensis Maury from the Miocene of
Santo Domingo, where it is the prevaling species, differs from
the antiguata by the reduced number of its ribs, which in some
cases are nearly obsolete and in having its surface covered with
a coarse, mesh-like sculpture of raised pustules.
In Costa Rica, the azzzqguata is the common species, but we
have also a single species of yagzenszs collected from the lower
Gatun near Old Harbor.
220 BULLETIN 39 392
Gatun Stage: Old Harbor.
(Gong. (Cs be
Genus DIPLODONTA, Brown
Section DIPLODONTA, s. s.
Diplodonta collina, n. sp. Plate 32, figure 15
Shell small, subcircular, convex; umbos wide, with small
inconspicuous beaks, slightly posterior of the middle; anterior
and posterior extremities well rounded into the base forming a
part of the same circle; surface sculptured with fine, even, con-
centric threads which are nearly lacking from the smooth umbos;
hinge normal, and a plain ventral margin.
Length 6.75, height 6.75, diameter of the right valve
1.60 min.
A small shell resembling the Dzplodonta nucletformis Wag-
ner of the Upper Chesapeake Miocene and the recent West In-
dian fauna, but less convex and more finely sculptured.
Gatun Stage: Middle Creek.
Section FELANIELLA, Dall
Diplodonta insula, n. sp. Plate 32, figure 21
Shell small, slightly convex, subcircular; beaks slightly
posterior of the middle so that the posterior is a little longer
than the anterior; beaks low and inconspicuous; both anterior
and posterior extremities widely rounded, the posterior and the
ventral margin forming a part of the same circle; surface with
fine, concentric threads heaviest on the extremities, but nearly
lacking from the umbonal area; interior of shell shallow, the
right valve with a strongly bifid posterior cardinal; ventral mar-
gin plain.
Length 6.60, height 6.50, diameter of the right valve
1.25 mm.
393 Costa RicA M1ockNE—OLSSON 221
A small, subcircular shell, neatly sculptured with fine, con-
centric threads. DY. mznor Dall from the Bowden beds is strong-
ly oblique.
Gatun Stage: Water Cay.
Genus MYRTAEA, Turton
Myrtaea limoniana Dall Plate 32, figures II, 23
Myrtea limoniana Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3,
pt. 6, p. 1358, pl. 52, fig. ro.
A small, thin and slightly convex shell, its surface sculptur-
ed with fine, thin and sharp, elevated, concentric lamelle. We
have several specimens from the coral limestones of Port Limon
as well asalarger specimen from Zone 4, of Red Cliff Creek.
This later shell measures: heigth 9.50, length 10.00, diameter of
the right valve 2.50 mm. It occurs also in the Bowden beds of
Jamaica.
Gatun Stage; Port Limon.
Coll. 4, Red Cliff creek.
Genus PHACOIDES, Blainville
Subgenus CALLUCINA, Dall
Phacoides radians Conrad, var. medioamericanus, n. var. Plate 32 fig. 16
The Phacotdes radians Conrad is a fossil of the Upper Ches-
apeake Miocene of the Carolinas but has continued through the
Pliocene to the recent fauna. The Costa Rican fossils differ from
typical radians in having their umbos sculptured with about
6 ribs much stronger than the concentric, while in typical
radians, the ribs are more numerous and about equal to the con-
centric lines. The medioamericanus is also smaller as may be
seen in the measurements given below, while vadzans is large
(20 mm) and often quite convex.
Length 9.5, height 9, semi-diametes 2.5 mm.
222 BULLETIN 39 394
Gatun Stage: Hleadwater of Middle creek.
Old Man Sam creek, 1 mile south of the beach.
Subgenus HERE, Gabb
Section CAVILUCINA, Fischer
Phacoides trisuleatus Conrad
Lucina trisulcata Conrad, 1841, Trans. Amer. Assoc. Nat. and Geol.,
vol. I. p. IIo.
Lucina trisulcata Conrad, 1845, Fossils of the Medial Tertairy, p. 71,
pl. 40, fig. 5. ?
Phacoides (Cavilucina) trisulcatus Dall, 1903, Trans. Wagner Free
Inst. Sci., vol. 3, pt. 6, p. 1369.
Few specimens from the Gatun of Middle Creek. They
are identical with typical specimens from the Chesapeake Mio-
cene of the eastern United States. The species also occurs in the
recent fauna.
Gatun Stage: Middle creek.
Old Man Sam creek.
Subgenus LUCINISCA, Dall
Phacoides costaricensis, n. sp. Plate 32, figure 5
Shell subcircular, moderately thick and but slightly convex:
beaks small and inconspicuous, a little in front of the middle;
dorsal area impressed and well defined; sculpture consists of
strong, regular, radial threads, widely separated by interspaces
2 or 3 times the width of the threads themselves; they are cross-
ed by distant but regular concentric ridges, the intersection form-
ing small beads or granulations, strongest on the anterior and
posterior extremities; the lunule is narrow, lanceolate and with
the concentric sculpture predominanting; interior of shell cavity
rather shallow, the ventral margin coarsely serrated by the ex-
ternal sculpture.
295 Costa Rica MrocENE—OLSSON 22
WwW
Length 11.50, height 11.00, diameter of the left valve
2.25 mm.
This species is recognized by its depressed or but slightly
convex shell and coarse sculpture, of which the radial is a little
stronger. Phacotdes cribrarius Say and nassula Conrad are both
more convex, with the concentric lamellee more distantly spaced
on the umbos and dominating over the radial. PP. murtcatus
Spengler of the West Indies is much more finely sculptured.
We have also a young shell from Water Cay which is pos-
sibly P. zassula, var. caloosana Dall of the Florida Pliocene. It
is very close to specimens of that species in the Cornell collec-
tion.
Gatun Stage: Middle creek.
Phacoides hocasensis, n. sp. Plate 32, figure 6
Shell thin, subcircular, depressed; dorsal area well defined
by a marked change in sculpture; beak small, but distinct, placed
a little in front of the middle; the sculpture on the disk of the
shell is fine, composed of subequal, radial threads, separated by
narrow interspaces; the ribs and interspaces are crossed by fine,
concentric threads or ridges which produce a finely scabrous
surface; the concentric threads are a little more widely spaced
on the umbos; the dorsal area is defined by a stronger radial, be-
yond which follows a wide radial band, strongly sculptured with
the concentric threads or lamelle but lacking in radials; the up-
per half of the dorsal band bears in addition to the concentric
lamelle, 2 or 3 weak but scabrous radial threads; the lunule is
very narrow, lanceolate and with a few strong, concentric lame-
lle; interior of the shell cavity shallow, with the ventral margin
finely crenulated.
Length 10.00, height 9.25, diameter of the left valve
2.00 mim.
A depressed and very finely sculptured species whose gener-
al form is that of P. muricatus Spengler of the recent West In-
dian fauna. It is more regularly sculptured than the P. hAzspar-
224 BULLETIN 39 396
zolana Maury of the Cercado formation of Santo Domingo but
the main difference lies in their dorsal areas, that of hzsbantolana
Maury being sculptured more or less like the rest of the shell;
in bocasensts, as described above, the sculpture of the dorsal area
is dominantly concentric, only the upper half bearing 2 or 3, low
but scabrous radials.
Gatun Stage: Bocas del Toro.
Subgenus PARVILUCINA, Dall
Section BELLUCINA, Dall
Phacoides actinus Dall Plate 32, figure 24
Phacoides ( Bellucina) actinus Dall, 1903, Trans. Wagner Free Inst.
Sci., vol. 3, pt. 6, p. 1385, pl. 52, fig. 3.
Dall gives for his typical specimens from the Bowden beds
of Jamaica, a height of 4.50 mm, while the Panama shells, all
from Bocas Island, do not exceed 3 mm in height but agree in
other respects. It is also recorded by Maury from the upper
part of the Cercado formation of Santo Domingo.
Gatun Stage: Bocas del Toro.
Genus DIVARICELLA, von Martens
Divaricella quadrisulcata d’Orbigny Plate 32, figure 20
Lucina divaricata Conrad, 1840, Fossils of the Medical Tertiary, p. 39,
pl. 20, fig. 3, but not of Linné.
Lucina quadrisulcata d’Orbigny, 1846, Voy. Am. Mer., p. 584.
Divaricella quadrisulcata Dall, 1903, Trans. Wagner Free Inst. Sci.,
vol. 3, pt. 6, p. 1389.
Lucina ( Divaricella) quadrisulcata Bose, 1906, Boll. de Inst. Geol. de
Mexico, numero 22, p. 77, pl. II, figs. 2, 3.
This species is very rare in Costa Rica. The shells are rath-
er thin and generally more finely sculptured and approach in
this respect the D. compsa Dall of the Florida Pliocene. Our
larges specimen measures: height 14 m, length 1o mm, diameter
397 Costa R1icA MrocENE—OLSSON 225
of left valve 4.50 mm. The D. prevaricata Guppy of the Mio-
cene of Santo Domingo and Jamaica measures only 8 mm in
height.
The D. guadrisulcata is a recent species of the West Indies
appearing in the east coast Miocene of the United States for the
first time in the Upper Chesapeake.
Gatun Stage:Coll. 4,5, Red Cliff creek.
Middle creek.
Genus CARDIUM, Linné
Subgenus TRACHYCARDIUM, Moerch
Cardium stiriatum Brown and Pilsbry Plate 27, figures 4, 5
Cardium (Trachycardinm) stiriatum Brown and Pilsbry, 1911, Proc.
Acad. Nat. Sci. Phila., vol. 63, p. 366, pl. 28, fig. 11.
This is most elegant of the several species of Cardium in
the Miocene of Panama and Costa Rica. The shell is high and
inflated with about 30 radial ribs which are peculiarly sculptur-
ed with twisted or obliquely projecting tuberclers placed on the
posterior side of the ribs and overhanging the adjacent inter-
space. Large specimens may reach a height 50 mm. It is re-
lated to the recent West Coast Cardium belcheri Broderip.
Gatun Stage: Gatun, C. Z.
Water Cay, Panama.
Hill No. 1, Banaxe River.
Cardium dominicense Gabb Plate 27, figure 2
Cardium )Trachycardium) dominicense Gabb, 1873, Trans. Amer.
Phil. Soc., vol. 15, p. 25.
Cardium ( Trachycardium) dominicense Gabb, 1881, Journ. Acad. Nat.
Sci. Phila., 2nd series, vol. 8, p. 344.
Cardium (Trachycardium) gatunensts Toula, 1908, Jahrb. der K-K
Geol. Reichsanstalt Wien, vol. 58, p. 720, pl. 27, fig. 4. Not Cara-
tum (Fragum) gatunensis Dall, 1900.
226 BULLETIN 39 308
Cardiuin ( Trachycardium ) dominicense Brown and Pilsbry, 1911, Proc.
Acad. Nat. Sci. Phila., vol. 63, p. 367.
A large species abundant in the Canal Zone but usually only
as internal casts. It appears to be rare in Costa Rica, our col-
ection containing only a few fragments of a large individual
from Hast Grape Point Creek. It is distinguished not only by
its large size, but very numerous (60+) fine ribs.
Gatun Stage: Gatun, C. Z.
East Grape Point Creek, C. R.
APOE, 3 Gara G-aoo))
Cardium costaricanum, n. sp. Plate 27. neurens
Shell ovate, solid, inflated; umbos wide and full with high
beaks; sculpture of about 24 strong ribs, rounded or subtrian-
gular in section with their apex leaning posteriorly except those
on the posterior slope which lean anteriorly; interspaces narraw,
formed by the sides of the ribs; the ribs are mostly smooth ex-
cept those on the anterior half of the shell which have their pos-
terior side more or less granulated; this granulation is found al-
so on a few of the central ribs near their ventral margin; hinge
narrow; internal margin fluted in harmony with the external
ribs.
Length 24.50, height 34, semi-diameter 15 mm.
This species is not uncommon in Costa Rica and may easily
be confused with the C. dominicanum Dall. It differs in its
larger side, narrow umbos, more pointed beaks and more flatten-
ed ribs.
Gatun Stage: Coll. 4, East Grape Point Creek.
Subgenus FRAGUM, Bolton
Cardium medium Linnzeus Plate 27, figure 6
Cardinm medium T,innzeus, 1758, Sys. Nat. ed. 10, p. 678.
Cardium medium Reeve, 1844, Conch. Icon., Cardium, pl. 6. fig.
30.
Cardium (Fragum) medium Dall, 1900, Trans. Wagner Free Inst.
SCiivolwa ap tsii pw lLOLe
3909 CosTa R1icA MiocENE—OLSSON 227
A recent West Indian species but also common as an Up-
per Miocene and Pliocene fossil along the Atlantic coastal plain
of eastern United States. The ribs number 24 to 27 in front of
the umbonal angle. Dall has described a Cardium (Fragum)
gatunensis from Gatun, on which the ribs number only 16 in
front of the truncation and 10 behind.
Gatun Stage: Port Limon.
Section TRIGONIOCARDIA Dall
Cardium heredium, n. sp. Plate 27, figure to
Shell small, solid, oblique, convex and subquadrate; strong-
ly carinate on the posterior-umbonal slope and truncated; umbos
high and prominent; shell sculptured anterior to the truncation
with 12 or 13 strong ribs, separated by interspaces about % to
Y% as wide; there are 7 or 8 smaller ribs on the posterior trun-
cation; the interspaces are finely sculptured with small, elevated
cross-threads; the anterior set of ribs on the type specimens are
wide, subequal and smooth; the posterior are unequal, with the
three, bordering the umboal angulation much the smaller and
bearing small and scattered pustules; on the posterior ribs where
the pustules are lacking, are small dot-like depressions.
Height 11.50, length 11.50, diameter 6.00 mm.
This species approaches the Cardium apateticum Dall from
the Oak Grove sands of Lower Miocene age in Florida. It dif-
fers by its more oblique umbos and proportionately higher shell.
The pustules are variously developed and on some shells are
more strongly developed and may also appear on some of the
ribs anterior to the truncation. These more strongly scalptured
shells may prove with more material to belong to another
species.
Gatun Stage: Rio Betey.
Banana River.
Fistrella River.
228 BULLETIN 39 460
Subgenus PAPYRIDEA, Swainson
Cardium spinosum Meuschen, var. Turtoni Dall Plate 27, figure I
Cardium (Papyridea) spinosum var. Turtoni Dall, tg00, Trans. Wag-
mer Free Inst., Sci., vol. 3, pt. 5; p. 1108.
The fossil shells from Costa Rica, are referrible to this var-
iety of spimosum, distinguished by having the interspaces be-
tween the ribs marked with a flat thread, between sharp grooves.
This variety is also found fossil in the Chesapeake Miocene at
Jackson Bluff Florida and in the Pliocene Caloosahatchie beds of
the same state. It the recent fauna it is confined, according to
Dall, to the eastern Atlantic, with the typical spzzosum in the
West Indies and a third variety aspersum Sowerby along the
Pacific side.
Gatun Stage: Port Limon.
Subgenus LAEVICARDIUM, Swainson
Cardium serratum Linnzeus Plate 27, figures II, 12
Cardium serratum Linneeus, 1758, Syst. Nat., ed. 10, p. 680.
Cardium (Levicardium) serratum Dall, tg00, Trans. Wagner Free
Insts Sciesivolai2s ptrs4) p.ULtos
Cardium (Levicardium) serratum Brown and Pilsbry, 1911, Proc
Acad. Nat. Sci. Phila., vol. 63, p. 367.
Cardium (Levicardium) serratum Maury, 1917, Bull. Amer. Pal., vol.
iy 1 27s) OLA) Ios) a8lreZ4 Xe)
Most of the fossil shells are indistinguishable from recent
examples of servatum from the West Indies. Others shells are
decidely oblique and approach in form the C. sublineatum Con-
rad of the Upper Chesapeake Miocene of eastern United States.
The C. venustum Gabb from the Miocene of Santo Domingo is
very distinct from serratum and characterized by its faintly flex-
uous outline, strong radial striz, andin having the lower cardin-
al tooth large and strong.
401 ' Costa Rica MriockENE—OLSSON 229
Gatun Stage: Gatun, C. Z.
Muddley Greek Gj 0h.
BO WA, (Go I
Genus PROTOCARDIA, Beyrich
Protocardia costaricensis, n. sp. Plate 27, figure 13
Shell small, rather thin, convex; beaks a little anterior to
the middle, with high, inflated umbos; the anterior end is wide-
ly rounded, the posterior somewhat truncated; the posterior area
is well differentiated and occupies the posterior one-third of the
shell and is sculptured with numerous small radial threads sep-
arated by wider interspaces; these interspaces are finely sculp-
tured with cross-threads; anterior of the posterior area the shell
surface is sculptured with small radial threads which are finely
beaded; on the anterior end of the shell, the beading is more ir-
regular and is seen to be made by the concentric sculpture.
Height 18, length 21, diameter of the left valve 7 mm.
This species differs from both P. jamazcensis Dall from the
Bowden beds of Jamaica and P. zslahispaniole Maury from the
Miocene of Santo Domingo by its smaller posterior area. It ap-
proaches more closely the Dominican shell but is shorter and
more coarsely sculptured.
The P. Newberryana Gabb from Gatun is based on a large
internal cast which is entirely unrecognizable.
Gatun Stage: Comadre creek.
Subgenus LOPHOCARDIUM, Fischer
Protocardia gurabica Maury Plate 27, figures 7-9
Protocardia gurabica Maury, 1917, Bull. Amer, Pal., vol. 5, p. 377, pl.
36, fig. Io,
Shell thin, inflated, a little longer than high and with the
greatest convexity about the center of the shell; the umbos are
high, prominent and wide, and project well above the hinge
230 BULLETIN 39 A402
margin; the anterior end is well rounded into the base; the pos-
terior end is more contracted and gaping slightly at its extremi-
ty; surface sculpture on the body of the shell, anteriorly con-
sists of narrow, concentric wrinkles which are best developed on
the anterior end but become obsolete on the middle of the shell
and posteriorly; in addition the whole surface carries fine,
sub-obsolete radial threads; on the posterior area, the sculpture
is of heavier and coarser radial threads, lacking the concentric,
except t he irregular growth lines.
Height 56, length 32, diameter 20 mm.
The subgenus Lophocardium, of which there are two recent
species along the West Coast of Mexico, differs from the true
Protocardia, in its gaping posterior end and lack of lateral teeth
as well-as in important difference in its anatomical structure.
The shells are strongly convex, with high, inflated umbos and
are extremily thin and fragile.
The P. gurabica was described by Dr. Maury from the Gur-
abo formation of Santo Domingo, and compared by her with the
Vickburgian P. dzversa Conrad, but it appears to me to be a typ-
ical Lophocardium and representing the first member of this in-
teresting subgenus to be found in the fossil state. As her speci-
men is very fragmentary, I have redescribed the species from
better preserved Panama and Costa Rican material. The poster-
ior end is gaping and the internal mold shows no impression of
lateral teeth along the hinge margin. It approaches closely the
L. Annette Yall, dredged from 8 to 27 fathoms in the Gulf of
California, except that it is proportionately longer and lacks the
concentric wrinkles on its posterior area. The L. Cumingz Brod-
erip, the type of the subgenus, carries between its posterior area
and the general surface of the shell, a thin radial lamina or crest,
extending from the umbos to the ventral margin.
The fossil shell is extremily thin. It generally occurs simp-
ly as internal molds which may still retains a part of the shell
or impression of its sculpture on its surface.
403 Costa Rica M1ocENE—OLSSON 231
Gatun Stage: Gatun, and Mt. Hope, C. Z.
Water Cay.
Margarita Trail, C. R.
Genus DOSINIA, Scopoli
Section DOSINIDEA, Dall
Dosinia acetabulum Conrad Plate 31, figure I
Artemis acetabulum Conrad, 1833, Fossils of the Tertiary Formations,
PZOvepl won fie. Te
Dosinia acetabulum Bose, 1906, Boll. de Inst. Geol. de Mexico, num-
ELO}2 2 py Oly iplo Tl, Ss. 17), Wl
Dosinia (Artemis) cf. acetabulum Toula, 1908, Jahrb. der K-K, Geol.
Reichsanstalt, Wien, vol. 58, p. 727, pl. 27, figs. 8, 8a.
The Dosinia acetabulum has been recorded by Bose from
Mexico and by Toula from Panama, but their determinations
were based on imperfect material or casts. The Costa Rican
collection however contains a large series of finely preserved Do-
sinias which are unquestionably the Dosinza acetabulum Conrad
of the Chesapeake Miocene.
The surface of the Costa Rican specimens are generally
neatly sculptured with even, concentric bands or ribbons which
on the middle of the shell disk average 2 mm in width. The
form of the shell is subcircular and like the typical D. acetabu-
Jum of the Yorktown formation of Virginia and North Carolina.
The Costa Rican shells are equally as large as the Chesapeake
shell measuring in height 70 mm or more.
The Dosinia acetabulum in the eastern United States, is a
most characteristic fossil of the Chesapeake Miocene, where its
range extends throughout most of the lower and upper Chesa-
peake formations. It is lacking from the Lower Miocene or the
Alum Bluff formation of Florida, as well as from the succeeding
Pliocene. Inits range through the Chesakeake Miocene, it
gives rise to several well-marked varieties of which the broad
222; BULLETIN 39 404
subcircular shell of the Upper Chesapeake is the typical form.
The Costa Rican shells are most like the Upper Chesapeake
typical acetabulum.
Brown and Pilsbry have described a smaller and more finely
sculptured species from the Gatun Spillway as delicatisstma.
This spceies is represented in our collection from the same place
and appears to be distinct. Toula’s specimen from Gatun, how-
ever, is a true acetabulum.
The Dosinia acetabulum is a common fossil in the Gatun
beds of Costa Rica, where it occurs most abundautly in the
lower part of the formation.
Gatun Stage: Gatun, C. Z.
Flotel and Hone Creek.
Pumbri Creek.
fiill No. 37, Banana River.
Coll. 6, Estrella River.
Genus CLEMENTIA, Gray
Clementia dariena Conrad Plate 31, figure 4
Meretrix dariena Conrad, 1856, Pacific R. R. Reports, vol. 5, p. 328,
pl. 6, fig. 55.
Clementia dariena Gabb, 1881, Journ. Nat. Sci. Phila., vol. 8, p, 344,
DLV AAN fet 6s
Clementia dariena Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3,
DEVON AI225.
Clementia daviena Toula, 1908, Jahrb. der K-K. Geol. Reichsanstalt,
vol. 58, p. 7:25, ple 27,, fges.o,) 10:
Clementia dariena Hubbard, 1920, Scientific Survey of P. R. and the
Virgin Islands, vol. 3, pt. 2, p. 118, pl. 19, figs 10-12.
This species is very common in the lower Gatun of the
Canal Zone. It is very rare in western Panama and in Costa
Rica, where it occurs in the lower part of the Gatun formation
and in the Uscari shales. Romanes* has recorded it from Bar-
* Quart. Journ. Geol. Soc. London, 1912, vol. 68, p. 125.
405 Costa Rica M1rocENE—OLSSON 233
ranca near Punta Arenas onthe Pacific coast of Costa Rica,
from beds probably equivalent to the Lower Gatun or to the
Uscari.
The shell is very thin and fragile, and hence the specimens
are very frequently much crushed and distorted in shape. Typi-
cally the shell is broad and high, the beaks anterior in position
and the surface sculptured with narrow, wave-like undulations
which are best developed on the umbos.
Length 70, height 60. diameter 34 mm (Sousi Creek).
Uscari Stage: Sapote, C. Z. (Gabb)
Gatun Stage: Gatun, C. Z.
Nancy's Cay, Prov. of Bocas del Toro.
Upper Hone Creek, and tts tributary Soust
Creek.
Genus GAFRARIUM, Bolton
Section GOULDIA, C..B. Adams
Gafrarium limonensis, n. sp. Plate 32, figure 18
Shell small, subcircular, moderately convex, umbos inflated,
beaks anterior to the middle: surface finely sculptured with even,
concentric riblets, more or less cancellated by fine, radial threads
which are strongest on the anterior and posterior extremities,
while the concentric riblets predominate on the middle of the
shell; lunule distinct, narrowly lanceolate; interior of the shell
deep, the hinge of both valves with 3 cardinal teeth and an an-
terior lateral; ventral margin concentrically grooved but not
tangentially asin 7vansenella.
Wengthia.75, height 4:25, diameter) ot) the lett: valve
1.50 mm.
This small Gouldia, is fairly abundant in the Miocene coral
limestones of Port Limon, and is characterized by its small, con- —
vex shell and subcircular form. Gabb has described a small
Gouldia from the Pliocene beds of the Limon Peninsula, but that
234 BULLETIN 39 406
species is more depressed, less circular and more trigonal in out-
line. Gabb’s species may prove to be but a small form of the
recent West Indien G. cerzza C. B. Adams,
Gatun Stage; Port Limon.
Old Man Sam creek, 1 mile south of the beach.
Gafrarinm altum Dall, va. costaricensis, n. sp. Plate 32, figures I9, 22
ef. Gafrartum altum Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3,
pt. 6, p. 1249, pl. 57, fig. 5,
Shell small, high, rounded trigonal, moderately convex;
beaks small and pointed slightly forward, in young shell nearly
central but becoming with maturity slightly anterior; anterior
and posterior extremities nearly similiar and evenly rounded;
surface with fine, concentric lines, most distinct and even to-
wards the ventral margin, and with faint, radial strize on the
anterior and posterior slopes; lunule large, lanceolate, defined by
an impressed line; interior of shell deep, with a concentrically
grooved margin.
Length 4.85, height 4.75, diameter of right valve 1.50
min.
The Costa Rican shells differ from the typical altum, de-
scribed by Dall from the Oak Grove sands of Florida, in being
more trigonal in form and with higher beaks. The sculpture of
its surface is mainly concentric, the radials showing only as
very faint striae on the posterior and anterior extremities.
Gatun Stage: Middle creek.
Coll. 6, Estrella River.
Genus MACROCALLISTA, Meek
Section CHIONELLA, Cossmann
Macrocallista maculata Linnzeus Plate 31, figures 6, 7
Venus maculata Vinnzeus, 1758, Syst. Nat., ed. 10, p. 686, ed. 12, p.
rri2%
Cytherea dariena Conrad, 1857, Pacific R. R. Report, vol. 6, p. 72, pl.
Ip, 1 0egg yA
407 Costa R1icA MIocKNE—OLSSON 235
Callista maculata Gabb, 1881, Journ, Acad. Nat. Sci. Phila:, 2nd series,
VOLS) p41 372:
Macrocallista (Chionella) maculata Dall, 1903, Trans. Wagner Free
IS tNSCi..) voles. pt. (Gn pos) 256:
Macrocallista maculata Brown and Pilsbry, 1911, Proc, Acad. Nat. Sci.
Plila., vol. 63, p. 370.
The Macrocallista maculata is a common and widely dis-
tributed fossil in the Costa Rican Miocene and is readily recog-
nized by its depressed, porcellaneous and highly polished shell.
In the usual sandy phases of the Gatun, the species is quite typi-
eal, but becomes high and broad in the coralline or transitional
phase.
A variety cuneata has been proposed by Gabb to include
shells in which this posterior-dorsal slope is straight and not
humped as usual. We have seen no examples of this type.
The Macrocallista maculata isa common recent species of
the West Indies, appearing for the first time, in the lower Mio-
cene beds of the Chipola River, Florida.
Gatun Stage: Headwaters of Middle creek.
Rio Blanco.
Genus CALLOCARDIA, A. Adams
Subgenus AGRIOPOMA, Dall
Callocardia gatunensis Dall Plate 32, figure 1
Callocardia (Agriopoma) gatunensis Dall, 1903, Trans. Wagner Free
Inst. Sci., vol. 3, pt. 6, p. 1260, pl. 54, fig.1.
Callocardia (Agriopoma) gatunensis Brown and Pilsbry, 1911, Proc.
Acad. Nat. Sci. Phila., vol. 63, p. 370.
A convex, cordate shell very abundant in the Gatun beds
of the Canal Zone, especially in the quarries at Mt. Hope. It
is a rare species in Costa Rica, but generally larger, more trian-
gular in outline and somewhat coarsely sculptured. These shells
are nearest to Dall’s variety mudltzfilosa.
236 BULLETIN 39 408
Dall also records the species from the Miocene of Santo Do-
mingo and Jamaica.
Gabb’s C. sapotensts, from Sapote, Costa Rica, we have not
seen. It isa higher and more Dosinoid shell with nearly cen-
tral beaks. It is possibly but a variety of this species.
The following measurements are of Costa Rican examples:
Length 46, height 28, thickness 35 mm.
533 44, 16 mm (right valve)
Gatun Stage: Gatun and Mt. Hope, C. Z.
fiill No. 1, Banana River.
Soust Creek (Upper Fone creek).
Genus PITARIA, Roemer
Section LAMELLICONCHA, Dall
Pitaria circinata Born Plate 31 figures 3, 9
Venus civcinata Born, 1780, Test. Mus. Vind. p. 61, pl. 4, fig. 8.
Cytherea alternata Broderip, 1835, Proc. Zool. Soc., p. 45.
Cytherea Juncea Guppy, 1866, Quart. Journ. Geol. Soc. London, vol.
22, p. 682, pl. 26. fig. 13.
Chione circinata Gabb, 1873, Tarns. Amer. Phil. Soc., vol. 15, p. 250.
Cytherea juncea Guppy, 1876, Quart. Journ. Geol. Soc. London, vol,
32, p. 531.
Pitaria (Lamelliconcha) circinata Dall, 1903, Trans, Wagner Free
Inst. Sci., vol. 3, pt. 6, p. 1269.
Pitaria circinata Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila.
vol. 63, p. 370.
Pitarvia (Lamelliconcha) ciycinata Maury, 1917, Bull. Amer. Pal., vol.
5, P- 379, Pl. 37, fig. I.
The Pitaria circinata Bornis a common, recent Veneroid
along the Caribbean coast of Panama and Costa Rica. It occurs
also on the west or Pacific side but ina slightly different form,
as the variety a/ternata Broderip. This variety differs from the
typical civcinata by its larger, more convex shell and more dis-
tant concentric ribs.
409 Costa RicA MIocENE—OLSSON 237
The fossil shells from Costa Rica are generally more like the
West Coast alternata, but are smaller and with somewhat more
pointed beaks. The species is also found as fossil, in Santo Do-
mingo, Trinidad and Venezuela.
Gatun Stage: Gatun, C. Z.
Lowe EN Saray Green Canine
Pitaria Hillii Dall, var. musanica, n. var, Plate 31, figure 5
Shell elongate, moderately convex and rather solid; beaks
situated at the anterior %, with the anterior extremity widely
rounded but obtusely pointed posteriorly; lunule small, ridged
in the middle and defined by a faint line; posterior-dorsal area
broad and flat; surface of the shell sculptured with low, irregu-
lar, rounded, concentric ribs, which are persistent along the
posterior-dorsal slope but lacking elsewhere near,the ventral mar-
gin; interior of shell unknown.
Length 42, height 27, thickness (left valve) 10.50 mm.
This species has a solid shell like that of Wacrocallista. It
differs from the true /7z//7zz described by Dall from the Gatun of
the Canal Zone, by its more rude and irregular concentric sculp-
Blane:
The P. planivieta Guppy, from the Miocene of Jamaica and
Santo Domingo is a much shorter and more regularly sculptur-
ed shell.
Gatun Stage: Hill No. 2, Banana River, C. R.
Pitaria Guppyana Gabb Plate 31, figure I1
Caryatis Guppyana Gabb, 1881, Journ. Acad. Nat. Sci. Phila., 2nd
series, vol. 8, p. 373, pl. 47, fig. 73.
Shell small, ovate, moderately convex and _ porcellaneous;
dorsal side straight; ventral side gently convex; posterior ex-
tremity sub-truncate but wide, meeting the dorsal side nearly at
right angles; anterior extremity much narrower, produced and
rounded; lunule narrowly lanceolate, sculptured with the con-
238 BULLETIN 39 410
tinuation of the concentric ribs and separated from the shell disk
by an impressed line; surface of disk sculptured with rather wide,
low or appressed ribs formed between incised lines, but the
shell as a whole is porcellaneous and smooth; as the ribs pass
over the dorsal-posterior area towards the dorsal margin they
decrease in number through fusion; interior concealed in a hard
matrix.
Length 32, height 18, diameter (right valve) 5 mm.
A small species of the coralline limestones and marls of the
Limon Peninsula, from which it was described by Gabb. It
approaches the P. planivieta Guppy of the Miocene of Jamaica and
Santo Domingo but is smaller and more rectangular in outlines.
It is rather convex with a smooth and polished surface and with
regular, even, concentric ribs.
Gatun Stage: Port Limon.
Pitaria VanWinklez, n. sp. Plate 32, figure 2, 3
Shell elongate, depressed; anterior extremity broadly round-
ed, posterior more acute; lunule small, lanceolate and defined by
an impressed line; escutcheon long and very narrow; surface
sculptured with narrow, closely spaced and reflected ribs; these
ribs which number about 100 on the shell disk are even on the
center of the shell, but generally uneven on the anterior extrem-
ity; the ribs are continuous from the edge of the escutcheon to
the anterior dorsal margin; the rounded posterior dorsal slope
carries two small, faint rows along which the concentric ribs are
slightly higher and occasionally raised into small incipient
spines.
Length 41.5, height 32, diameter (left valve) 10 mm.
This species is quite similiar to the P. vosea Broderip and
Sowerby of the West Coast in its depressed shell and general
form. ‘The concentric ribs are very regular, even and continu-
ous across the entire surface of the shell. Where these ribs cross
the posterior-dorsal slope, they occasionally become raised into
4AXY Costa RicA M1ocENE—OLSSON 239
two rows of incipient spines. In P. rosea there is only one row
of incipient spines and the dorsal-posterior area is smooth.
This species is named for Miss Katherine Van Winkle of the
Paleontological Department of Cornell University and who is at
present engaged on a Monograph of the American species of
Veneride.
Gatun Stage: Hill No. 3. Banana River, C. R.
Pitaria boucaryensis, n. sp. Plate 31, figure 10
Shell rounded quadrate, thin and moderately convex; pos-
terior and anterior extremities rounded, umbos rather large;
lunule small and feebly defined by a faint, impressed line; sur-
face of disk sculptured with about 4o even, distant, concentric
lamellae; interspaces generally sinooth or irregularly sculptured
with growth lines; interior of shell concealed.
Length 31, height 22, diameter (right valve) 10 mm.
As the single type specimen is fragmentary and imbedded
in a hard sandstone, the above measurements are but approxi-
mate. It isa very distinctive species, differing from the other
fossil Costa Rican Pitarias by its more delicate and convex shell
and more distant, even, concentric lamellee.
Gatun Stage: Boucary creek, C. R.
Genus ANTIGONA, Schumacher
Antigona multicostata Sowerby Plate 30, figure I
Venus multicostata Sowerby, 1835, Proc. Zool. Soc., p. 22.
Venus multicostata Reeve, 1863, Conch. Icon., vol. 11, pl. 3, fig. 9.
Cytherea ( Cytherea) mutlticostata Dall, 1902, Proc. U.S. Nat. Mus.,
vol. 26, p. 390.
This is a large species living on the south or Pacific side of
the Isthmus. A perfect specimen was collected from the Gatun
beds of Water Cay, identical in every respect with recent ex-
amples from the Bay of Panama. It is another instance among
the many which we have had occasion to note in the course of
240 BULLETIN 39 412
the present study of the occurrence of West Coast species in the
Caribbean Miocene, proving conclusively from a purely paleon-
tological standpoint the rather late separation of the Atlantic
and Pacific through the final uplift of the isthmian lands.
The recent A. Lister? Gray is a smaller, more elongate
species and with finer and sharper concentric lamelle.
Length 107 mm, height 102 mm, diameter 69 mm.
Gatun Stage: Water Cay.
Antigona tarquina Dall Plate 30, figure Io
Venus magnifica Gabb, 1673, Trans. Amer, Phil. Soc., vol. 15, p. 249.
Not of Sowerby, 1853.
Cytherea tarquina Dall, 1900, Trans. Wagner Free Inst., vol. 3, pt. 5,
pl. 38, figs. 2, 2a; vol. 3, pt. 6, p. 1274.
Antigona tarquina Dall, 1915, Bull. go, U. S. Nat. Mus., p. 147.
Antigona tarquina Maury, 1917, Bull. Amer. Pal., vol. 5, p. 381. pl.
37, fig. 4.
A small shell identical or closely related to A. tarquina
Dall was collected from Zone 3 of Saury Creek. It is similiar
to Dr. Maury’s specimen of 4. farguina from Santo Domingo.
Typical A. targuina Dall is a common fossil of the Orthau-
lax pugnax fauna of the Tampa Silex beds of Florida. It was
also collected by Professor Gabb and Dr. Maury from the Mio-
cene of Santo Domingo.
Gatun Stage: Zone E, Saury creek, near Cahutta, C. R.
Section VENTRICOLA, Roemer
Antigona rugosa Gmeiln Plate 30, figure 4
Venus rugosa Gmelin, 1792, Syst. Nat., p. 3276.
Venus rugosa Reeve, 1863, Conch. Icon., vol. 11, pl. 7, fig. 23.
Several specimens from the coralline limestones of Port
Limon. ‘They are practically identical with recent examples of
A. rugosa from the West Indies. This species is distributed
throughout the West Indies but is also found living in the Gulf
413 Costa Rica MrocENE—OLSSON 241
of California. This distribution on both sides of the Isthmus is
in harmony with its occurrence as a Miocene fossil.
Gatun Stage: Port Limon.
Antigona Blandiana Guppy Plate 30, figures 5, 6
Venus Blandiana Guppy, 1874, Geol. Mag., London, Decade ii, vol. 1,
P4366, ply 17.) fs. 6:
Cytherea (Ventricola\ Blandiana Dall, 1903, Trans. Wagner Free Inst.
Sci Voli 35 pt. 6) p. 1277.
Antigona (Ventricola) Blandiana Maury, 1917, Bull. Amer. Pal., vol.
5, P- 381, pl. 37, fig. 5.
This species was described by Guppy from the Bowden beds
of Jamaica but also collected by the Maury expedition in the
Miocene of Santo Domingo. Dali also records it from the lower
Miocene of the Chipola beds and from Curacoa, Dutch West
Indies. It is asmall species of nearly circular outlines and
sculptured with distant primary lamelle and finer secondary
threads in the interspaces.
Our Costa Rican shells consist of two specimens, one from
the coral limestones near Port Limon, the other from the lower
Gatun of Comadre Creek near Cahuita. They differ from Dr.
Maury’s specimens from Santo Domingo in having the primary
set of lamella more distantly spaced and their interspaces more
closely striated with the secondary threads.
Length 30, height 24, diameter 9g mm (left valves) Com-
adre.
Length 24, height 23, diameter 8.5 mm (left valve) Port
Limon.
Gatun Stage: Comadre creek, near Cahuita.
Port Limon.
Antigona Harrisiana, n. sp. Plate 30, figure 3
Shell small, convex and nearly circular in outline; umbos
full with the beaks situated a short distance in front of the mid-
dle; lunule small but broadly cordate and sculptured with the
242 BULLETIN 39 | 414
continuation of the concentric ribs and lamella; escutcheon lan-
ceolate, bordered by an angled ridge from the shell disk; disk
sculptured with about 28 even, concentric ribs which appear
as close undulations of the shell surface; the ribs and inter-
spaces are in addition sculptured with fine lamelle-like threads,
each finely crenulated or radially striated; these lamellee vary in
size, those of the interspaces or troughs between the ribs are fine
and 5 in number, with 2 large ones forming the tops of the ribs
themselves, so that the ribs appear as if mesially divided; the
radially striated character of the ribs and lamellz is much strong -
er on the anterior and posterior submargins; interior of the shell
deep but concealed by matrix in the type specimens.
Length 21.5, height 19.5, diameter 6.25 mm _ (right
valve).
This elegant species occurs in the coralline phase of the
Gatun formation in Costa Rica. Its relations are with the 4.
Blandiana Guppy and the recent A. strigillina Dall, but differs
from both in its more elegant sculpture. This sculpture consists
of close, regular, wave-like undulations or ribs, the crests of
which carry 2, low lamine, the troughs or interspaces, 5 or 6
finer threads. ‘These fine thread-like laminz are delicately etch-
ed with fine radial striations.
It is named for Professor G. D. Harris of the Paleontologi-
cal Department of Cornell University.
Gatun Stage: Port Limon.
Genus CYCLINELLA, Dail
Cyclinella beteyensis, n. sp. Plate 31, figure 2
Shell thin, Dosinoid or sub-circular in form, the margins of
the shell nearly a perfect circle; slightly convex; beaks small
but distinct, situated at the anterior % of the shell ; sculpture
with very fine, concentric threads, which are a little elevated and
strongest on the anterior end, nearly smooth on the middle; lu-
nule small, lanceolate and defined by a small, incised line; hinge
45 Costa R1icA MiockNE—OLSSON 243
normal, the left valve with 3, narrow; cardinal teeth and a wide
ligament.
Height 39, length 38? diameter 5 mm left valve.
Represented by a single, imperfect specimen from the Gatun
of Rio Betey. It is but slightly convex, and in form approach-
es very closely a species in the Newcomb collection from the
Bay of Fonseca, which is probably the C. Avoyeri Philippi. The
fossil shell is more perfectly circular in form.
Gatun Stage: Rio Betey.
Cyclineila subquadrata Hanley, var. quitana, n. sp. Plate 31, fig. 8
Shell small, thin, convex and subcircular in form; beaks
small but distinct, with inflated umbo and small beak situated
at the anterior %; the anterior end is narrow and slightly pro-
duced, while the posterior end is wide and with the hinge-margin
is subquadrate in form; the surface is sculptured with fine, ir-
regular, concentric growth-lines, which are strongest and more
regular on the posterior dorsal submargin; the interior 1s con-
cealed.
Height 28, length 31, diameter of the right valve 7 mm.
This is a smaller and more convex species than the beteyen-
sts and like that species, it seems to find its recent analogue on
the Pacific side, in this case the C. swbqguadrata Hanley. On the
Pearl Islands in the Bay of Panama I collected a few small
valves which are probably the C. subguadrata Hanley, but they
are not so produced anteriorly as is shown in Reeve’s figure.
These Pearl Island shells are very similiar to the Costa Rican
fossil, differing mainly in their more inflated umbos and in
slight difference in form.
From the Gatun beds of the Canal Zone, Dall has described
C. gatunensis, but that species is very distinct from the two
Costa Rican Cyclinellas.
Gatun Stage; Outtana creek.
244 BULLETIN 39 416
Genus CHIONE, Megerle von Miihlfeld
Chione Rowleei, n. sp. Plate 30, figure 2
Shell rather large, convex, ovate; lunule broadly cordate
and concentrically sculptured; escutcheon long, narrow, smooth
and defined by a small cord-like ridge; surface of the disk with
about 30 slightly elevated concentric ribs which on the umbos
are thin and lamellar but lower on the shell disk are wide and
trianglar at the base, but with a thin zigzag or fluted lamellar
ridge on top; the interspaces, much wider on the umbos is sculp-
tured with even rib-like cords which also flute in harmony the
dorsal face of the concentric ribs; the ventral face of the ribs are
finely striated by radial incised lines which form cord-like ridges
along % as wide as the radial cord of the interspaces; interior of
the shell deep, with a small pallial sinus and subequal adductor
scars; basal margin finely crenulated as well as the lunular mar-
gin and the extreme posterior extremity.
Length 42, height 37.5, thickness (right valve) 17 mm.
This large, elegant Chione is quite common in the Gatun
beds of Costa Rica. It also occurs in the lower Gatun at Gatun,
usually in the form of casts. It recalls in a general way, the re-
cent West Coast C. amathusia Philippi, but differs in its heavier
shell, higher and fuller umbos and difference in details of its
spiral and radial sculpture. In the Costa Rican shell, the radial
cords are simple, relating it more closely with the small C.
Woodwardi Guppy of the Miocene of Jamica and Santo Domin-
go, while in C. amathusia, they are double and alternating.
The C. tegulum Brown and Pilsbry we have not seen. It
is a smaller shell, related to the C. Woodwardi. In C. tegulum
the concentric lamellee are much closer and the wide cordate
lunule is sculptured only with radial lamellee.
It is named for Professor W. W. Rowlee, who collected a
few specimens from the Banana River in the course of botanical
investigation of the Balsa tree (Genus Ochroma), a very light
wood used in the construction of life-buoys, etc.
417 Costa RicA Mi1ocENE—OLSSON 245
Gatun Stage: Gatun, C. Z.
Banana River.
Coll. 6, Estrella River.
Chione costaricensis, n. sp. Plate 32, figure 4
Shell of moderate size, convex, ovate-trigonal; the dorsal
portion of the type specimen is lacking; the surface is sculptured
with fine, subregular, concentric lamella spaced on the center of
the shell disk about .75 mm apart; the interspaces and the ven-
tral surfaces of the lamellee are marked with regular, incised lines
forming radial bands of an average width of .40 mm; a faint rad-
ial sinus extends from the beaks to the posterior; ventral margin
of the shell is slightly arcuate at its posterior end; interior of
ventral margin finely crenulated.
Length 34, height 29, diameter of the right valve 10
mm.
This shell will be distinguished from the other Costa Rican
Chiones by its more crowded, concentric, lamelle and regular,
radial stria. The striz occur on the ventral faces of the lamel-
lze and on their interspaces.
Gatun Stage: Hill No. 3, Banana River.
Section LIROPHORA, Conrad
Chione mactropsis Conrad Plate 30, figures 7, 8
Gratelupia? mactropsis Conrad, 1856, Pacific R. R. Reports, vol. 5, p.
328, pl. 6, fig. 54.
Chione (Lirophora) mactropsis Dall, 1903, Trans. Wagner Free Inst.
Sci., vol. 3, pt. 6, p. 1294.
Chione (Lirophora) mactropsis Brown and Pilsbry, Ig911, Proc. Acad.
Nat. Sci. Phila., vol. 63, p, 369.
Chione (Lirophora) ulocyma Brown and Pilsbry, to11, /dem, p. 369.
Not of Dell. 1903.
Until the construction work on the Panama Canal had com-
menced in earnest, the abundant and finely preserved fossils of
246 BULLETIN 39 4i8
the Canal Zone, so easily obtained at the present time, were not
available, and the only fossil remains collected by the early trav-
vellers were generally in the form of poorly preserved casts. In
1856 there appeared in the Pacific Rail Road Reports, the de-
scription of an internal cast as Gratelupia? mactropsts, together
with scarsely recognizable Turritella gatunensis and altilira.
These specimens obtained by Mr. W. P. Blake from the
Isthmus are the first noticed from the fossiliferous rocks in
Panama and Central America.
The Livophora mactropsis described by Conrad from an in-
ternal cast as Gratelupia ?, is one of the most abundant fossils in
the Gatun beds of the Canal Zone, associated with the small-
er and more finely sculptured holocyma Brown and Pilsbry. JL.
mactropsis is a species of variable size, but moderately convex,
longer posteriorly and usually with the basal margin more or
less arcuated about the posterior extremity. The sculpture con-
sists of irregular, more or less confluent concentric lamelle,
which are overrun by fine radial lines. These radial lines may
remain strong or become obsolete as usual with specimens from
the Canal Zone. A narrow foliaceous band is developed on each
side of the escutcheon and another about the anterior extremity
below the lunule. The Costa Rican examples are often very
much larger than those of the Canal Zone, the concentric lamel-
le are finer and less confluent and with more persistent radial
lines. The following measurements will illustrate the range in
size:
Length 30, heigth 23, thickness 15 mm, Central Zone.
ce ce
37) 28, 19
41.5, 22) 9.5 mm (right valve)
Old Man Sam Creek, C. R.
Bilas) Rhigyen 21.5 mm, Sousi, C. R.
Gatun Stage: Gatun, and Mt. Hope, C. Z.
Old Man Sam creek, C. R.
Soust creek (Upper Fone creck)
Banana River.
419 Costa Rica MrocENE—OLSSON 247
Chione holocyma Brown and Pilsbry Plate 30, figure 9
Chione ulocyma Dall, variety holocyma Brown and Pilsbry, 1911, Proc.
Acad. Nat. Sci. Phila., vol. 63, p. 369.
Associated with mactropsts, in the Central Zone is a more
convex and finely sculptured shell, allied to the wlocyma Dall of
of the Floridian Miocene. It is the /olocyma of Brown and Pils-
bry. We have seen no typical wlocyma from Panama.
From mactropsis, the holocyma will be distinguished by its
generally smaller, more convex and more circular shell. Its
concentric lamellae are more numerous and less confluent and
with fine but persistent radial strice. It is generally best distin-
guished by its smaller and less clearly differentated foliaceous
area on its anterior extremity, which in mactropsis is as wide
an area as that along its posterior dorsal margin.
Heigth 29, length 36.5, thickness 19 mm.
Gan Stage Mt. Flope, C- Z.
Chione hotelensis, n. sp. Plate 32, figure 7
Shell small, subtrigonal, heavy, but slightly convex; sculp-
ture of about 15, thick, rounded and evenly spaced concentric
lamellee; on the posterior-dorsal slope, the ends of the lamellce
become attenuated, thin and somewhat appressed as in glyptocy-
ma Dall of the Flordian Miocene; the lamellze themselves are
smooth and rounded but with their interspaces finely radially in-
cised by small lines; escutcheon rather large, smooth; lunule
small, cordate and smooth except for the growth lines,
Length 23, height 19, thickness 12.5 mm.
This elegant little species is distantly related to the C. gly-
ptocyma Dall from the Oak Grove sands of Florida, but isa
smaller and more compact shell. The radial striations commence
on the ventral surface of the lamellze and are continued across
the interspace to the base of the next rib.
All of our specimens were collected in a hard sandstone in
the lower Gatun of Hotel Creek near Old Harbor, Costa Rica.
248 BULLETIN 39 420
Uscart Stage: Hotel creek, C. R.
Chione chiriquiensis, u. sp. Plate 32, figure 9, Ic
Shell small, solid, depressed; lunule rather large, cordate,
smooth; escutcheon large, smooth and defined by an angled
ridge; surface of the disk sculptured with about 8, large, irregu-
lar ribs, like those of Chione latilirata; these ribs do not extend
to the edge of the escutcheon, but commence a short distance in
front leaving a narrow band which is smooth and sculptureless;
the ribs are of variable size, flat-topped and smooth; they are
rounded on their ventral side, but shelving or recurved back-
wards on the dorsal face; interspaces smooth or only sculptured
with fine growth-lines; interior of shell of moderate depth, with
a small pallial sinus and with a slightly larger posterior adduct-
or scar; basal margin finely crenulated.
Length 13.75, height 12.50, diameter 9.50 mm.
The few specimens which we have from Water Cay, are
probably immature, but they possess such distinctive characters
that the species cannot be mistaken.
The heavy, irregular concentric ribs recall certain varieties
of Chione latilirata Conrad. ‘They differ in being abruptly cut
off or flattened a short distance behind the dorsal-posterior angle
leaving a band asin C. paphia Linnzus, but smooth and sculp-
tureless. The interspaces are smooth,
Gatun Stage: Water Cay.
Section TIMOCLEA, Brown
Chione grus Holmes, var. Plate 26, figure 21
Tapes grus Holmes, 1858, Post-Pliocene Fossils of S. Car., p. 37, pl. 7
fig. 5.
Chione ( Timoclea) grus Dall, 1903, Trans. Wagner Free Inst. Sci.,
vol. 3, pt. 6, p. 1299.
We have but a single and not entirely perfect specimen of a
Timoclea closely related to C. grus Holmes. It is somewhat
42t Costa Rica MI1ocENE—OLSSON 249
more convex than typical C. grvus from the Carolinas and more
finely sculptured. More material is needed to show whether or
not these charaters are constant. The C. gvus appears asa
fossil in the Upper Chesapeake Miocene of the eastern United
States and is recent from Cape Hatteras to Yucatan.
Gatun Stage: Coll. 4, Red Cliff creek.
Genus TELLINA, (Linné) Lamarck
Teilina rovala, n. sp. Plate 26, figures 7, 8
Shell elongate, slender, subequilateral with the beaks situ-
ated near the center of the shell; anterior side long, with a
straight dorsal margin and narrowly rounded at its extremity;
posterior side much narrowed and rostrate, the left valve with 2
radial bands forming the rostrum, the dorsal one a fold, the oth-
era wide sinus; ventral margin rounded anteriorly, nearly
straight about the middle and a wide sinus posteriorly; the sur-
face is sculptured with even, concentric lamelle; this sculpture
on the rostrum is heavier and on crossing the fold and sinus be-
come reversed in position.
Length 29, height 12, semi-diameter 2.5 mm.
36? T4 3 mm.
We have two specimens of this typical 7e//ima from Water
Cay. The larger specimen is fragmentary but probably measur-
ed when complete about 36 mmin length. The species recalls
the recent West Indian 7. zxzterrupta Wood (7. antonzz Philippi)
in its form and sculpture but lacks the finer radiating strie.
Gatun Stage: Water Cay.
Subgenus ARCOPAGIA, (Teach)
Section MERISCA, Dall
Tellina crystallina Wood Plate 26, figure 10
Tellina crystallina Wood, 1815, General Conchology.
250 BULLETIN 39 422
Tellina (Merisca) crystallina Dall, 1901, Proc. U. S. Nat. Mus., vol.
Pew oye ener ols 7s) never.) 3(6)
Tellina (Merisca) crystallina Maury, 1917, Bull. Amer. Pal., vol. 5, p.
387, pl. 38, fig. 4.
This Zel/ina, the type of the section JZerisca, is recognized
by its thin, depressed shell, strongly sculptured with distant, but
regular, raised, concentric lamellee. It is very rare as a fossil,
our collection containing but a single specimen from the Gatun
beds of the Banana River. It has previously been recorded by
Maury as a Miocene fossil in Santo Domingo.
In the recent fauna, Tellina crystallina is distributed along
both the Atlantic and Pacific coasts of Central America. I have
collected a few specimens from the beaches of Bocas Island and
more abundantly at Bucaru, Province of Los Santos, on the
Pacific side of Panama.
Gatun Stage: Hill No. 2. Banana River.
Tellina nasua, n. sp. Plate 26, figure 17
Shell sub-rhomboid, slightly convex, thin; beaks small, in-
conspicuous, posterior to the middle; the anterior end longer,
the dorsal-anterior dorsal margin descending, the extremity
rounded into the gentle curve of the base; (the posterior side
broken in the type specimen); the greatest convexity of the shell
lies along the broad, umbonal slope anterior to the beaks; the
surface is finely sculptured with regular, sharp, concentric
ridges, separated by narrow, band-like interspaces; interior of
the right valve with a large pallial sinus which extends to the
anterior 1% of the shell; hinge appearing normal for the section,
but partly broken away, but showing a large anterior lateral
tooth.
Length 20? Height 15.25, diameter of the right valve
3.00 mm.
Our specimen if fragmentary, the posterior portion of the
shell being broken away. There is however little doubt that
the species is a true JZerisca, allied to 7. egutstriata Say and
423 Costa RicA M1ocENE—OLSSON 251
TZ. lintea Conrad. It differs from eguzstriata in its greater dor-
sal anterior slope and finer sculpture, and from 7. /zztea of the
recent fauna, in its somewhat greater proportional height and
greater convexity.
Gatun Stage: Coll. 4, Red Cliff creek.
Section PHYLLODINA, Dall
Tellina lepidota Dall Plate 26, figures 4, 5
Tellina (Phyllodina) lepidota Dall, 1900, Trans. Wagner Free Inst.
El mVOla 35 Pts 5, .p: LO22. pliv AO whoa les
Dall’s type of this species is based on a fragmentary right
valve from Gatun. We have several complete and large speci-
mens from Water Cay, where the species is fairly common. The
right valve is flat and depressed, the left more convex. Both
valves are nearly similarly sculptured with 2 or 3 wide undul-
ations of the beaks, which are soon replaced by even, sharp, con-
centric lamellee, of which 3 or 4 are higher and more lamellose
on the dorsal margin.
Length 24, height 12.5, diameter of the left valve 2.75
mim.
Length 21.5, height 11.00, diameter of the right valve
eA sonben,
Gatun Stage: Water Cay.
Section EURYTELLINA, Fischer
Tellina costaricana, n. sp. Plate 26, figures 6, 9
Shell oblong, depressed; beaks small and inconspicuous
about the middle of the shell so that the anterior and posterior
ends are of nearly the same length; posterior end widely round-
ed the anterior but slightly narrowed and less _ round-
ed at its extremity; anterior and posterior dorsal margins nearly
straight; surface of the disk finely sculptured with even, con-
252 BULLETIN 39 424
centric lines which are slightly lamellose on the posterior-dorsal
margin; interior concealed.
Length 32, height 19, semi-diameter 2 mm.
This species is related to the recent West Indian 7. angulosa
Gmelin and its Pacific analogue, the 7. rubescens Hanley. The
fossil shell differs by its more central beaks and sharper concen-
tric sculpture, which is continued across the entire surface of
the disk.
Gatun Stage: Hill No. 3, Banana River.
Tellina dariena Conrad Plate 26, figure 3
Tellina dariena Conrad, 1857, Pacific R. R. Reports, vol. 5, p. 328,
pl. 6, fig. 53.
Tellina semilevis Gabb, 1861, Proc. Acad. Nat. Sci. Phila., vol. 12, p.
567.
Tellina Dariena Gabb, 1881, Journ. Acad. Nat. Sci. Pnila., vol. 8, p.
343, pl. 44 fig. 13.
Tellina Rowlandi Toula, 1908, Jahrb. der K-K. Geol. Reichsanstalt
vol. 58, p. 728, pl. 28, fig. rr.
Tellina Rowlandi Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 368.
Conrad’s figure of 7e/lina dariena is very poor but probably
represents this species, the most abundant 7Ze//zma at Gatun.
Conrad’s figure shows a subtrigonal shell, with a produced and
truncated posterior extremity. In 1861, Gabb described Jellina
semtlevts together with Arca chiriguiensts from the Gatun beds
of the Chiriqui Lagoon. This species Gabb however later con-
sidered synonymous with the earlier 7e//ina dariena of Conrad.
Toula’s TZellina Rowlandi is certainly identical with Gabb’s
Tellina semilevis as figured by Gabb.
Shell but slightly convex, subequilateral with the beaks sit-
uated a short distance posterior to the middle of the shell; anter-
ior and posterior dorsal slopes straight and evenly descending;
posterior end rounded and subtruncate, the anterior rounded ;
beaks low; surface striated with very fine concentric lines and
425 Costa R1icA MIocENE—OLSSON 253
the shell may sometimes appear nearly smooth and polished
especially on the umbonal areas; hinge of the left valve with a
bifid anterior cardinal, a small posterior and 2 laterals; the an-
terior adductor is bordered on the posterior side by a thickened
internal ray; pallial sinus long, ascending above in the middle
of the shell and touching the anterior adductor scar.
Length 34, height I9.5, semi-diameter 3.50 mm.
Gatun Stage: Gatun, C. Z.
Water Cay.
Chiriqui Lagoon and Oronli creck (Gabo)
Subgenus ANGULUS, Mergerle
Section ANGULUS, s. s.
Tellina umbra Dall Plate 26 figures 12, 15
Tellina (Angulus) umbra Dall, 1900, Trans. Wagner Free Inst. Seine
vol. 3, pt. 5, p. 1033, pl. 46, fig. 13.
The Costa Rican specimen consists of a small individual
with both valves. It is somewhat smaller than typical wmora,
from the Upper Chesapeake Miocene of the Carolinas, but agrees
in form and in the fine surfrce sculpturing.
The Tellina umbra is a common fossil in the Upper Chesa-
peake Miocene of the eastern United States and in the Pliocene
of Florida, It is closely allied tothe recent Tellima sybaritica
Dall of the West Indies, differing most importantly in its less an-
gular posterior extremity.
The Costa Rica specimens measures:
Length 8.00, height 4.15, diameter 2.25 mm.
Gatun Stage: Coll. 4, Fast Grape Point creek.
Section SCISSULA, Dall
Tellina cercadica Maury
Tellina (Scissula) cercadica Maury, 1917, Bull. Amer. Pal., vol. 5, p.
388, pl. 38, fig. 9.
254 BULLETIN 39 426
The Costa Rican specimens are fragmentary but there can be
little doubt of their identity with the TZel/ina cercadica Maury
from the Miocene of Santo Domingo.
The shell is thin and strongly depressed. ‘The surface is
polished and obliquely striated with fine lines. It is related to
the recent Pacific 7ellina delicatula Deshayes.
Gatun Stage: Coll. G, Saury creek.
Estrella River.
Genus TELLIDORA, Moerch
Tellidora cristata Recluz Plate 26, figurce 1, 2
Lucina cristata Recluz, 1742, Revue Cuvier., p. 270.
Tellidova cristata Dall, 1900, Trans. Wagner Free Inst. Sci., vol. 3,
pt-.4) \p) 1037.
A recent species of the West Indies and previously knowa
as a Pliocene and Pleistocene fossil from Carolinas. The fossil
Costa Rican shells are frequently larger than the recent form
and with the dorsal margins more nearly in line. It is not rare
in the form of impressions but the perfect shell is seldom found.
Gatun Stage: Hills ra, 2 and 3, Banana River.
Genus STRIGILLA, Turton
Strigilla pisiformis Linné
Tellina pisiformis Linné, 1758, Syst. Nat., ed. 10, p. 677.
Strigilla pisiformis Dall, 1900, Trans. Wagner Free Inst. Sci., vol, 3,
pt. 5, p. 1038.
Strigilla pisiformis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 389, pl.
39, fig. 6.
This species is rare asa fossil in Costa Rica. It is very
common asa recent shell on the sandy beaches along the en-
tire north coast of both Panama and Costa Rica, where it occurs
with Donax denticulata Linné and afew other species. It is
427 CosTA RicA MiocENE—OLSSON 255
also found as a Miocene fossil in the Bowden beds of Jamaica
and in Santo Domingo.
Gatun Stage: Middle creek.
Strigilla musanica, n. sp. Plate 26, figure 18
Shell subtrigonal, moderately convex, subequilateral; the
umbos are broad and wide, situated about the middle of the shell,
with the greatest convexity of the shell lying along the anterior-
umbonal slope; the anterior end is broad and rounded, the pos-
terior slightly narrow, but not produced; the sculpture is divid-
ed between three areas, the posterior, central and anterior, each
of which is set off from the adjacent one, by a marked change in
sculpture along a well-defined line; on the anterior area, it con-
sists of a concentric series of wave-like incised lines, the crest of
the wave lying near the posterior side and directed as usual dor-
sally; on the central area, the lines are vertically directed, com-
mencing on the lines bordering the posterior side; on the poster-
ior area, the lines commence on the anterior side and pass verti-
cally, forming therefore with the middle lines very acute an-
gles; the interior is concealed in our specimen.
Length 13.00, height 11.50, diameter of the right valve
3.00 mm.
Differs from Strigzlla carnaria Linné of the recent West In-
dian fauna, by being more inequilateral, more convex and in de-
tails of its sculpture which is sharply divided between three areas
as described. ‘The interior of the type specimen is concealed, so
that the features of the pallial line and sinus cannot be deter-
mined.
Gatun Stage: Hill No. 3, Banana River.
Genus METIS, H. and A. Adams
Metis barba, n. sp. Plate 29, figures 3, 4
Shell subquadrate, inequilateral, with the beaks situated at
the anterior two-thirds of the total length of the shell; anterior
256 BULLETIN 39 428
end longer with a sloping dorsal margin and rounded anterior
extremity; posterior side shorter, with a dorsal margin sloping
more rapiding to the wide and flexed posterior end; the left valve
is convex but depressed along the narrow posterior submargin or
wing; the right valve is less convex with a broad depressed area
extending from the beaks to the ventral margin; this depressed
zone is sharply separated from the posterior-dorsal slope or wing
by a strong angle or flexure which corresponds to the sinus in
the middle of the wing on the left valve; surface finely striated
with concentric growth-lines.
Length 48, height 37, diameter 19 mm.
Pilsbry and Johnson have described without figuring,
two species of Metis from the Miocene of Santo Domingo (the
M. efferta and postrema). ‘These species, they compare with the
M. chipolana Dall, from the Chipola beds of Florida and men-
tion in their description the nearly central location of the beaks.
In the Costa Rican shell the beaks are at the posterior one-third
of the shell. It is very distinct from the (7. chipolana Dall. JZ.
trinttaria Dall from ‘Trinidad and Porto Rico differs in its more
central beaks, fuller umbos and has the right valve much less
flexed posteriorly.
Gatun Stage: Betey creek.
Genus MACOMA, Leach
Macoma costaricana, n. sp. Plate 26, figures 14, 16
Shell broadly ovate, thin and slightly inflated; dorsal and
ventral margin straight and parallel; the posterior dorsal side
sloping to the rounded ventral extremity; anterior extremity
rounded; the posterior side is very slightly flexed; surface
smooth except for very fine, incremental growth lines; interior
of shell concealed.
Length 29, height 19, semi-diameter 4.5 mm.
The reference of this species is Macoma is doubtful as the
429 Costa Rica M1ocENE—OLSSON 257
hinge is completely concealed in all our specimens. It has the
form of the JZ. Conradi Dall, from the Chesapeake Miocene of
United States but is higher and differs in many other respects.
It is shorter and higher than 4. gatunensis Toula with which it
miay occur.
Gatun Stage; Hill No. 3, Banana River.
kio Betey.
Subgenus PSAMMACOMA, Dall
Macoma gatunensis Toula Plate 26, figure 13
Macoma ( Tellina) gatunensis Toula, 1908, Jahrb. der K-K Geol. Reich-
sanstalt, vol. 58, p. 729, textfigure Io.
Tellina gatunensis Brown and Pilsbry, IgtI, Proc. Acad. Nat. Sci.
Phila., vol. 63, p. 368.
This shell is extremely similiar to the recent M. elongata
Hanley of the Pacific coast of Panama and I have failed to find
any character which will distinguish the fossil shells from the
recent ones. Our shells were all collected loose near Hill 3, of
the Banana River and were probably derived from beds low in
the Gatun formation.
Gatun Stage: loose, near Hill 3, Banana River (Terry).
Macoma panamensis Dall, var. canatis, n. var, Plate 26, figure II
Cf. Macoma (Psammacoma) panamensis Dall, 1900, Proc. U. S. Nat,
Mus., vol. 23, p. 324, pl. 4, fig. 3.
The Vacoma panamensis Dall was described from a left
valve dredged from 33 fathoms of water in Panama Bay. ‘The
fossil shell, also a left valve from the Gatun of Mt. Hope, differs
most importantly from Dall’s figure in being somewhat higher.
The shell is thin and extremely fragile, with the surface simply
scultured with the fine growth lines. The interior is concealed
so that the characters of the pallial sinus cannot be seen. The
specimens measures as follows:
258 BULLETIN 39 430
Length 30.50, height 15.00, diameter of the left valve
3.00 mm.
Gatun, Stage Mi Fope. (Gi Zh
Genus SEMELE, Schumacher
Semele levis Sowerby, var. costaricensis, u. var. Plate 29 figure 1
Cf. Amphidesma levis Sowerby, in Reeves, 1853, Conch. Icon., pl. 7,
fig. 50.
Shell large, subelliptical, but slightly convex; beaks very
nearly central, the posterior side wide and well rounded, the an-
terior somewhat narrowed, contracted and. slightly the longer;
the surface is nearly smooth and porcellaneous, faintly marked
on the anterior and posterior extremities by fine, concentric
lines.
Length 72, height 65.50, semi-diameter 8 mm.
This large Semele is quite common in the Lower Gatun beds
of the Banana River. It is closely related to the recent Pacific
S. levis Sowerby, but differs in being slightly longer and more
evenly rounded at its posterior extremity.
Gatun Stage: Hill No. 3, Banana River.
fiiull ra, Banana River.
Semele Sayi Toula Plate 29, figure 8
Semele Sayt Toula, 1908, Jahrb. der K-K Geol. Reichsanstalt, Wien,
vol. 58, p. 730, pl. 28, fig. 17.
Semele Sayt Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila.
vol. 63, p. 368. ;
Shell convex, rather thin, with the umbos nearly central.
The anterior side is somewhat narrow or contracted, and round-
ed at its extremity. The surface is sculptured with fine, hair-
like, concentric threads. The species was described by Toula
from the Canal Zone. Our record from Costa Rica is based ona
single, imperfect specimen from Sousi creek, where it occurs as-
sociated with Clementia, Chione mactropsts and Turritella altilira
431 Costa R1rcA MIocENKE—OLSSON 259
variety costaricenszs in the lower part of the Gatun formation.
Gatun Stage: Sousi creek.
Semele Claytoni Maury Plate 29, figure 2
Semele Claytont, Maury, 1917, Bull. Amer. Pal., vol. 5, p. 227, pl. 35,
fig. 9.
The S. Claytont was described by Dr. Maury from the Cer-
cado formation, Santo Domingo of Lower Miocene age. Itisa
large species, reaching a length of 66 mm or more and charact-
eristically sculptured with even, concentric lamellz like the re-
lated species from the Tertiaries of eastern United States as chipo-
lana Dall, Leana Dall and perlamellosa Heilprin. From these,
the S. Clayfonz is distinguished in having its radial strize nearly
obsolete. Our specimen from Costa Rica is fragmentary but
probably measured when perfect 38 by 30 mm.
Uscart Stage: Hotel creek.
Genus ABRA, (Leach) Lamarck
Abra triangulata Dall Plate 29, figure 10
Abra triangulata Dall, 1900, Trans. Wagner Free Inst., Sci., vol. 3,
pt. 5, p. 997, pl. 49, fig. 4.
A small, rotund or trigonal species abundant in the shale
beds of Bocas Island. ‘The valves are nearly equilateral, the an-
terior end slightly the larger and inflated. Surface smooth and
shining. The type was described by Dall from the Bowden beds
of Jamaica. Our specimens are a little larger, measuring:
Wength 7.25) height |\6450, diameter of) the) left) valve
1.75 mim.
Gatun Stage: Bocas del Toro.
260 BULLETIN 39 A432
Genus PSAMMOSOLEN, Risso
Psammosolen gatunensis Toula, var. costaricensis, n. var.
Plate 29, figure 7
Solecurtus gatunensis Toula, 1908, Jahrb. der K-K. Geol. Reichsan-
stalt, vol: 58, p. 732, pl. 28, fig. 12.
Solecurtus gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci.
Phila., vol. 58, p. 372.
Shell rather large, elongate, convex, the dorsal and ventral
margins straight and parallel; the low, inconspicuous beaks sit-
uated at the anterior 4, the anterior end rounded, the posterior
subtruncate; two low, radial depressed rays extend from the
beaks obliquely to the ventral margin; the central part of the
shell and the anterior end is sculptured roughly with irregular
growth lines; the posterior dorsal slope has strong, vertical or
slightly oblique lines which cross the finer concentric growth
lines; in addition the whole surface of the shell is roughened by
small, short threads as if the surface had been painted with a
stiff brush.
Length 63, height 25.50, diameter of the right valve
14.50 mm.
Toula’s figure of an internal mold shows a smaller shell, a
strong anterior-umbonal angle and a somewhat shorter anterior
end. It is possible that the two forms are distinct but we lack
a good comparative series of the Gatun shell. A smaller and
more delicate species of Psammosolen occurs at several localities
in the Costa Rican Gatun. It is not the P. strigzl/atus (L) of
Toula, differing in its pointed anterior end.
Gatun Stage: Rio Betey.
Genus SANGUINOLARIA, Lamarck
Section SAMMOTELLA, Blainville
Sanguinolaria alouatta, n. sp. Plate 29, figures 5, 6
Shell rather large, elongate; and rostrated behind; left
433 Costa RicA M1ocENE—OLSSON 261
valve nearly flat, the right valve much more convex; the right
valve carries a strong umbonal keel from the tip of the beaks to
the posterior extremity; this keel is but poorly developed on the
left valve; anterior end broadly rounded with the dorsal and
ventral margins nearly parallel; the posterior side is much nar-
rower and with its extremity pointed; the surface is smooth; the
dorsal-posterior slope of the left valve is less smooth and carries
coarse, irregular growth-lines; the interior is not known.
Length 54, height 30, semi-diameter 4 mm (a left valve)
Length 60, height 27.5 semi-diameter 7 mm (a right
valve)
This species stands intermediate between the S. Smzthwood-
wardt Maury from the Miocene of Santo Domingo and the re-
cent West Indian S. oferculata Gmelin. It differs from the S7zz¢h-
woodward, in being much larger with a less depressed left valve:
From oferculata the Costa Rican shell has higher and more
pointed beaks as well as being longer posteriorly. In full-grown
specimens of operculata, the surface generally shows radial lines
and striz which may become quite strong near the posterior
ventral margin. The Costa Rican shells are smooth except for
the growth lines.
Gatun Stage: Hill No. 3, Banana River.
Genus TAGELUS, Gray
Tagelus cebus, n. sp. Plate 29, figure 9
Shell elongate, moderately convex and solid; beaks central
with the anterior and posterior ends nearly equal, and with the
dorsal and ventral margins parallel and nearly straight; anterior
and posterior extremities subtruncate; a broad depressed zone
extends from the umbos to the ventral margin of the valves;
surface marked with irregular concentric lines of growth; inter-
ior concealed.
Length 36, height 13, diameter of the left valve 3.00
mim.
262 BULLETIN 39 434
The Szligua subegualis Gabb, from the Miocene of Santo
Domingo is doubtless a 7agelus, belonging to the Section J/es-
opleura Conrad. It differs from the present shell in being thin-
ner, more depressed and more pointed posteriorly. Gabb’s species
approaches very closely the recent 7. divisus Spengler of the
West Indian fauna, judging from the specimens in the Maury
collection at Cornell, differing mainly in its larger size.
The interior of the Costa Rican shell is concealed so that the
presence or absence of an internal rib cannot be determined.
Its heavier shell and general form, is more that of typical 7age-
lus differing from the recent gzbdus Spengler in its more central
beaks.
Gatun Stage: Rio Betey.
Genus MACTRA (L.) Lamarck
Subgenus MACTRELLA, Gray
Kactra exoleta Gray Plate 28, figure 2
Mactra exoleta Gray, 1837, Mag. Nat. Hist., p. 372.
Mactra exoleta Reeve, 1854, Conch. Icon., Mactra, fig. 16.
Comparision of our fragmentary shell with recent examples
from the Bay of Panama show no inportant differences. The
species is notable for its high, narrow, inflated umbos, strongly
depressed posterior-dorsal margin and alation of its dorsal-pos-
terior angle.
At the present time, this species is restricted to the Paciliic-
side, but represented on the Caribbean coast by the next, the
Mactra alata Spengler, common along sandy beaches. It is
therefore of more than usual interest to find these two species,
together in the Miocene beds of the Caribbean slope of Costa
Rica.
Gatun Stage: Boucary creek.
435 Costa RicA M1iocENE—OLSSON 263
Mactra alata Spengler Plate 28, figure 3
Mactra alata Spengler, 1802, Skriv. Naturch. Selsk., vol. 5, part 2,
Pp. 99.
Mactra alata Reeve, 1854, Conch. Icon., Mactra pl. 8, fig. 29.
Mactra (Mactrella) cf. alata Maury, 1917, Bull. Amer. Pal., vol. 5,
P- 395-
A thin, fragile species distinguished by its sharp dorsal-pos-
terior angle and hatchet-shaped shell. It is related to the exo-
feta, already referred to, but is a very much broader shell. In
exoleta, the dorsal-posterior area is flat or concave, with a slight
raised line which is bounded on each side by a markly depressed
or sunken band. In a@/afa this area is convex.
It is living today on the Caribbean side of the Isthmus, be-
ing quite common on sandy beaches.
Gatun Stage: Boucary Creek.
fill No. 3, Banana River.
Section HARVELLA, Gray
Macira estreilana, n. sp. Plate 28, figure 1
Shell rather large, ovate, thin, slightly convex; surface reg-
ularly, concentrically plicated; the fragmentary type specimen
about 38 mm in height has about 32 plicee which on the center
of the shell disk are spaced about 2 mm apart; these pliceze com-
mence as in typical arvella at the dorsal-posterior carina which
bounds the escutcheon and extends across the surface to the an-
terior extremity; the surface is, in addition, finely, concentrically
striated; lunule large elliptical, smooth or finely striated; escut-
cheon smooth; interior not known.
Length 50, height 38, semi-diameter 8 mm,
Although this is not a very rare species in the Costa Rican
Miocene, its thin, delicate shell renders the collecting of good
specimens very difficult and it is usually secured only as distort-
2604 BULLETIN 39 436
ed internals molds. It is related to the recent Pacific 7. elegans
Sowerby, but differs in being less convex, wider and with more
numerous and closer concentric plicee.
Gatun Stage; Coll. 6. Estrella River.
Coll. 2, Quitana creek.
Macira sincola, n. sp. Plate 28, figure 4
Shell thin, broadly subtrigonal, depressed, nearly equilater-
al; beaks small and pointed anteriorly; lunule absent; escutche-
on linear-lanceolate and defined from the shell disk by an elevat-
ed carina; the surface is sculptured with strong, wave-like, con-
centric plicee which are continuous from the anterior-dorsal mar-
gin to the edge of the escutcheon; on the type specimen there
are about 30 pliceze on the center of the shell disk, spac ed about
I.75 mm apart; the intervals between the concentric plicz
are deep and trough-like and in width equal to the plice them-
selves; in addition the whole surface of the shell is finely sculp-
tured with small, radial, vermiculate threads, best seen on the
summits of the plicze; the escutcheon is ridged on the middle
and smooth; the hinge, largely concealed in the sandy matrix, is
mactroid.
Length 36, height 27, diameter of the right valve 5.5
mm.
The systematic position of this interesting shell is very
doubtful. Its general surface sculpture of strong, concentric
plicze and well-defined escutcheon resembles that of Havrvella.
The surface is covered with fine, vermiculate threads as seen on
Labiosa.
Gatun Stage: Saury creck.
Genus LABIOSA, (Schmidt) Moller
Labiosa lineata Say Plate 21, figure II
Lutraria lineata Say, 1821, Journ. Acad. Nat. Sci. Phila., vol. 2, p.
310.
437 Costa Rica MIocENE—OLSSON 265
Mactra Nuttallii Reeve, 1854, Conch., Icon., Mactra, fig: 125.
Labiosa lineata Dall, 1898, Trans. Wagner Free Inst. Sci., vol. 3, pt.
4, p- go6.
This species is represented in our collection by fragments of
the hinge and a perfect mold of the interior which is figured.
This mold compares very closely with recent specimens of Ladz-
osa lineata Say.
Dall records this species as fossil in the Pliocene Caloosa-
hatchie beds of Florida and we have several fragments in the
Cornell collection from the Upper Chesapeake Miocene of Nat-
ural Well, North Carolina.
Gatun Stage: Saury creek.
Coll. 6, Estrella River.
Genus MULINIA, Gray
Muiinia lateralis Say, var. sauryensis, n. var. Plate 28, figures 23. 24
The species of Mulinta are extremlly variable and we have
therefore associated this form with the common /atevalis of Say,
although it may later prove to be a distinct species.
The general form of the shell is like corbulotdes Deshayes,
the common Gulf variety of lateralis of Say. It differs most im-
portantly from /aferalis in having the beaks situated a short dist-
ance above the hinge-line as is seen in the Chesapeake Miocene
M. congesta Conrad. In other characters, the Costa Rican shells
are more closely related to the J/atervalis, than to the congesta.
The shell is fairly convex and thin.
Length 20, height 16, semi-diameter 6 mm.
Gatun Stage: Zone E, Saury creek.
Genus MESODESMA, Deshayes
Mesodesma costaricensis, n. sp. Plate 28, figure 18
Shell ovate, moderately convex, inequilateral, thin; anterior
266 BULLETIN 39 438
end about % again as longas the posterior, rounded; beaks
high, triangular; posterior and anterior dorsal margins straight;
ventral margin evenly rounded; surface nearly smooth; but with
the dorsal submargins finely concentrically grooved; interior
concealed.
Length 22, height 16, semi-diameter 4 mm.
A rather thin, Dozax-like shell from the Lower Gatun beds
of the Banana River. Its surface is smooth and polished with
concentric grooves on its posterior and anterior dorsal area.
Gatun Stage: Hill No. 3, Banana River.
Genus GASTROCHAENA, (Spengler) Cuvier
Gastrochena rotunda Dall
Gastrocheena ovata Sowerby, variety rotunda Dall, 1898, Trans. Wag-
ner Free Inst. Sci., vol. 3, pt. 4, p. 825.
Gastrochena rotunda Dall, 1915, Bull. N. S. Nat. Mus., No. 90, p. 157,
pl. 19, fig. 2.
A single, small specimen of a Gastrochena was collected in
the Gatun of Red Cliff Creek. It is widely expanded about the
posterior portion and probably represents a young shell of G. ro-
tunda which Dall records from the Bowden beds of Jamaica.
This species is alsofound in the Tampa Silex beds and in the
Lower Miocene of Florida.
Gatun Stage: Coll. 4, Red Cliff Creek.
Genus CORBULA, Lamarck
Section ALOIDIS, Megerle
? Corbula vieta Guppy Plate 28, figures 15, 16
Corbula meta Guppy, Quart. Journ. Geol. Soc. London, vol. 22, p.
580, pl. 26. fig: 8.
Corbula disparilis Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
247.
Corbula (Aloidis) vieta Dall, 1898, Trans. Wagner Free Inst. Sci.,
vol. 3, pt. 4, p. 859.
439 - Costa RicA MiocENE—OLSSON 267
Corbula (Aloidis) vieta Maury, 1917, Bull. Amer. Pal., vol. 5, p. 395,
pl 39, fig. 13.
The members of this group of Corbule are difficult to sepa-
rate satisfactorily as all are quite similiar in their characters. The
right valve is broad, high and coarsely sculptured with regular
concentric coste, while the left valve is much smaller, longer
than high, with its surface smooth or faintly lined with un-
equally distributed radial threads.
The C. viefa was described by Guppy from the Manzanilla
beds of Trinidad and most authors have regarded the A7ycina
tensa described at the same time as its left valve, an indentifica-
tion which is open to question. Guppy’s figure of fezsa, shows
a Veneroid shell strongly sculptured with even radials and
fine concentric lines, and quite unlike the left valves of Corbule
of the A/ozdzs section.
? Corbula vieta from Costa Rica is identical with Maury’s
specimens from the Miocene of Santo Domingo. It is a smaller
and more circular shell than the Coréula heterogenea Guppy
which we have from the Canal Zone.
Length 6.25, height 6.50, semi-diameter 3 mm. (right
valve)
Gatun Stage: Water Cay.
Hull 1a, 3. Bananavketver
Section CUNEOCORBULA. Cossman
Corbula oropendula, n. sp. Plate 28, figures 12, 13, 14, 26, 27
Shell small, solid, elongate, both valves of nearly equal con-
vexity but the the right valve somewhat larger and overlapping
on the left; beaks nearly central, the anterior end well rounded,
the posterior somewhat contracted, slightly rostrated and with
a well-marked keel; ventral margin more or less rounded, ter-
minating atis posterior end ina small sinus, just anterior to
268 BULLETIN 39 440
the keel; the basal margins of both valves are folded in and
flattened as seen in C. contracta Say but toa less extent and
least in the right valve; surface sculpture of rather heavy con-
centric costee, most widely spaced on the umbos; the surface
may also show fine radial lines of small pustules or puncte
which are best developed on the rostrum; no lunular pit.
Length 6.5, height 4.25, diameter 4 mm (Banana River ).
9 6 semidiameter 2 mm (Saury).
We have chosen as the type of this spceies, the smaller and
more abundant form as found in the Gatun bed of the Banana
River, rather than the larger shells from the more inaccessible
Saury. It is related to the C. contracta Say, but differs in hav-
ing the ventral margin more curved, and the posterior extremi-
ty more emarginate. Its surface is sculptured with rather
strong, concentric costee which are more widely spaced on the
umbos, and submicroscopic radial lines of fine punctze or small
pustules, best seen on the rostral areas.
Gatun Stage: Hills ra, 3, Banana river (abundant).
Zone G, Saury creek.
Gol On Rea Oli enecr:
Coraula Dietziana C. B. Adams Plate 21, figure 8
Corbula Dietziana C. B. Adams, Contr. Conch., vol. 12, p. 235.
Corbula (Cuneocorbula) Dietziana Dall, 1898, Trans. Wagner Free
Imsty Seis) voli2. pt Ayip isso:
A recent species quite common in the coralline limestones
of the Limon Peninsula, from which it has previously been re-
ported by Dall from specimens collected by Hill.
In its earlier stages of growth, this species resembles the
C. eguivalvis Phil._(C. cubantana d’Orb) in its rectangular from
and strong dorsal keel. Later in life, the shells become strongly
distorted, the ventral margin is abruptly bent downwards nearly
at right angles to the former plane of the shell as is seen to a
much less degree in’the recent C. contyvacta Say. Its surface is
441 Costa RicA MIocENE—OLSSON 269
coarsely concentrically grooved, the two valves being sculptured
alike.
Gatun Stage: Port Limon.
Corbula hexacyma Brown and Pilsbry Plate 21, figures 12, 13
Corbula hexacyma Brown and Pilsbry, 1912, Proc. Acad. Nat. Sci.
Phila., vol. 64, p. 518, pl. 26, fig. 4.
A species of the Gatun beds of the Central Zone. The shell
is rather large, solid and with its valves similiary sculptured
with a few, deep, wave-like, concentric ribs. Its right valve is
slightly more convex than its left. Our specimens measure:
Length 20, height 14, diameter of the left valve 6.00
mm.
Length 20, height 13.50, diameter of the rigth valve
5.75 mm.
Gatun Stage: Gatun, C. Z.
Corbula orosi, n. sp. Plate 21, figures 9, 10
Shell elongate, convex with the low inconspicuous beaks
about the middle of the shell, so that the posterior and anterior
ends of the shell are about equal in length; the right valve is
slightly larger and its basal margin folds over slightly that of
the left valve; anterior end rounded, the posterior narrow and
more pointed but with the keel only very poorly developed; the
surface is finely and evenly concentrically ribbed and both valves
are similiarly sculptured; no lunule.
Length 20, height 11.5, diameter of both valves 9.5 mm.
A rather large species with nearly equally convex valves;
surface sculptured with fine, regular, concentric ribs. It
resembles the C. sarda Dall, from the Chipola Miocene but is
much larger.
Gatun Stage; Hill No. 3, Banana River.
270 BULLETIN 39 442
Section BOTHROCORBULA, Gabb
Corbula viminea Guppy Plate 28, figure 25
Corbula viminea Guppy, 1866, Quart. Journ. Geol. Soc. London, vol.
22D A2Oa nla Zon iong Tere
Lothrocorbula viminea Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15
p. 247.
Corbula ( Bothrocorbula) viminea Dall, 1898, Trans. Wagner Free Inst.
Sci., vol. 3, pt. 4, p. 850.
Corbula (Bothrocorbula) viminea Maury, 1917, Bull. Amer. Pal., vol.
5, Pe 39754 Dl- 39) es. 20.) 2)
This is a very characteristic species, distinguished by its
large, deep lunule and its surface strongly sculptured with
coarse, concentric, wave-like coste. It is known from the Mio-
cene of Jamaica and Santo Domingo.
The Corbula hexacyma Brown and Pilsbry, still known on-
ly from the Gatun of the Canal Zone is similiar in form and
sculpture but lacks the deep lunular pit.
Length 15, height 10, semi-diameter 4.85 mm (right
valve).
Gatun Stage: Middle creck.
443
Fig.
Costa RicA MiocENE—OLSSON 271
EXPLANATION OF) RE AduES
PLATE 1 (4)
Page
I. Terebra bipartita Sowerby, Saury Creek, height 77 mm........ 35
2. Terebra dipartita Sowerby, Middle Creek, height 53 mm.
3. Terebra cf. haitensis Dall, Sousi Creek, height 37 mm......... 35
4. Terebra gatunensis Toula var., Gatun, height 58 mm............ 36
5. Terebra gatunensis Toula, Gatun, height 52 mm.
6. Terebra gatunensis Toula, Gatun, height 42 mm.
7. Terebra spirifera Dall, var. midiensis, n. var., Middle Creek 39
8. Terebra gausapata Brown and Pilsbry, Mt. Hope, height
RUUWbe cba NARA MAAN RAO Hoa ra cab Woan Suan Kae AUABBO be 38
9. Terebra estrellana, n. sp. Estrella River, height 7.5 mm...... 40
Io. Terebra gausapata Brown and Pilsbry, Mt. Hope, height
PXS) NOW oO peKeedeasccqdsooRaccQEHSeS SGouuacboaseBseupoasoogde 38
11. Terebra Wolfgangi Toula, Mt. Hope, height 38 mm.............. 37
12. Terebra Wolfgangi Toula, Mt. Hope, height 26 mm.
iewlerebraspintiera Dall iBananapRaivetpecsecnee se seeeea eae a tees antes 37
14. Conus gracilissimus Guppy, Bocas del Toro...... .........2.--0+55 52
I5. Conus tortuosostriatus Toula, Bocas del Toro..................-...- 49
ROwMMverebra estrellanay nm.Wsp', Estrellay Rivet nes. yal meee ain aaans ate 4o
17-18. Terebra Berlinerze Maury, Boucary Creek.. ........ PE Asse A 39
19. Terebra costaricensis, n. sp., Banana River, height 13 mm... 40
20.) (Conus marginatus Sowerby, Red Clift Creel nee eee seo eee ee 51
Zig COnuUS mW tS triatusVvBOSe Water Calycsauneeseeceesensyeescee cece 54
2249) COMMS MUScensis. ns Spy, Bamana Ravietesece amy cenchecseeeaeeeestece as 47
22a Conus muUltistriatus BOSE Water) Caine teases seecceee ee eeeee 54
DARN COMMS HMUSZENSIS) My Spy. bata WRU CT enseetaeeneseeceacer ete cee 47
25- 26. Terebra benthalis Dall, var. bocasensis, n. var., Bocas
Daf
Caled Wid oy oy MAU ian NE EA a aR URE 4I
B72
Fig.
Fig.
Fig.
BULLETIN 39 444
PLATE 2 (5)
Page
I-2. Conus molis Brown and Pilsbry, Banana River............ ..... 42
3-40) /Conus proteus) /Euwassi) NViaitern © aiyesie. eats arene ane eeeane 43
5. ConusiViea teh ty yaknspy MMW eit ergs Cay euch aera Ue eae ie Unc nts a manuae 44
6. Conus imitator Brown and Pilsbry, Mt. Hope....... .....-.....00- 45
Yo \ Orbis) WdosroreraIsS,) sole Gio; | IROIKO (Cals ccssopicnodiccscddcesudcoo eeoBobeuoade 48
8), Conus Veatch misma a NVave rth @aly aula suse ease menene 8) 0 icsaee 44
93) Conus! recognitus) Guppy. Banana Rivenroar eee ue .cee eee 46
PACD) 2400 6))
jig) (CONS Je levewisi, tol, Siow. IBkewatebnel IRIE. shocc wssosoneeaes! \aune ;cbosodosooce 53
2. Conus|stenostomus) Sowerbys) Bone Mu im Oneeeee senna. oe see ene 42
3. Conus floridanus Gabb, var. costaricensis, n. var Banana R... 45
AE: Cloyohntsy JeyouRel'd akahqobbl JBxoRS. | NWE\dese (OAy7 155 sooosodcosucsoocseascocab0SazC 52
6))) Conus tortuosopunctatus Moulaw Mit. elope. ee a ease ene eee 54
7freys) (Clovonbis}, IhhooVoyoVeraisyisy 014 /G}o},4) Jetorae ILpbooVha aqcoopocadécoosdocbndebocaods AT
g. Conus floridanus Gabb, var. costaricensis, n. var., Banana R. 45
Lov Conus planilinatus) Sowerby MeOnts lei O mts tren seaee eae tee ene 50
Tt.) \Conustortuosopunctatus)Moulav Mth Hiopersice-csssseencceeeeeees 54
12. Conus cf. interstinctus Guppy, Banana Rivetr..................0.00+ 48
13.) Conus planiliratus) Sowerby, Port, Wimonm sc: se4 ee sseeee cece 50
14. Conus planiliratus, var. bocasensis, n. var., Bocas del Toro.. 57
152), Conus pranozonatus) Guppy, Rio) Blanco ws wesse.sseeneeceee ese eeae 50
16. Conus planiliratus var., bocasensis, n. var., Bocas del Toro.. 51
PLATE 4. (7)
1-2, Turris albida Perry, slightly reduced, Toro Cays............ 58
3. Scobinella Morierei (Laville) Cossmann, length 48 mm, Toro
CaS eae Ee awa aaa sistent eet tees 79
4. Scobinella Morierei (Laville) Cossmann, length 52 mm.,
Moro) Cay sys oseaucnweaeecece sete 79
5. Drillia papaya, n.sp., length 32 mm., Rio Cocles............... 63
6. Turricula lavinoides, n. sp., length 54 mm., Banana River.... 55
Halia americana, n.sp., length 66 mm., Mt. Hope............... 79
445
Fig.
CORSON CME ee Oo UE
22.
23.
Costa RicA Mr1ocENE—OLSSON 273
Page
Drillia consors Sowerby, Grape Point Creek, length 60mm... 61
Drillia venusta Sowerby, Banana River, length 30 mm........ 61
Drillia consors Sowerby, Mt. Hope, length 33 mm......... ..... 64
Drillia carruca, n.sp., East Grape Point Creek, length 18mm. 71!
Turricula lavinoides, var. limonensis, n. var., length 97 mm.,
MOrOy Cary Sey CMe aati castle 56
Drillia consors Sowerby, length 4o mm, Banana River......... 61
Bullaria paupercula Sowerby, Middle Creek, length22 mm... 34c
Drillia limonica, n. sp., Port Limon, length 19 mm.............. 66
Ancistrosyrinx Dalli, n. sp., Toro Cays, length 14 mm......... 60
Ancistrosyrinx elegans Dall, var., Banana R.,length 15 mm _ 60
PLATE 5. (8)
Drillia Cristobali, n. sp., Bocas del Toro, heigth 20 mm........ 72
Drillia chirquiensis, n. sp., Bocas del Toro, height 25 mm.. 64
Turricula taurina, n. sp., Bocas del Toro, height 25 mm........ 57
Turricula Terryi, n.sp., Bocas del Toro, height 23.5 mm..... 57
Drillia bocasensis, n.sp., Bocas del Toro, height 22 mm..... 73
Drillia bocatoroensis, n. sp., Bocas del Toro, height 23 mm.. 67
Driliiarcoccos,s 1) Spis)\Cocovelum\sheigiae 2mm) sees ecee se. 64
Drillia limonensis, n. sp., Port Limon, height 20 mm......... 71
Drillia limonensis, n. sp., height 15 mm.
Drillia limonetta, n. sp., Port Limon, height 12.25 mm......... 70
Drillia theobroma, n. sp., Bocas del Tcro, height 21 mm...... 68
Drilliacitriat nusp..\/Lort Wimontohieleipyien santa sense erste 65
Drillia citria, n. sp., Port Limon, height 11 mm.
Drillia cocosina, n. sp., Bocas del Toro, height 12 mm........ 75
Drillia aurantia, n. sp., Port Limon, height 11 mm..... ...... 68
Drillia aquanica, n. sp., Water Cay, height mm.................. 65
Drillia aquanica var.
Drillia lithocolletoides, n. sp., height 10.75 mm.,
IBXOY CRIS GIS NoidO\an nds neu ausecosendhesoqsseads 74
Leucosyrinx chloris, n. sp., height 14.50 mm.,
BocasrdelgMorou tee mean wasn uae! 69
Cytharella limata, n. sp., height 9 mm, Port Limon.............. Ta
Cythara terminula Dall, var. costaricensis n. var., height 13
mm., Hill 1a, Banana River...... 77
Cythara terminula Dall, var. costaricensis n. var., height 13
mm., Hill ra, Banana River......... 77
Borsonia cocoensis, n. sp., Coco Plum, height 16 mm............ 78
2 Woe G74 BULLETIN 39 446
Page
Fig. 24. Borsonia cocoensis, n.sp., Coco Plum, 14 mMl.........-.....2006 78
25. Drillia aquzensis, n. sp., Water Cay, height 15 mm.............. 73
26. Drillia musa, n.sp., Banana River, height 16 mm............... 74
27-28. Drillia musacina, n.sp., Banana River, height 9 mm ..... 69
29-30. Glyphostoma moinica, n. sp., Port Limon, height,
Eni}: 0dbaDY, ond Gon dooasopaseeaoneedoRonooDseagce 76
Fig.
PANE) 361199)
I. Mitra Swainsoni Broderip, var. limonensis, n.sp., Port
IWibesvoyay, oKeRKelole * /Al: TaBWEO,. «Ass nogaaeaqocdodse 16606 100
2-3. Cancellaria Plummeri, n.sp., Toro Cays, height 34 mm... 85
4. Cancellaria toroeusis, n.sp., Toro Cays, height 29 mm ....... 84
5. Cancellaria Maury, n. sp., Cercado de Mao, Santo Domingo,
JaKesGegalty /GX6), SOUTH ag aenoonvosadoogaosodccoogs oooaECdosS 82
6. Cancellaria Barretti Guppy, slightly enlarged, Banana River. 81
7. Cancellaria Rowelli Dall, East Grape Point Cr., height 30mm. 84
8. Cancellaria dariena Toula, about natural size, Toro Cays........ 80
g. Cancellaria Cossmanni, n. sp., somewhat enlarged, Banana R. 81
Lo.) Matra longa Cabby iGatun) heiglit) oy mimeo se pasccecnacssacseees IOI
11. Cancellaria Cossmanni, n. sp., somewhat enlarged, Banana R_ 81
12. Cancellaria islacolonis Maury, enlarged, Red Cliff-Creek........ 86
13. Mitra cf. rudis Gabb, Hone Walk Creek, height 33 mm.......... 102
14-15. Marginella MacDonaldi Dall, Banana R., height 28. mm... 95
16. Marginella mindiensis Cossmann, Gatun, height 9 mm.......... 96
17. Marginella MacDonaldi Dall, Banana R., height 26 mm......... 95
18. Mitra Almagrensis Toula, variety coralliophila, n. var., Port
Limon, height 17 mm.......... 103
19. Marginella MacDonaldi Dall, Banana River, height 25 mm.... 95
20-21. Mitra poas, n. sp., about natural size, Port Limon.............0. 102
22. Marginella leander Brown and Pilsbry, enlarged, Gatun........ 98
23. Marginella mindiensis Crossmann, Gatun, height 17.5 mm...... 96
24. Mitra Almagrensis Toula, variety coralliophila, n. var., Port
Limon, height about 17 mm........ 103
25. Mitra dariensis Brown and Pilsbry, Bocas del Toro, height
2) G5 0008 ARB HAH ER aE i Sore OF ese asdabacée IOI
I. Oliva cylindrica Sowerby, Old Man Sam Creek.......... Hocesclnae 88
2-3. Oliva brevispira Gabb, Red Cliff Creek.......... awa aciaaas Bhansee 88
447
Fig.
Fig.
Costa RicA MIocENE---OLSSON 275
Page
4. Oliva brevispira Gabb, var. Giraudi Cossmann, Middle Creek. 88
Sm Olivarcatinensis: Loulaw Gatun eesccecn ecco oe een ccretanseceeee 89
6-7. Oliva Sayana Ravenel, var. immortua Pilsbry and Brown
BananaiRiverncnsewudevossescieeeeccet 89
8-9. Oliva mancinella, n. sp., East Grape Point Creek......... sesenT PO)
LOmeeAnelllariajaqiuzensis, ni/Spi Water Cay.iueeesesacesseneseerese sce ce 94
11 Olivella muticoides Gabb, East Grape Point Creek................ 92
12-13. Oliva testacea Lamarck, var. costaricensis, n. var., Banana
RAVE TA eee ANU ARLE) SU aL 90
12. Olivella muticoides Gabb, Red Cliff Creek... .............. ceceeeee ee, 92
15-16. Olivella Boussaci Cossmann, var., Banana River............... 93
17-18. Olivella muticoides Gabb. Red Cliff Creek. ..................5. 4 92
19-20. Olivella limonensis, n. sp,, Port Limon.................60eeseeeee 93
21. Marginella avena Valenciennis, Banana River ..................0+- 97
pom Olivelia Goliath, im. Sp Red Ciith\ Greeks usec sesseseseeencee- 92
23. Olivella Goliath, n. sp., Banana River
24-25. Olivella limonensis, var., bocasensis n. var., Bocas del
TOTO Te Se Mane MN e Ua EU OMaLEE UIs Lue IU 94
26-27. Marginella collina, n. sp., Banana River................0000eeeeees 97
28. Marginella avena Valenciennes, Port Limon.....................-:. 97
PLATE 8. (If)
1. Fasciolaria MacDonaldi, n. sp., Banana River, height 70 mm. 106
2. Voluta Alfaroi Dall, Banana River, height 63 mm............... 99
3. Solenosteira chiriquiensis, n. sp., Water Cay, reduced........... 114
4. lLatirus taurus, n. sp., Toro Cay, slightly reduced................. 108
5. Fusus miocosmius, n. sp., Bocas del Toro, height 100 mm..... 103
6. Fusus honensis, n. sp., Hone Walk Creek, height 63 mm ..... 104
7. Solenosteira Vaughani Dall, variety medioamericana, n. var.
Toro Cary Sis aera neal iar un) iain 2
8. Solensoteira Dalli Brown and Pilsbry, Gatun, slightly en-
langed seni i Men at wataae cr 113
g. Fasciolaria Gorgasiana Brown and Pilsbry, Rio Betey, height
Ko Vb cab oe aM anUON NAW enue eto tata UML Ae nag 105
10. Latirus infundibulum Gmelin, Pumbri Creek, slightly re-
GUC ed eset ee ued GOS en Ns 107
11. Peristernia insula, n. sp., Water Cay, height 23 mm............ IIO
12. lLatirus irazu, n. sp., Port Limon, height slightly reduced..... 107
13. Peristernia tortugera, n. sp., Port Limon, height 29 mm....... 110
BULLETIN 39 448
PLATE 9. (12)
Page
Melongena consors Sowerby, Old Man Sam Creek, height
TEINS) (SBODBHIS Gaba0G54 Hansa pup oabodoonbEedodcdec5 112
Phos Veatchi, n. sp., Hill 1a, Banana River, height 43 mm.. 121
Phos Veatchi, n. sp., Hill 1a, Banana River, height 41 mm.. 121
. Phos gatunensis Toula, Gatun, height 30 mm................-..5. Mie)
Phos beteyensis, n. sp., Rio Betey, height 26 mmu................. 122
Northia northiz Gray, variety miocenica, n.var., tip, Banana
ISOMEIEY Sretondessancos sooBdOuSeRABRposacHodona 124
. Phos Moorei Guppy, variety costaricensis, n. var., Banana
RVs Jolemelate, 2X6) weoknalsesoaqeac GudonoodC 118
Phos mexicanus Bose, Water Cay, height 29 mm...... ........5. 117
Phos mexicanus Bose, Water Cay, height 27.5 mm.
Phos elegans Guppy, variety limonensis, n. var., Port Limon
leraerel nl 237/, TaabIal se ccosddondooaboooosds eo00e 119
Phos elegans Guppy, variety limonensis, n. var., Port Limon
length 22.50 mm
Northia northie Gray, variety muocenica, n. var., Banana
VET Sh el pac Gu ASEM ROn Rae HaRAA MR nen aie aio 124
Phos snbsemicostata Brown and Pilsbry, Saury Creek, height
AD) MGSO oa seoeadce BBen Aan nee sone aenboe se 121
Phos metuloides Dall, Mt. Hope, height 19.5 mm............... 122
Phos estrellensis, n. sp., Estrella River, height I9 mm......... 120
Phos estrellensis, n. sp., Estrella River, height 16 mm.
Alectrion merenda, n. sp., Old Man Sam Creek, height 7 mm 124
Alectrion ranuncula, n. sp., Banana River, height 14 mm..... 123
Alectrion losquemadica Maury, Port Limon, height 11.50mm 123
Alectrion ranuncula, n. sp., Banana River, height 14 mm...... 123
PLATE 10 (13)
Marginella latissima, variety Pilsbryi, n. var., Banana R...... 96
Marginella latissima, variety Pilisbryi, n. var., Gatun
. Clava costaricana, n. sp., Banana River, height 40 mm........ 144
Metula Harrisi, variety limonensis, n. var.. Port Limon,
InlSsheloly) 7/ Swaheolyonane? Shouse soqsobacdosedco 116
Metula Harrisi, variety limonensis, n. var., Port Limon,
eto hey 25 TAT esescs pennant 116
Strombina matima, n. sp., Banana River, height 12 mm........ 127
449 Costa RicA M1ocENE—OLSSON 277
8. Strombina costaricensis, n. sp., Middle Creek, height 15 mm 128
g. Strombina ambigua Guppy, Banana River, height 26 mm...... 126
to. Metula Harrisi, n. sp., Coco Plum, height 27 mm................. 116
11. Metula Gabbi Brown and Pilsbry, Mt. Hope, height 28.5 mim 115
12. Metula cancellata Gabb, Bocas del Toro, height 18.5 mm...... 115
13. Strombina Lessepiana Brown and Pilsbry, Gatun, height
AS Sanvad Landay. aude jnppaauAaoooebadbaaonoca 126
AMe SECO bia Chiniguiensisy maiSPilucamatnnene re acee ce ee eneeee oie eecesns 130
15. Typhis alatus Sowerby, Toro Cays, height 23 mm................- 132
16-17. Strombina costaricensis, n. sp., Middle Creek...............0. 128
18. Strombina cricamola, n. sp., Water Cay, height 16 mm......... 127
19. Strombina sincola, n. sp., Banana River, height 7 mm.......... 129
20. Strombina Lessepiana Brown and Pilsbry, Gatun................. 126
12. Strombina sineola, n. sp., Banana River, height 7 mm......... 129
22. Typhis linguliferus, variety costaricensis, n. var., Bannan R. 132
22y eoistrum) nodulosum ©. Bs Adams, Red Cliff) Creeks co .5.5-. 133
PAM StLOm bia Chiriquiensiss das )Spi isan uen eel oe wasulatesleclsicele sels salle 130
25. Marginella musacina, n. sp., Banana RiveT........ 0.0.0.0... se000 98
26-27. Strombina costaricensis, variety musanica n. var. Banana R 129
28. Cerithium costaricensis, n. sp., Red Cliff Creek, height
PX oii DOOD AA ANUN GRRE AA BANE aaa SeRaeReToe see ae 143
29. Typhis linguliferus, variety costaricensis, n. var., Banana R. 132
30. Clava alajuela, n sp., Middle Creek, height 17 mm............. IA5
31-32. Drillia estrellana, n. sp., Estrella River, height 7.50 mim... 69
33-34. Columbella submercatoria, n. sp., Red Cliff Creek.............. 125
PLATE 11 (14)
Fig. 3. Xancus scopulus, n. sp., Banana River, height 265 mm......... TIt
PLATE 12 ({5)
Big: 1. Serpulorbis papulosa Guppy, Banana River........................-. 145
2. Malea elliptica Pilsbry and Johnson, Banana River, height
G3 STAY ee Ne SS et aera 139
3. Sconsia levigata Sowerby, variety Gabbi, n. var., Toro Cays,
cbvedondlyy, SrallahsegeC onda aouasdanopodad se: 136
4. Siliquaria modesta Dall, variety limonensis, n. var., Port
Limon, somewhat enlarged......... 147
5. Siliquaria modesta Dall, variety limonensis, n. var., Port
Limon, slightly enlarged............ 147
278
Fig,
Fig.
BULLETIN 39 450
Page
6. Siliquaria modesta Dall, variety limonensis, n. var., Port
Limon, slightly enlarged............ 147
7. Sconsia cocleana, n. sp., Rio Coles, height mm.................... 138
8. Cypreea cinerea Gmelin, Rio Blanco.............0..cccccceesescereeese 140
g. Cyprea almirantensis, n. sp., Water Cay, height 39 mm...... 140
10, Cypreea parisimina, n. sp., Port Limon, height 39 mm......... 139
11, Phalium moniliferum Guppy, Rio Betey, height 35 mm........ 135
12. Sconsia bocasensis, n. sp., Bocas del Toro, height 35 mm...... 137
13. Sconsia bocasensis, n. sp., Bocas del Toro.
PLATE 13. (16)
1-2-3. Strombus pugiloides Guppy, Banana River...................05- 142
4. Fasciolaria tulipa Linnzeus, variety Port Limon................... 105
5-6. Strombus gatunensis Toula, Water Cay, somewhat reduced.. 141
7. Polinices Stanislas-Meunieri Maury, Banana River, slightly
SRO NEKO los ngnocdoquosadesedddn sles 157
8. Natica Youngi Maury, varietv cocleana, n. var., Cocles
Creek, slightly enlarged............. 155
ov Natica canrena Ieimmecisy Gatun eee see ee epee eeneee ee eee seeneee 155
10-11-12. Architectonica granulata Lamarck, Banana River......... 154
13-14-15. Natica Guppyana Toula, Banana River...............:06 see 156
16-17. Polinices subclausa Sowerby, Banana River..................000+ 157
PLATE 14. (17)
i. Turritella oreodoxa, n. sp., Banana River, height 49 mm........ 152
2. Turritella exoleta Linnzeus, variety limonensis, Port Limon,
LOSS olle Ate Fools 4 Gescenugaopscasedoceoace 152
3. Turritella exoleta Linnzeus, variety limonensis, n. var., Port
Limon, height 36 mm
4. Turritella altilira Conrad, variety chiriquiensis, n. var.,
Banana River, height 52 mm...... 150
5. Turritella mimetes Brown and Pilsbry, Gatun, height 72 mm 149
6. Turritella altilira Conrad, Mt. Hope, typical, height 50 mm 150
7. Turritella altilira Conrad, Gatun, typical, height 36 mm...... 149
8. Turritella altilira Conrad, variety chiriquiensis, n. var., Wat-
er Cay, height 116 mm. ............. 150
9. Turritella altilira Conrad, variety chiriquiensis, n. var. Ban-
ana River, height 4o mm............ 150
Fig.
Costa Rica MioccENE—-OLSSON 279
Page
. Io. Petaloconchus sculpturatus H. C. Lea, Water Cay, height
YAM oD OTB EBS RBA MACHO aa AN OR SAM SiN cane anr
It. Turritella altilira Conrad, variety costaricensis, n. var., Sousi
Greek Sheiolt QAi mms ease
12, Turritella gatunensis Conrad, Water Cay, height 36.5 mm.....
13. Turritella gatunensis Conrad, Water Cay, height 24 mm.
14. Turritella altilira, variety chiriquiensis, n. var., Red Cliff
Creeks iheiabtig aim malay eaeeeeseeeeees
I5. Petaloconchus sculpturatus H. C. Lea, East Grape Point
Creek Whleioliia a yirarreen ares see seen
PVA TS len)
1. Dentalium uscarianum, n. sp,, Rio Cocles, natural size.........
2-3. Dentalium. bocasensis, n. sp., Bocas del Toro, somewhat
enlarged. a eat NEO ANE I NE
4. Astralium brevispinum Lamarck, variety basalis, n. var., Old
Man Sam Creek, natural size......
5. Astralium brevispinum Lamarck, variety basalis, n. var., Old
Man Sam Creek, natural size......
6-7. Calliostoma castilla, n. sp., Port Limon, slightly enlarged.....
8. Calliostoma limonensis, n. sp., Port Limon, about natural
STZ NaS MUU UT SOAK eae UE A ED
g-10. Calliostoma mancinella, n. sp., Red Cliff Creek, about nat-
WRAY SIZ SEN eR Ve uig acts N AUST Dba at
tt. Calliostoma limonensis, n.sp., Port Limon, about natural
SUZ OY UNM SD ANCA AT MEA PEED seat
IMuboysaltuss n> sps, Red: Clit Creek height mmh ees) ee
13. Scaphella costaricana, n. sp., Cocles Creek, height 29 mm.....
14 Ptychosalpinx ? dentalis, n. sp., Cocles Creek, about natural
15. Actzeon costaricensis, n. sp., Banana River, height 19.5 mm.
16. Chlorostoma costaricensis, n. sp., Red Cliff Creek, diameter
17. Crepitacella limonensis, n. sp., Port Limon, height 8 mm.....
18. Ptychosalpinx ? dentalis, n. sp., Cocles Creek, about natural
19. Bursa crassa Dillwyn, Banana River, about natural size........
20-21. Acteocina bullata Kiener, variety costaricana, n. var., Mid-
die Créeks heiehtiropmmeses.scscr:
150
146
166
166
Fig.
Fig.
is)
BULLETIN 39 452°
Page
Neritina viridemaris Maury, Middle Creek, enlarged............ 159
Alaba turrita Guppy, Estrella River, height 5 mm............... 153
PLATE 16 (19)
Pecten MacDonaldi, n. sp., type, Gatun (Toro limestone)
right valve, height 96 mm.......... 198
Pecten MacDonaldi, n. sp., cotype, same locality, left valve
height 79 mm.
Pecten gatunensis, Chagres Spillway, right valve, height
COPA oTb Gols Aas nao ha MERE RE ARE Hone hada 197
Pecten gatunensis, same locality, left valve, height 48 mm.
Pecten levicostatus Toula, Hill No. 2. Banana River, slight-
hf) sREYG NE CeL=YE Ae aon uaanoBooo sdonosdoed odor 199
Pecten levicostatus Toula, same locality, left valve, slightly
reduced
Pecten levicostatus Toula, same locality, right valve slightly
reduced
PLATE 17 (20)
Amusium luna Brown and Pilsbry, Chagres Spillway, some-
Wiltatine duce disease esata en esc eee 205
Pecten preglyptus, n. sp., Cocles Creek, natural size (type).. 202
Amusium bocasensis, n.sp., Bocas del Toro, natural size,
COLY POH ase dee sciseeeoneemeeessee eer 206
Amusium bocasensis, n. sp., Bocas del Toro, interior of right
valve.
Pecten coralliphila, n.sp., type, Port Limon, slightly en-
Pare nce ee CS ee Ue aa “t-99
Pecten coralliphila, n. sp., Port Limon, left valve................ 199
Pecten preglyptus, n.sp., interior view of type, natural size 202
PLATE 18 (2i)
Ostrea megodon Hanley, Banana River, natural size ........... 195
Glycymeris canalis Brown and Pilsbry, Comadre Creek, about
MatuTnalisize nesses wens sue 177
453 Costa Rica MrockxE—OLSSON 281
Page
3. Glycymeris canalis Brown and Pilsbry, Banana River, about
pacing INH Aer ANE AMR Ee At. Atta acts Neon 177
4. Glycymeris canalis Brown and Pilsbry, variety, East Grape
POM Cy Cree ke aN a eye aa Nofy
5. Glycymeris canalis Brown and Pilsbry, variety, East Grape
Pointy Greeley eee uu yy veakne enna tues Mola)
6-7. Glycymeris canalis Brown and Pilsbry, Banana River......... 177
Pecten costaricensis,n. sp., Red Cliff Creek, right valve........ 201
g. Pecten costaricensis, n. sp., Red Cliff Creek, left valve.
Io. Pecten costaricensis, n. sp., Red Cliff Creek, right valve.
Ir. Pecten costaricensis, n. sp., Red Cliff Creek, left valve.
12. Leda ensinoides, n. sp., Banana River, length 54 mm........... 174
13. Glycymeris castaneus Lamarck, Banana River,length 30 mm. 180
14. Glycymeris castaneus Lamarck, Banana River,length 23 mm.
15. Leda Balboz Brown and Pilsbry, Banana River, enlarged..... 173
16. Pecten almirantensis, n. sp., Bocas del Toro,heigth 4.75 mm. 203
IPppecten aOLus, ni sp: Port Limon, height 4i5o,mam.-.34) +... 204
18. Pecten aotus, n. sp., Port Limon, 4.75 mm.
19, Leda quitanensis, n. sp., Quitana Creek, length 26 mm........ 174
20uWeda Davila, m.\Sp.) Bocas deli Moros lenges 5-75) tate... sese-2 7a
21-22-23-24. Nucula cahuitensis, n. sp., Saury, length 3.50 mm..... 171
25. Nucula tenuisculpta Gabb, Red Cliff Creek,length 4.00 mm. 172
PLAGE Loy (22)
‘Fig. 1. Glycymeris carbasina Brown and Pilsbry, East Grape Point
Creek,aboutimatunalllsizensseisce sce 179
2. Glycymeris carbasina Brown and Pilsbry, same locality,
AOUb Ta tually Siz Cleese enna ease 179
3-4. Glycymeris jamaicensis, Red Cliff Creek, about natural size 178
5. Glycymeris carbasina Brown and Pilsbry, East Grape Point
Creek, small shell about natural
SUZ Gs ameter eer eA near Me ERLE NAM IUD ERO Uy 179
6-7. Glycymeris decussatus Lamarck, Old Man Sam Creek, nat-
Trad SIZE WAN saiaranees UNA Aen Aen re 180
PLATE 20 (23)
Fig. 1. Spondylus chiriquiensis, n. sp., Water Cay, height 151 mm... 206
2. Spondylus chiriquiensis, n. sp., young shell to show the small
foliaceous attachment area.
3. Thracia Tristani, n. sp., Rio Betey, length 42 mmi............... 211
282
Fig.
Fig.
nun
Io.
BULLETIN 39 454
Page
Crassatellites Reevei Gabb, Banana River, slightly reduced.. 212
Spondylus chiriquiensis, n. sp., Water Cay, young right
Spondylus chiriquiensis, n. sp., Water Cay, young left valve.
PLATE 21 (24)
Spondylus gumanomocon Brown and Pilsbry, Hone Walk
Creek, shell greatly reduced....... 207
Ostrea costaricensis, n. sp., Rio Blanco, about natural size... 196
Placunanomia lithobleta Dall, Banana River, height 36 mm.. 210
Placunanomia lithobleta Dall, Banana River, height 49 mm
Placunanomia lithobleta Dall, interior of same specimen as
figure 3.
Anomia simplex d’Orbigny, Middle Creek, height 34 mm..... 209
Corbula gatunensis Toula, Gatun, length 21 mm.
Corbula Dietziana C. B. Adams,Port Limon, length 15.5 mm. 268
Corbula orosi, n. sp., type, Banana River, length 19.5 mm... 269
Corbula orosi, n. s9., type, Banana River, length tg mm.
Labiosa lineata Say, Saury Creek, length 44 mm................ 264
Corbula hexacyma Brown and Pilsbry, Gatun,length 26 mm. 269
Corbula hexacyma Brown and Pilsbry, Gatun,length 18 mm.
PLATE 22 (25)
Arca occidentalis Phil., Port Limon, somewhat enlarged...... 181
Arca umbonata Lamarck, East Grape Point Creek, slightly
enlarged eee sy ree ee sae 182
Arca auriculata Lamarck,,Old Man Sam Creek, young shell.. 190
Arca Mauryze, n. sp., Port Limon, right valve of type, length
Arca candida Gmelin, Port Limon, left valve, length 18 mm. 183
Arca candida Gmelin, Port Limon, right valve length
13 mm.
Arca\Mauryce. (ns Spr ittenon Wlewy Onity Pea eene se eel-ieettanseee 182
Arca honensis, n. sp., Hone Walk Creek, left valve type,
slighitlys;nediiced wus teeesencee cee 187
Arca honensis, n. sp., same locality, right valve of cotype,
Sligdhtlyasne diced etn naieeheruecnescaee 187
Arca dariensis Brown and Pilsbry, Water Cay, about natural
SIZ ORME a Re A ICR NR tat 184
Fig.
Fig.
Fig.
1a &
Pao
3.
Costa RicA MrockENE—OLSSON 283
Page
Arca dariensis Brown and Pilsbry, Water Cay, about natural
Arca dariensis Brown and Pilsbry, Gatun, about natural size.
Arca dariensis Brown and Pilsbry, Gatun, about natural size.
PLATE 23 (26)
Arca Veatchi, n.sp., Water Cay, left valve of type, length
GMAT Gata ecine sesame cionainieigs eel neige 189
Arca Veatchi, n. sp., Water Cay, interior of left valve of type.
Arca Veatchi, n. sp., Water Cay, right valve of type.
4-5. Arca golfoyaquensis Maury, variety medioamericana,n. var.,
6.
Hone Walk Creek, valve of type,
Ikesaverdoy’ Gey Semboouogousaqnesodaqomabadeosde 188
Arca golfoyaquensis Maury, variety medioamericana, n. var.,
Hone Walk Creek, left valve of
type, same locality.
7-8. Arca actinophora Dall, Red Cliff Creek, nearly natural size.. 185
I
De
Be
6.
7
Xe
14.
aes
PLATE 24. (27)
Arca cacica, n. sp., type, Comadre Creek, length 36 mm...... 190
Arca Pittiern Dall Sauny) Creek mabunalisizessnuniecsssess secre ces 192
4-5. Arca-Pittieri Dall, Hill 1a, Banana River, natural size.
Arca Pittieri Dall, Hill 1, Banana River, natural size.
Arca Hindsi, n. sp., type, East Grape Point Creek, natural
RIYAS ei socsanerdodasnimacasods sdscoosoosesdeE 193
Arca Hindsi, n. sp., co-type, Rio Betey, natural size.
Arca Hindsi, n. sp., co-type, Hill-3,.Banana River, natural
size.
Arca Lloydi, n. sp., co-type, Red Cliff Creek, length 18 mm.. 192
Arca Lloydi, n. sp., type, same locality, diameter 18 mm.
Arca Lloydi, n. sp., co-type, same locality, length 15.5 mm,
Arca Henekeni Maury, Water Cay, length 30 mm................ 186
Arca Henekeni Maury, Water Cay, length 28 mm.
PLATE 25 (28)
Arca costaricensis, n. sp., type, Hill 1a, Banana River, nat-
TRA SIZE Peet ede cme atigauigunecnar tas 186
284
Fig.
nn
10.
is)
Lal
OR Or Oe ON Oates 2
20106
2s
13.
14.
BULLETIN 39 456
Page
Arca costaricensis, n. sp., co-type, Hill 1a, Banana River nat-
ural size.
Arca actinophora Dall, typical form, Mt. Hope, slightly re-
GUC CU ree Sak Shee aati eeaatate 185
Arca MacDonaldi Dall, Hill No. 3, Banana River, nearly nat-
ural size, interior view to show
ihighWicardinalltancateeers sac sesesacs
Arca MacDonaldi Dall, exterior of the same specimen.
Arca MacDonaldi Dall, variety subreversa, n. var., Rio Betey
exterior of left valve.
Arca MacDonaldi Dall, variety subreversa n. var., Hill No. 3
Banana River, interior of right
valve, to show form and low card-
inal area.
Glycymeris Lloydsmithi Pilsbry and Brown, Red Cliff Creek,
right valve, slightly reduced.......
Glycymeris Lloydsmithi Pilsbry and Brown, same locality
left valve.
Glycymeris Lloydsmithi Pilsbry and Brown, same locality
right valve.
PLATE 26 (29)
Tellidora cristata Recluz, Hill 3, Banana River, nearly nat-
MAGA STZ Oars a SE Sie a terest
Tellidora cristata Recluz, Hill 1a, Banana River, nearly nat-
ural size.
Tellina dariena Conrad, Water Cay, natural size.................-
Tellina lepidota Dall, Water Cay, length 24 mm..................
Tellina lepidota Dall, Water Cay, length 21 mm.
Tellina costaricana, n. sp., Banana River, about natural size
Tellina rovala, n. sp., Water Cay, slightly enlarged.............
Tellina rovala, n. sp., Water Cay, (type) slightly enlarged.
Tellina costaricana, n. sp., Banana River, about natural size
Tellina crystallina Wood, Banana River, enlarged nearly
Macoma panamensis Dall, variety canalis n. var., Mt. Hope,
eroverdol inky Senhiol sm hoocbondeeKd6) | coSbo00
Tellina umbra Dall, East Grape Point Creek, enlarged about
Bi abst) ocounsqoeegeocnubboouesHonDb05d0
Macoma gatunensis Toula, Banana River, about natural size
Macoma costaricana, n. sp., Rio Betey, about natural size.....
194
181
251
249
257
253
257
256
457 Costa Rica MiIockNE—OLSSON 285
Page
15. Tellina umbra Dall, East Grape Point Creek, enlarged about
BWA OBITS a Hagt eacsacoadsogoGce soud05 253
16. Macoma costaricana, n. sp., Banana River, about natural size 256
17. Tellina nasua, n. sp., Red Cliff Creek, about natural size...... 250
18. Strigilla musanica, n. sp., Banana River, length 13 mm....... 255
Ig-20. Venericardia Conradiana Gabb, variety limonensis, n. var.,
Portueinions leneth) 7mm)ys....senie 217
21. Chione grus Holmes, variety, Red Cliff Creek, enlarged about
AlGiTMesSeaaeiews: PAIGE A) ta ASIA Ban Ar SR 248
22. Cardita caribbeanensis, n sp., Banana River, length 18 mm. 214
23-24. Limopsis monilis, n. sp,, Old Man Sam Creek, length 1.75
ED FT oe daca dtonecate s6obh bene ucnvbpocadeapeade: 177
PLATE 27 (30)
Fig. 1. Cardium spinosum Meuschen, var. Turtoni Dall, Port Limon
aboutmatunallisizes nies aeeceh seieicece 228
2. Cardium dominicense Gabb, East Grape Point Creek, slight-
hi? SEC IWKOETE cocagcosatocdeogeqoneoosD jo6o0H 225
3. Cardium costaricauum, n.sp., East Grape Point Creek height
DIAN) CON en ANKE Soc S A SRO AAG rosa AHAB BABEe OnE 226
4-5. Cardium stiriatum Brown and Pilsbry, Banana River, slight-
Ny SSO BLES | so od bdomnododoosy sescaspanodcedo 225
6. Cardium medium Linné, Port Limon, about natural size....... 226
7- Protocardia gurabica Maury, Gatun, length 29 mm....... bedbesor 229
8. Protocardica gurabica Maury, Water Cay, length 25 mm.
g. Protocardia gurabica Maury, Water Cay, diameter 19 mm.
1o. Cardium heredium, n. sp., Rio Betey, length 11.50 mm........ 227
11-12. Cardium serratum Linné, Middle Creek, natural size......... 228
13. Protocardia costaricensis, n. sp., Comadre Creek, length 21
PADDED 3S dE MS SHIN AE UUM RNase 229
PLATE 28 (31)
Fig. 1. Mactra estrellana, n. sp, Estrella River, length 50 mm......... 263,
2. Mactra exoleta Gray, Boucary Creek, about natural size....... 262
3. Mactra alata Spengler, Banana River, about natural size...... 263
4. Mactra sincola, n. sp., Saury Creek, length 36 mm .............. 264
5. Echinochama yaquensis Maury, Old Harbor.............-..00:0000 219
286
Fig.
19.
20.
BULLETIN 39 458
Page
Plicatula marginata Say, Water Cay, the lower or attached
valve, enlarged about 2% time....
Plicatula marginata Say, Water Cay, the lower or attached
valve, enlarged about 2% times.
Hchinochama antiquata all R1o Blanco sececea-c-5 hoe eee
Pteria inornata Gabb, Red Cliff Creek, natural size. ...........
Limzea solida Dall, Red Cliff Creek, height 2.5 mm..............
Chama congregata Conrad, Red Cliff Creek, enlarged about
GwiGeyaeves ibddougueaauouanpeHAsDsusooaoDdad
Corbula oropendula, n. sp., Banana River, length 6.5 mm.....
Corbnla oropendula, n.sp., Banana River, same valve but
less magnified, length 6.5 mm
Corbula oropendula, n. sp., same locality, right valve, 6.5
mm.
Corbula cf. vieta Guppy, Banana River, right vaive, length
(OPA hoohes We Man ec AemaseaisnaaascasbooC0
Corbula cf. vieta Guppy, Banana River, left valve, length
6.25 mm.
Leda Dalliana, n. sp., Port Limon, length 10.50 mm.............
Mesodesma costaricensis, n. sp., Banana River, length 22
Nucula orbicella, n. sp., Port Limon, length 9.5 mm............
Nucula orbicella, n. sp., Port Limon, length 11 mm.
21-22. Yoldia bocasensis, n. sp., Bocas del Toro, length 9.5 mm...
23. Mulinia lateralis Say, variety sauryensis, n. var., Saury
Creek, about natural size............
24. Mulinia lateralis Say, variety sauryensis, n. var., Saury
Creek, about natural size.
25. Corbula viminea Guppy, Middle Creek, length 15 mm.........
26-27. Corbula oropendula, n. sp., Saury Creek, length 9 mm......
PLATE 29 (32)
1. Semele levis Sowerby, variety costaricensis, n. var., Hill 3,
Banana River, length 72 mm......
2. Semele Claytoni Maury, Hotel Creek, natural size..............
3. Metis barba, n.sp., Rio Betey (type), right valve, length
AS ier Th ea Oe ee eee OA Severe
4. Metis barba, n. sp., Rio Betey (type), left valve, length
48 mm.
208
208
218
195
209
218
267
266
270
267
258
259
255
Fig.
Fig.
to.
Tes
PIA Re SH
ae
©
Costa RicA MrocENE—OLSSON 287
Page
Sanguinolaria alouatta, n.sp., Banana River, right valve,
Neraverelay (eyo) seobee A secan doroAdicaechoda eeauA 260
Sanguinolaria alouatta, n. sp., Banana River, left valve,
length 54 mm.
Psammosolen gatunensis Toula, variety costaricensis, n. var.
Rio) Betev, length 63) mimis5. 5.2... .. 260
Semele Sayi Toula, Sousi Creek, slightly enlarged .............. 258
Tagelus cebus, n. sp., Rio Betey, length 36 mm.................. 261
Abra triangulata Dall, Bocas del Toro, length 7.25 mm........ 259
Crassatellites midiensis, n.sp., Middle Creek, length 3.25
TTT Aunt setleciee eal an cea ace ma 213
Crassatellites bowdenensis Dall, variety costaricensis, n. var.
IKsiakestel at, eh. Ryo) saubaol, codssacon.conbannnoanaoae 21g}
PLATE 30. (33)
Antigona multicostata Sowerby, Water Cay.......-....c.c.ceeseeees 239
Chione Rowleei, n. sp., Banana River, length 42 mm.......... 244
Antigona Harrisiana, n. sp., Port Limon, length 21.5 mm... 241
Antigona rugosa Gmelin, Port Limon, about % natural size 240
Antigona Blandiana Guppy, Comadre Creek, length 30 mm.. 241
Antigona Blandiana Guppy, Port Lomon, length 24 mm.
Chione mactropsis;Conrad, Gatun, length 30 mm................. 245
Chione mactropsis Conrad, Sousi Creek, length 51 mm.
Chione holocyma Brown and Pilsbry, Gatun, length 36.5
PTL CIN EO UA eS acetal 247
Antigona tarquina Dall, Saury Creek, about natural size...... 240
PLATE 31 (34)
Dosinia acetabulum Conrad, Rio Betey, somewhat reduced... 231
Cyclinella beteyesis, n. sp., Rio Betey, height 39 mm........... 242
Pitaria circinata Born, Saury Creek, about natural size......... 236
Clementia dariena Conrad, Sousi Creek, height 60 mm ...... 232
Pitaria Hillii Dall, variety musanica, n. var., Banana River,
Wesalegeol 07) S007 Aa cuubpeadu cjesoddousosoe 237
. Macrocallista maculata Linnzeus, Middle Creek, about nat-
ebENU SILAS. uebaabaded SIR Bette iaraaaan stan ea 234
Cyclinella subquadrata Hanley, variety quitana, n. var.,
Quitana Creek, height 28 mm..... 243
288 BULLETIN 39 460
Page
9g. Pitariacircinata Born, Saury Creek, about natural size......... 236
10, Pitaria boucaryensis, n.sp., Boucary Creek, about natural
. SUZ ET OR Man UNTER OU ee) Oana na 239
11. Pitaria Guppyana Gabb, Port Limon, about natural size. ..... 237
PLATE 32 (35)
Fig. 1. Callocardia gatunensis Dall, Banana River, about natural
oI She Han rela hea eeReR MEE ONS andl cach 235
2-3. Pitaria VanWinklee, n.sp., Banana River, about natural
SIZE a user hed ie cs eae 238
Chione costaricensis, n. sp., Banana River, length 34 mm..... 245
5. Phacoides costaricensis, n.sp., Middle Creek, length 11.50
ELUTED re sres ee ee ore ee ra Mada eae Natta lale 222
6. Phacoides bocasensis, n. sp., Bocas del Toro, length 10.00
SET ee aM Se ON rata 223
7. Chione hotelensis, n. sp., Hotel Creek, slightly reduced...... 247
Cardita matima, n.sp., Old Man Sam Creek, about natural
SIVA SHoncbdose UuAseiiBboodoued) ebsododeccemDaced 215
g-10. Chione chiriquiensis, n. sp., Water Cay, length 13.75 mm.. 248
11. Myrteea limoniana Dall, Red Cliff Creek, height 9.50 mm..... 221
12-13. Venericardia Terryi, n. sp., Quitana Creek, natural size.... 216
14. WVenericardia scabricostata Guppy, East Grape Point Creek,
aboutematunalli sizeasessesese see eaeaet 216
15. Diplodonta collina, n. sp., Middle Creek, height 6.75 mm..... 220
16. Phacoides radians Conrad, variety medioamericanus, n. sp.,
Middle Creek, height 9 mm........ 221
17. Venericardia scabricostata Guppy, Comadre Creek, about nat-
(SOREN L SHV AD nonaat aah oaodeespesosope0nq000000 216
18. Gafrarium limonense, n. sp., Port Limon, length 4.75 mm 233
1g. Gafrarium altum Dall, variety costaricense, n. var., Estrella
River, enlarged about 3 times...... 234
20. Divaricella quadrisulcata d’Orbigny, Middle Creek, height
TY Who ahaa Mie ANOLE Nas Ta RreRe CaS MCAN bonds 224
21. Diplodonta insula, n. sp., Water Cay, height 6.5 mm............. 220
22. Gafrarium altum Dall, variety costaricense, n.sp., Middle
Creek, height 4.75 mm............... 234
230 Moyateoa llamoniana Dall (Port Wim mses ssicesclelacsasiecnleile ae 221
24. Phacoides actinus Dall, Bocas del Toro, height 4.50 mm....... 224
%* *
*
INDEX TO VOLUME 9
Note.—Figures refer to the continuous numbering of the volume (not
of the separate Bulletins); Roman numerals refer to pages; [talics, to plates.
Abra triangulata 32.........200, 431
Acanthochites pygmeeus........... 170
SPICULOSUS!Recscacee case) vceesemsis 169
Acanthopleura granulata......... 169
PIC CAE eiseeesws ne sictoistie lulnaieseewicels 169
Nicimiceaycandeanal . eis. .c.eese. 151
Heicoplemmaversssccoscesisns sea: 152
muelaMnoOleucay ves.cest ss: ferent
PUUMTCeulatay vy. ee. dsauneeae I51I
Actzon costaricensis /78........ 206a
CIIDXSTICIG sesasdcea ede uBSd eee eR sacs 45
GMI CALS eas Siseeseseacnesceencnes 46
Exc SMe eee eet tee BOL 45
PAMVESLOMESIS: eee seeaccssssae se: 47
AI CTS ISR aaere stoic te cienale sauces 46
TURE KG INS cesaataeed en eee aEHE aH ES 46
DETMOLALUS ee tse eens poet eee a 46
PUNCLALUI ST oe ciiseveciclssnisisnseeess 45
PUMCLOSERIAtUS Heese nub: 45
PUSS eee aees saan 46
Acteocina bullata......... 47, 190, 199
bullata, var. costa-
TA CATIA D7 Se bose lesan 2064
Canali cullataeeveescchiecsoneecces 48
CAM Ge IRs le Sd cli 48
ThE log ooogoopbend 47, 190, 194, I9I,
206b ;
triticum-tritonia................ 194
WIG WORT Shae Bae nanan AN ey 48
INGeorbis) adamsile. ss y.csesuese sees 159
SUP TATU S hapa wtee eee ee oe 160
Admete microscopica................- 75
PNESOPUSISLEATMS Ia ere neiee iancen secs 94
PMA a LELVATICOSAN ee rcecaseuscnesel: + 138
{GDN Gch NE RL a SME 199, 325
AulaloiiavadiarlSitaaesese see wen ee reeas 127
Cenithioidesye ste seen eae 128
NIC CEMOMVACHEUS Meeeeetneace cy seeeaeee 89
AD ae inva U AN a eau 89
Ehoall oN eA DIDS 55a rlonsboooaqedaosHeHodos 89
Abo a Dl eye bhTo yn oss Gouoomseonddoooaos 89
DeatimMOntenSis daeessceeee sete 9!
Caididissimiayserns-cee seen eoeee 89
CONSENS USH eee emo rena ans go
DORE SN OISV SI NG aa ee aS UC 89
Pal VEStOMENSIS eee eeeaccase 90
OUOKSSISH SS Thy 344. ndodghign InoueconSoses 90
Losqidemodicay wis ees. naercuces 195
losquiemodicajer2) eis cen. ss. 295
TMCLCN CAN e Nie tees. cee caee 296
ObSOLSUIS HEN eelnes west acee ae sie go
LAM TL CUA 2 eee eee 295
(EBUEUNWESOUIS) cote crises sodoostoduaacos go
ball oreo. taal A SERB SNA fab NARS 89
Auluimy Bilis erteshe meas seee eee see 202
7a (op: Fe Hea Aa ae a aaa HARA Ma Nn else 57
Amanra tombigbeensis.............. 26
Amphiperas canadiensis........... 112
Ampullaria caliginosa. ............ 141
GISFOmRSS ss doobeoosoodconsuodunses I4I
PUN SA Ey BNA Ny an 141
Amusium bocasense 20.......... 378
DEBS PA eA ae acoA sa ANAS aa 200
Wy ODT EUS dae ce sauceaieoneuaal 187
TL OGLOMIG eee eee 200, 377
EO Unil ae is Si SED leon aac a 186
Anachisiall bella ne eile, 93
Bhaeoyalabiy sll. cod sadoocodoodseeBoboLe 92
290 BULLETIN 39 462
avara, var. Ssemiplicata......... 92
Ware Sim tlisnyees see meeenesees gI
Vat jtrams linatassd seed ssenenesee 92
cancellatayy iia ue sea ninnane 92
Corel Wes oleh 2 Wea Re A aa Sa 92
un Sie Eig SoH Eee 186
ihoOtesSseniatnal wey eeney. sneer ns 93
Obese a rae a ees be un alie 92
ODPM) hod edocs nc ooopabagdsay maoadoHbond 92
OStrelcola ieee Weta 92
h OP CEN Eines eR M Ean Ns Rue TUE 93
Ancillaria acquaensis 70......... 266
Chipotle ays iN Meee eu 267
| OyU USA DUIS MP SERS NAMB USE Aeneas 267
Subslobosay an haiaene 7
Ancillopsis tuomeyi /7.............- 12
IATICISEFOMASISS eee eee neato 158
Ancystosyrinx dalli 7... ........ 232
elegans 7a eee eer. 190, 232
Tawa talent sue eA SEEM ABE 63
Anomiayeplanp pile seeeeece cet 381
Sim plexeeeeesesee 189, 194, 200,
DOS N27 Os SR ada se nee RUnieie 381
Nan 2 BSP MGB AA ea Sins ane UG ARE ROR Ep 35
Antigona blandiana 33........... 431
lean oINehoeh jasoudonsencsdadsesuedue 188
Marrisianayiyeemeee essences 413
multicostata 33......... 189, I9I
AII
TUL ZOSAM GRR ae esacaaiaae 189, 442
tarquinia 3}...... 196, 194,
412
Aphera islacolonis zone............ 193
Ja\y 0) bASN Wee Mico Gon Ao Aabdebocnucaubonob dents 53
Arcalactinophiora 26 lier s seas 357
auriculata 25......189, 193, 199
362
CACTI CANNER ee eter ncas see ssa nteaes 362
Candiday 25) tiareaessce. 189, 355
Chinigiiensisvysecsssersnacscse 193
ConSObrinan yee posse seats 358
Costaricensis 25)... see: ee 359
darienSis! W25ncsesese secs 187, 356
SalI CASIS me dee eeeee eeeee eer 356
golfoyaquensis...............0+5. 193
golfoyaquensis, var. medio-
AySoCAiney’ | AO) ssoecoccadase0e 360
STAN Gis wy hits eae IQI
henekeniy 2722221934 OO 258
Wad S27 eee owen yaneee 365
LoKOsoKSINISIS: Abyatseodod ohandennse 359
Pham opeKOr EAN io) Gaesnanncddocdes! sauce: 354
UK (0 by “haa AeaMeee Aon acco Aobede 364
macdonaldi 28...187, 191, 366
TMAWUYeS) |) 25.25 eee AN LOS
occidentalis 252...) 44" 189, 193
199, 353
OOM TENS sadcsesosaconccsogascooc 356
Oza KENSISWMghe eee te eee
ON OKs CIB BAL ura ha ad uRAAna sclatc IQ
DALCISK eZ 7) meee 187, 364
LEVETSAN cua una uaa ncbUnnn mee IQ!
SObrimal iieshqiaenaaalaeneeere eee 358
umbonata 25.....189, 193, 354
WEENUO ON Alo Sho csacuaeeecouaeace 261
Architectonica granulata 79...138
190, I9I, 196, 200, 326
perspectivaeeeastee ee 138, 326
Aspella, var. obeliscus............. 117
Vane PAallpereulaseaw ese Ree eee 117
Scalariord eswy ene eee trees 117
Assiminealaubertaman.ccescussteer 142
Astralium brevispinum 78...... 190
334
(CHS ENGUIID d oooac Shonen sadose 190, 334
Astyris) diaphanaeens ssa 94
dueclosianmay (ewes eee ease aes 94
Tuatha eer ees re 93
Atlanta perOmtteneee-nssseeeeee cess: III
Atty S TObUStOId esuareseaeesscacy Meccsees 26
Ait ClANCOTON ane enn ese eC EEE EEE 56
Oe ee Ne Pa 55
PEllUCens es Mae eeesoasseecee 54
Auriculina insculpta ................. 46
463 Costa RicA MiocENE— OLSSON 291
PMETACUIASEGUEMY V4. oa sececcaccbeesenoee 54
/ANEVTTRIG, GLI OE mendasessnaecppeA cauaHeone 82
FINCA DMA sscciee secrete cea escis 3
OMUSLAL es lose callessaisio ns wesioceee 8I
AW ACU ITMOTM ATA n+. <+-scsscccse 367
B
Balanophylla haleana. ............... 27
TBiailaonientb tad a5 ae ae weer ae Ame 43
Barneamallatovdeas. 2.0.0.0 /. 5. 7
BaASwliSSalalttasccse cs s-0cesee- 157
COSHUMa tae sees ccoe cle sssees sses 158
SUPE I Darneeni see cucew suse oseees 155
watdelicatiuilayesay cms ecel. 158
WEEE, GIG DIKESREI A) Wocanangesdocseee 158
BaSterotianypraima, fe 20
Bathoxsyplalistaests. 2 vest svc alate sos 38
Belawinlike raven anon) seavane ney cua. 68
Betanodolinnm es eh we bE Il7
Benthonella fischeri................. 140
SAAR nnMea HLL Mee Va GARIN 140
TAVISKOVED ISS Rau ema a NEE See Gan 140
Bitinaemnboiplexs ese ase. 128
Sint MIO ores sbecdeusuacede: ebake 128
PaAlweStOnMensSes ny ve eee 129
BAUS DYSeEU Baa GhAsconadeadonesneoner 128
TEN CATTT TT AA eae Eee 128
HAUS ALO TAUTAN es nase se ee seeise seen 186
VIANNA soe aes al esata hee eae We 128
DY MUCALS CATANIA) eee ateisiefoeeiels leis 135
ISYIONEHA, GoxXGsc@lsconscdnaccoouseseuesue 132
WBA TT ral ease ee eat NL 56
SLC KOCIitaM ensues ovis babe Meee 57
jOCINeHGlay San Sea sAbtae ABE) Bee uonmemaE 57
IBOKEOLTOPMOM: 0. ssieeeslsee ocean oe bee a9
Borsonia cocoensis S.............- 250
Bothrocorbula viminea............ 442
Bowden beds, Jamaica............. 199
BUCCIMUM ODES. no sass sesesoc ace: 92
OT DIT Sayles eres ene uses 87
arbre) ae een AN ke 99
ZO TS aerate nen tauan ante eau 94
Bulings arora y eee eae ates 55
I BY DUN ke WORN eee LH A A Ue ea 48, 49
Di bhica tars pnd aw Weal ae aaa 50
canalliculatame weceunrarenceueetee: 47
Pawperenilaeemesceee keene 206d
HEC Cer oe ae Han eee Nene 206a
Bullariajabiyssicolas. |... sssseees: 51
lanl AeXGIe VE) es seddan Baccioass booted 206d
OCCidental sy eae een 50
pauperculia 7...194, 199, 206c
PAM SUIS CUA se wt ae eA ET
Solid any aN hn eee ea 51
Sta bare Ngee marci oll lat iin eral 50
Burrs al arta as eevee maa ae gen 114
crassa J8..... 190, 195, 200, 306
Busycon canaliculatum ............. 85
CATA CA re MUU AN Ab §
Sli CSA TIS AUN oeE Me ve ae 56
DELVERS UAT sie Weve eae eee 86
DY AU ee Sous SaSdobRSoaHoseedes 55
C
Gaduilastanctusiuan cou) senna 4!
AMT AIC Sue HOUR UN RRINS LTH 40
CAMO EATS Seen racen se ta eeak Huet 4o
Clo meas he Ak, Cee 40
MTA OTA UaREE Nee UE neh aN 4I
PA VSS es ee ea ae 4o
POC TH NO AE oN 40
quadnidentatusme: spsepecenees 40
HErer HS TOTS inate eens Al
Ce SiS eae eee 26
CzecumWvannlatume nee 130
bipartibuman Sarees tie esc: 130
Carolinas eee ee eee eee 130
COSTAE TAR Gre eee eRe nee 131
STAD RUT: Kare ee ee 131
TAS PIAL BUDTTD eee er eek ee 130
TREE Ql aera Uae Ae Ms aka ee NNR 130
TORT Cinna tls Lea eS con 132
poeta tara eee ees 131
292 Costa Rica MrocENE—OLsson 464
Honida mums ee yee saat 130
Calliostoma asperrimum........... 156
Castile 7S ee Na aU SE 337
CIFCUMCINCtum yon ee 154
CORD IG TU HO Beene a 155
Cushy ptusiy cou eeanawen aes 154
TLPU AUCH HOWE we NN AA Ne 155
NIMOMen Sighs. ees ena 336
Mmaneinellan 73k Menai 336
PUl Chery NM ee 155
VATE AW SOM Iie see aul e eens niee 156
TOSCO UN Cee Rea a eased 155
CATT PD AS TNS TSP ese eee Mean ert 165
DEN EEN He LE es es SAL MSs a 155
VESTER SUNNY NUMA O atta 155
yucatecanum...... SS MPa are 156
Callocardia gatunensis 35....... 407
Gallo gazau Zee Wns uay Me) Wiuaess 154
Galyptreea, candeana 22) eee, 145
JOE WENVABU NTIS Os Sonny HacenedcaoosadaueD 145
Calyptraphorus valatus, var. com-
PLESSIS Scans Nee Uses aw ae ea NU ee 26
Cancellariaagassizil in oa 74
Gell cea ee OC MIA A AUER A IA |
barneth aig aes an 200, 253
LODE UU Fei es ee a SIN te ee ROL 191
CONGO Haima Sau Manag ek iets 388
Clavatunl apse Gee ate aay 74
COnradiana wen SE BAas 74
COSSIMATIM I OW eee a anes 253
COSC ba EN AUN Na Sei |
darienay rom cananune Sos 252
Epistomifera wisi eel O5 255
galvestonensis................008. 74
gillbentin i 7s we Meni oeiTi
islacolonis 9. ..... 187, 191, 195
196, 258
ELL NUTSV ZO eens nares 195, 254
TOOLSET EMM aca aN an nanny 255
DATIOTICS yt ey ANNO Aalaaoe 7
Plummet vomey ee eee IQI, 257
FELICIA ta ae ae ae Ge 74
CON KEUET)) @sdagnacssoosonnbos 195, 256
SIMPSOn i ewe 75
[ten oar gee eR Se Annan MEE 74
tesselila tame neeseeeeeenes I9QI, 258
MOSKOS | O)esaussccgodacen IQ, 256
var. trachyostraca.............. 252
tiberculatayn iiss) anal aNeE 191
Cantharns limbatus...:..eeen nee 88
ORDIQMYA as... coe 87
CINCH See 2), eee eae 87
Capulustintontus.:,..ee eee 143
Cardata caribbeanensis 29...... 386
laticostatal i... l.e semana 387
TACIT Ai) 95.22). ae eeneene on eee 387 |
SCAtD ia COStatar yy sseeeeeee cea 388
Cardium, var. alternata............ 189
APAtetICUmay. Whee ee ee eee OG
Costaricanuml)| (GOs... eee 398
dominicense 30.........194, 397
fap b ab bar UR ARC Man Se REM S RAINS 186
ferehobhalerelS,pagunddoscoasoces 186, 307
JOKESRSONMDbOON! EXObsadodsaonddn Boodss 399
medium 30...189, 194, 205, 398
Serratum Ou... £94, 205, 400
JOO TOS I dota 189, 205
Webeg WUNAKONON: | Or-cdccooncdecooos 400
Stila tuna iy Ones seeeee eee 188, 397
Sublimeatum sass ee eee ee eee 201 .
Gareliopstsie i aMule Sei ees 110
Caricellaldolitas anne sun a eames a
Caryatis guppyana........ ......... 409
@assidaria laevigata. ts.-:css0s5m 308
CaSsisicaileo eases eee eee 118
Alar imea eau eu gec maaan 307
IMA atay NN) seach eee ee eee ee 118
podYOVAHU NUKE loa donudegnacoodsoucsH4cS 307
LECTUS AV LN Oe 307
Snilciteralanwenauaaaes 195, 200, 306
(HSI OTOBI EIS. Aas oadoddcodenanuoossios 118
PuUberOsal ee Ae eae 118
465 BULLETIN 39 293
CAvolinaniniexay vee oe 42
TRS PNMOSAN chase ue Us sen Uae 42
ITACHI ALAA ant secre et ait 42
CercadoMtornmation. (i.5...2.ceccosees 193
Cembmicdeaicostatal..../.2.-....6.- 126
IS RENIIONG nee n ances nee Ge sEaHaRe 127
Scallanifonmis...-...--:-.-.4. 126
lI Ietal beeen eee oscienindcs ooaseoes st OO,
Wag COSAME ee esc. otes, os senn sss 2O
Cerithiopsis burkvillensis.........123
Chy Stallman oe. ese ese. J 123
CHIEKSOMI nsec ie sees slain uote 124
EstelllensiSia esate scene aS
BTCCM OW esse scan teecessoeeteeasts 123
TED ETISTIS Gootadcene ce a ranee eeBeeee 122
Sto mlatarecnesecocsee, seems. <4. 124
ai TTA sskekss. (cese se 123,
Certthivm) algicola ...3............. 12
COStATICEMSE| 7 e oe s25s 52s. 315
dislocatumiye es eeetetcnsceaeoes: 59
GID TIFT suooaudeo onoddbnoelees 125
Ord dani tbeeesen eae ST OM
PialWeESLOMCHSE! cre... 4. 404-402 120
RECN e sesh sn ldeck vanes t 123
Inte SreaTEwTT Gos souose. “Goddoesecee 125
THATTHPTNEN TT neooobesseue sskanpeBsne- 125
VATA SESCEMS 2.25. sac sascnes- 125
IAT CANT yee atseis eee ew eee 125
PROLEXstT TAY ecm ia-tas sisson ace 59
PUNCLatwIME eens seeeeeceeeeeee 124
tencbrallasmpen sun cesee ceca i ueeune 12
ATS EDLOU Se eee alas aia ie 125
Cheetopleura apiculata.............. 168
Gama antiquata. is...) 0.0.6.5... 390
ACIS acer mee ne eked scan tc2k 391
CONTE Pata EG eee cme 203, 390
congregatoides......... We ae 390
CWhreileatequestris...22.6:.42 6225.5 .5- 143
PME CEOIMTAM eee eat le oes eteE 186
(CINE STHVoST ATES a nsnac Ser aaa a ESnE PERE 108
Chesapeake Miocene............... 202
Chione chiriquiensis 35.......... 420
COStATICENSIS gbnestenossseaee 417
SUSU ZO mr eee eee 189, 205, 420
holotyanaly eopiee- nesses ses 419
MOtELENSIS ig sanecencesMensesseee 419
AMACLTOPSISH NG Feeleceseseee 188, 417
THONG BGssoocudodornnsdce 188, 416
Leg uillrmma eee eee see e eae eee eee 187
NOWWATHEI. sodoabeacoos 200, 201, 417
CPO Tees LLU nae tee uni 168
MAGI OLACUS meeeceseeseeeeeeeceee 169
tuberculatusieeeeseeeeeeeeeaee 169
Chi cores eas oan ta iibeaua: auatcinae 95
Chlorostoma costaricensis 78...335
C horistella pompholyx ........... 161
Cicculuspliratusyeeecee eee eee 160
[BCDB D-CA ANGR ar RE AAU HRN te 160
Clathurelilayy ewettitnes.cs---ncessee: 70
Clavatalaywelaiigeniaeseeeeeeee 317
chipolama Le anes 126
COstameanayai7ige) wcoseesess 316
Clementia dariena 34....... 187, 404
Cleodora Tanceolatal.:. 200. .sse eae 42
Cliogne curva ness aus 43
Dy Lai dataleessses estes eee 42
Cochliolepis parasitica.............. 159
Striata cular seas een cee. 159
Columbella ambigua. .............. 298
IMEDCAtOnan. sere. Aslea eee eins 93
FAWISROKCLONC KES) co oagnoooncocsudoaqosoue 93
submercatoriS 73............0 297
Columbrariaytestaceaynseessee 115
Comipressidensi tea tues 39
Conomiitra ee Aue aes 82
Connsiicemiulators ween enna 186
AGASSIZ Iv. ehaes eek aun 200, 226
Bhacly ol awieE FEES, cococabocodosnesosscoNe 62
Der salisite ee eee een eee (eseun 205
Vat DOSCEMSIS I MOl-eereecsee eas 223
burckhartiviOmne ees 200, 224
Catenatusrsn cca senses rt neasees 215
EEMEMTT Oe as ate eeeemn eR yees 61
294 BULLETIN 39 466
Conus)ichipolanuss.3 62, 217 tortuosostriatus 4............ 221
CONCAVILEC tama vee Meee es 215 NASH Ob Ny eae a ranma uuabeodoradc 216
(LovaiKoloyGbaels) soy) papeuoonose 186, 195 Coralliophila abbreviata............10T
var. costaricensis 6.......... 217 debungiae eee ae eeee ee eereee IOE
C6 Fe BTA MUN IE eT NA 217 lactwealy akin) IOI
delessentiamusieenss essen 61 Corbula dietziana 24. .....189, 440
demiungsuse een. Lon TOES 216 dispartlis ye eee 438
domingensis#ay- ae 186, 195 SHAN HOYOS OISTS Asjeacec Gadde coooases¢ 186
SENSORS, 5 sAcldaonosdaonoddon aA 61 hexa qyrmmal2 7 ee eee 441, 442
floridantarics. hse tesescere 61, 189 SHOUD UT UDIC GRAD ARM RE MARAAAA iol oeoboce 25
BAZAN hen NL Wee Runt 226 oropendmlay io iene ee ee eee 439
gracilissimus 4...195, 199, 224 OLOST 1927.66.22 ae aen eee eee AAI
granozonatus 6......... 199, 222 Serica dike iy ae eee 186
RAT TISIWMOvN Ie Rey aes me eee 225 Splenisss.20.s oheeae eee eee 186
hay tenSishe ease ee 186, 194 Subcompressaii cee eee
TOR PUBEN KONE yes caemaoasgcocauadd 217 WC LAR ITAL ti ane aenune 194, 438
interstimctus) Oi. .e) se. 199, 220 viminea 37...194, 196, 200, 442
[EOTERSIS eee Nee ene 215 Costa Rican Geosyncline ........ 205
Nexo yah OSL 84 ae Nenee a eUG anaes 61 Costa Rica Stratigraphy..........- 181
laVOTMeHSIS) USe7 ee ee 188, 219 Granopsisiic.saiceascccusecesceeeeee 167
marginatus) 7..seee 195, 223 Crassatella antillarum.............. 384
ANONMS (io. eoecoeaeetn ena 188, 214 Crassatellites bowdeninsis, var.
Varimultilinatus: cess ane, 200 COStATICENSIS HG 2t enc eencaet 385
ml tilinatus snes eens 195, 201 Fo712)1 6) OVW GAMe BEB ARE SoH anBeo Asbo sGb6s 25
multistmatusi ze ence 226 TIE CACTISIS| Hg Ze ea een cies e eee 385
HMSACTISIS qibuicsNaelie sus cana: 219 mediamericanus................ 384
pealiee enw EULA STU ORC UR an 62 DEEVE MMM Za entee ae tent mene ne eee 384
planiliratus 6.....188, 199, 222 Sepulcolllascessesseossereeseseeeteee 25
proteus 5......61, 189, 195, 215 GCremiidesy\eeseeesece secs emcee meseeacee 16
ON ASHITE DS aa agadscboeaauauducseedans 62 Crepidullayaculeatae eee I4A
pytitormisy..2s:-4ee une le IgI, 218 COUDVERaN a rane nnccicece nee neete 144
recognitus 5...... IQI, 195, 199 POT Ca talks slednseseeee eee 143
518 Plana eases 144, 190, 199, 331
GEA TS ees LuSe an eee nese 222 Crepitacella limonensis 76...... 315
STATI deat aue mele 194, 220 Creseis subula.... cilia sieuiateinaleeeaaaetieet 43
Solidi i or Crucibulum auricula................ 145
RVers PARAL a AMON eE pPlamatiwins os ewesces esses 145
SUCA TH SIN price adsaeecuese ace 62 :
Scutalla tums eeesaeeeeeeeeeceeeee 145
ene Shocliseh, Gh, “Gy Straten ma eee Cee a 145
i Ua Cnyptonaticasesasse cere eceeere 147
BROSHS I )eetn eicosccooa acca: aha Cucullzea macrodonta............... 25
tortuosopunctatus 6........ 226
SatoOcday: MEAN ee 25
467 Costa RicA MIocENE—OLSSON 295
Cuvierina columnella................ 44 @ythexeavaltenmatayensescuscsses-ta 408
Cyathodonta gatunensis...........383 loleraslianaes -bsdnane sadagdacacadac 200
Wy clostrema afiine,.../......56 1-144: 161 daiienaly a arseu seme asneaetinecse. 406
OMA AO Vex, seco enece yess LOM HIEDGKCKE) Cub dWansnadudennonpoocodoue 408
HS etl OS UMMA pel Ws loyslfelee sjniecls= (sles 160 CAnChadiMaeeccostseeeloocsesblarss se 412
ERO CMOTG ES ee eae eees sees eee: 161 D
Cylichinajbidentata:..0) 5221/24).205€
dacs ns : Daphnella hormophora.......-+. +» 73
TRENT Ss soso NS GRE EE SRS EE POR BBB 26 leucophlegma..............:2000+- 73
Cyclinella beteyensis 34...191, 414 Atria Claas Ae nat aaa enemas 73
CPARIEMENSISt tS A) ehe aN. 187 Wehr aia clay clay taemeauensectdetseceensias 72
Hea Oyy, CUI ees seie velo sy sis Saison cise 191 perpauxilla ...............4. seeeees 72
SMMDEAG GALA =. +...¢seraee-senne 191 Dentalium antillarum...........35, 36
Wate iguuitanal) 34..6.5-..c00+4: AI5 barblebeivmeeee i naeeeeas elev ucts ce 36
Cylichnella bidentata................ 50 bocasensis 18.........s.sss000 338
var. galvestonensis.............. 50 Callipeplum............00-seee00+- 37
bidet CUIM=trAtOMIS4s.-.6-2-0-- 22: 206¢ Callithrix..............seseeeeeeeee 36
Cylindrobulla beaui.................. 52 CITES, catobobacsuobdaokooscags dbaaoonos 36
Cymatium chlorostomum ........ 116 CSiAshH RNID \codoscasnIpgaadbboauOGuoUde 35
CHy/MOS) oO mveY lubaols ear boncabosaeacne 116 COMPTESSUM.-.. 0-0-2... -eeeere nee 39
STA CH SMe eM iy 116 GidymMUm. 0.2.22. seseeeeeeseeeeees 39
Olearinmy Nee ee 116 Cisparile.............cceeeceeee esses 35
Cymatosyrinx centimata............ 67 (2) DYOFMENEN Col sw osodesdonoconaadouLaHoOGNS 37
{TUOSETY, SSSR US HHUA RHO ABHa EBON GeneUs 67 ENSICUIUS «00.0.0... veeee eeeeeeee £8
@yphomialeibbosa...:......6-...4-5; 112 HUM ....... eee eeeeee eee eee 38
Cypreea alminatensis 75......... 312 IBKOACIEMES,.. Nadsenosnoccocobadnous 339
CHMEKEA 15. scdecccovsesses D134, 312 EXOTOIICLET s-.cdunodebsenocosungacaceooons 34
dominicensis.. ............ 196, 312 laqueatum...........0...ssseeeeeeees 34
ESESMMEMSIS. | 12... ¢5sciensceasiensee: 14 Te ptuma to coe eee enon o-noeesenstene ae 37
ERAMUMETMAN A). se snsscaknerees 113 MHALATA, ..........0eseeeseeer eeeeeeees 37
Wemelkenin guile 186, 196 mie diavAdenSe ss sssaeeee tse: 26
DEIISATTT, Tel sasanealoanedeoese 311 <n ESSAI OPOEMSs ss coos gqgoaadcacosBes 36
BALE esse ck soles. vats AS OPhiodon.........ee- sees reseeseees 39
SPURCArestnetecesstiaccedaccsecs =: U1 perlongum.................sseeees 38
Gyiprde CASSIS a skesceecscscesedeaecersess 118 PTESSUI, 2.020. e eee ee scence ee eeee 39
Cythara heptagona ........... 186, 195 pseudohexagonum.............. 35
terminula, var. costaricen- quadrangullaney ese eens 39
SUS ONE EME Shon It 249 TEDECCACTISISH Mn Mincneecuenes 34
Cytharella cymella.................... 713 semistriolatum...................- 37
@alvestouensis. oe ea. 73 S@mi Ca tutes aaiecc nanan nanan 37
irr atiay i Sereee ener ae ema yn ana: 249 sexangulare,...........00065 0.2. 34
296 CosTA RicA MIOCENE—OLSSON 468
’
Dentalium’sowenbytsceae sae 38 Carre Cay7 ii icnecen ele Maar 243
Vat wpellicenttee eee 38 Chiriquiensisiy Sessereeeeees 236
ey ay SLUT ee Ue hiemedaealnbe saree 35 Co) oly lB reper MES MU ESR Sa) enon aD 72
tetragonum......... secede eae eae 39 CINCH Volcan teense eee ooate 64
Weigh nabboolyA ea aie ye 34 CONG aby CT RORAMANne nts CCN 237
Vaio OS tT gua wenvea sta 34 COCCOS SL ASIN Lay eee aN 236
uscarianum /78.......... 187, 338 COCOSIM Ay Sass. see eeeiec ee 247
DD emitis by layin o vane ee aNN Mita | MINE 156 Comatotropisy G4 eee B72
Detra clay ae OA NO Ml ieaninn gs 56 consors 7....195, 199, 200, 233
Dialeniawe eee BLP A eu AM ea Eh 42 Cristobaligy (6) see EEA a:
Diplodonta: coblinay igs. seis ee 392 dalla, war. cestrota.. ose... 66
PSU Aygo eee Ce eee 392 Getectays ye say te epeeoenneel: 64
LED VU Se Mees ea ae Re a 7 CS] DYBE che AREAL SABES clei at enone 65
Distortio, var. clathratus......... II5 eliveivyaly ee Aoi eas 72
PaStuMensis eee eee saseee 306 (SabaeSnKepaskel 4555, suscdocoonqocoues 186
HELICula tastes eeen any 115 estrella nay 77 eae smseee eres. 241
Stet ira ae seein cela 195 CXASPeratalyi essed naesnoeae 65
IAS COnbrioxs) sama einai a II5 S:ccfohallh Oy tuWaousdoneondnaaeusieadog 73
Simi biases eean yn auen ie 305 eucosmia, var. canmna........... 64
Divaricella quadrisulcata 35...189 ADIETHEONPTOHIS,, bean coagoqoou.cseadoood 238
200, 205, 396 BUSHING MMU eas Joie evar 186
Dolton Tee seianeatane 119 PALMMCUSIS Ey -cuseece eens ce 186
Caminray ee DO es ALU AR ANG 196 haliostrephis............... 65, 235
ellipticals. au nia NS as 196 AVA CHSISH ences ss caseeaeee sas 67
ellipticuan resi a cee evan ie 311 HORNA econ dacadobqadonobes access 73
Donax acutangula 3? .............. 22 NOPKCOVT EY orsnocopancdennscebaoobeD086 64
Dosinia acetabulum BA LIS) 201 sbanVevaVSAISS) | Chapnoscoodoaccoosa0. 243
203, 403 limone tian Sac ss-escnse eco 242
delicatissima......... SHOP eee 187 IbbsaKosekerey Oi:7Ap se AReenbandgendacooos 238
Drilliavacestran soon aon Lak 65 LiSsotropisys eee Heese eeeee ete 66
albicoma: tN oo ce2 wale alMaenie 64 lathocolllectayenensssneseeaetstast 67
alllaaod aitalyeren ee uMn aoe sane 65 = lithocollectoidesiioeenrcsr sce 246
alesidota, var. macilenta...... 64 TOMS Cae eee aseashcsasenisciseses 7
AGUZSTSIS) (iSene ee euene tee aaa 245 TT OCTISIS) Sete Un aneenaeuaeene 235
AGMATILC AG San bie aiiin Ulalananlns 237 SPOTBISE), | Gibcbongadopea ueddooesoebooo. 246
AUT ATLA Son nue eee nema tay 240 SHONBISHNEHWOLEY |! labo aadgouabacodonodee 241
LRN aKOKSI a ea A IP ay Ie 72 MM CleAtan else ueenauetesen see 66
bOcatoroensis/iSeeawew eee ess 239 OHSAS nooscu copgaspo00 ‘eobsd00 64
IDOSCEMSIS) MiSinti inne we uee ose oe 245 POR NEROXG ABU EY“ "GoocaddacodonasesoesKsod 65
var. carminura........ ate selma 66 WN ORME | 7/edoasacogaboanba09). cade 235
469 Costa R1cA MrocENE—OLSSON 297
Dida pellagial wane cscs seem ecwccecle 72
MEUM LAS Ase e ca stetanWastes eae 71
POINT CT I misane a reaenGaeebeses coor 65
SHTT/OSOIO Wegasconsonscaboeosea adscos 66
GIMIDENG Eis easepaaheaaaseaborsee se oasacs 73
TELEK = sa pc bBo Re Ieee ae IS roaare 66
PAEOMLOM AN nOsaeecesiensaeceeeaee 240
lelicclleeisteectcloutiere vvislecissiciveece ceases 72
ETS ETC Au en ese ssida as traealiniekees 65
VAUIMIMSET I vecsceieaeiaceccsecieet 186
\/SioS82), -Weosncmnncos 195, 200, 233
aC etlllllmaemetitere: is Scceclome secu oases 66
ZOOM is esene scien ssiccveneans 186
E
Kehinella nodulosa.................. 137
Echinochama antiquata 37..199,390
fccrebny dl erimiaseeeses sce ance ce se. 391
yaquensis 37.............194, 391
HI SCT MMA enna Weliavnseaioveces 24
Egerella subtrigonia...... ............ 7
Emarginula compressa............. 167
TMT APLAR A Neaats aac tai 167
Ey mibolusmmiblatts...s.ce.s.4-+.-csesceee 44
Epicynia multicarinata..:......... 161
ES MAS IOI ees cee sare nacneine Woeces anaes: 38
Epitonium angulatum.............. IOI
CllaiPairbbiel Aogasansnseneananeeoseer 102
COC eae ess 103
contorquatum.................... 104
Genticwlatiumy se eneee cece 102
FAIVEStOMEMSE Ms eee ctmee sesiee 103
IbboKSrN EB NOO OS GanncconeeoonceetaeoBBe 102
EOOTKC] EI bl 3550400 nesacuscacbenadonn LO}
FTNOGIES DEO \aunade) qacoqneadcans 104
saeaikailbbowbierqbbsst Wfeodeoneacosdaso II
TMUNGIStriAa CIITA sae aseeese cess 104
AUIS ELA EMM 7G eee ee eel IO
Toute EN WoT os eScHoooandacaboneceoe 102
GANG oO vtagdooocecdeooaneasanees 102
SCIPIO tn udneeneueesouseisetss 102
Seni Ciiilawcayeeercms tee eeaceu aud. 103
Suiba Crit utraleze accra ees sctines - II
(WEIENEIE, sodcuscusbbs sasgooons onaes600C 102
CUCINA e sl eseeeeeinesce cians 103
EM atom lanai Stracscun coccercscten ses sete: 114
OUNCE pao saaqeearibusoocoodhsq00 114
EVM ge mi ti Caw ag ccsenssece-ee eee 21
DOLICAN Ee mayictad saan seems bacceceels 22
Mthaliameclusarnc.ys.ccctees sese sens 162
SONG coeocasasanancooodasosoeedoac 162
SUPPRESSAbeesceceteteccnscteas cea 162
Ulf ail anasawars cements sclatclssclsecisesinecissces 73
I aller a eas See eaten esis ciate 105
Eulimella unifasciata............... 107
A Deb aah cel ea Naas SAN U A Nea NAD an EN aig 124
Eunaticina semisulcata............ 151
Hupleurallican data ssss--sscssesnees 99
var. sulcidentata........... 99
AUG CILOLIMIS Sees esse eects: 99
AEG S Oh ez ease dbcrecer eed sRaeHadeebenSeuocE 147
TENE RECOV ATE 4racadecoossoobecoosouadsae™ 155
cilia percracCiliS wancusmsscssaes Fas Of
Fc
Pas ciolariatdistansy.scccnnleccseseees 83
PAG AML AM wacnesei/eaaceeeeeeeresee 83
gorgasiana JJ........ JeCobSoqoE PG]
macdomal digas ae asses 278
MeClaian aye saiasewcsmeceseaetest 84
(qEUIBGOY). | VCE odacocsouson 83, 205, 277
Havarntiacelil ml OSaecueccssceaseasessees 96
VaTalevi CUlassuesnssseen ce 97
intermediarce sss PA Ge 97
paWBKOYSNNISY CuarcuacacemosbadoN GosSsaubS 96
Jeb Geriwtesane=s bebe Gogseepsnqgououosoodeaboe 36
F issurella alternata........ 166
ASTIIGIANMA Ms oseueceseee se cee sweets 167
barbadensiseeeeenseaeeeccsecees 165
Fissurellidea fasciata............... 165
Fissuridea alternata.................166
Galy,CHleNSiSweenuaecsencenceaeesnee
ISSURISEPtamacseamaccesrneesteccseess 167
Flabellum conoideum............... 235]
Fusinus cepynotus...............2.055 85
alcimiuse ee deteuasicccecince 84
AIMTATICUS eee eee seee cee one seioae 84
298 CosTA RicA M1ocENE—OLSSON 470
COMIC ee ME iat sabia 34
SUCOS TITUS ay wena een alee 84
ELE ESS UTS NR SUITE ante Fan nts 84
MO TIETSISI MiZpZeee en aera naa 276
MAM OCOSIAINS pT eeeae nye eee 275
HILO Tee ace vue cae ea Beane 26
OLLOMIS ese yO ieee 26
CUEKEON IS Hee renner sass DO
eSehla tise etc w sees Died 8I
(MOY ABTS NC AeA Gs a 26
G
SNOB oe oese Gacueaoes cedee doce uadocdoon 4I
Gafrariam altum, var. costari-
CONSE aban waa sre nenanee 406
IiIMOMEMSE! iz 5eteeeee eee 188, 405
Gastrochzena ovata...... PN Ag 433
EO CULCa eaea 200, 438
Gatimistageieiey ease ee meeee 181, 162
Gazairfis cheney een aes Ba, 153
SUPER Da ees Waetae Taiaccuet 153
WALSOMM see oeseo ses FA aia £54
Genota didyma........ SEN Oa 63
TATE OA ea eae ae allanol 63
Gilbertia estellensis 2.........13, 26
Glycymeris aviculoides...........0 25
CADMAS Dies eee sda e ae 349
CarbaSina)i22u eee. ahtesihes 351
CaStaneus 270s We AT OQua 52
decussatus 22...... SEH gaa 352
Glycymeris decussatus. .......... 189
jamaicensis 22...194, 196, 199
350
lloydsmithi 28......... Seis 352
EDtlObICOSta Use een eee 349
Glyphostoma dentifera 195,190,248
(27210) OF WERE Soca n CN Ana SAu A hag ah 69
ES HDD Pe ae SRN A «269
aaTOVHODLSE! opeo MARU neat e 248
Graptacmien was we Ieee ee ey er pees Bil
Gratelupia mactrop3is..............417
Gurabo formation................ aOR
GyGiT Swi eee aes eae nae 115
|
aliavamericarialya 7a eae 251
HMamineavantillariunm-s sss. .e eee 52
antillarum, var guadalup-
TIS TSA, Sa a ane en 52
Cle sia SR eee eae Annem 5I
Sul ara ae) 5 an ee ee 51
OYE ICH se eae EME Hauer KAUR 52
SUC CIMe ANU aaa Eee 52
VARESE OTIS Heo iC: skis ee a 52
FTAUStA LOT 0. os 2a sel eee 134
elie depressus) see eee eee eee IAI
Feteroschisiiialies ss heee eee ee 36
Ely datinalaplaysise steer eee 53
H
Mnonitbs sob oyeblbban yo yale sodesndcadnoos 145
Isapishebsoletaeneus sca saee 137
Ischnochiton limaciformis........ 168
JPEVOMINOSBIS Cecgdenocas) coucsoeosoes 169
PULUIpLasScenis ieee ae 168
J
Ueborbab ee yRe ip ee ee eRe osueEaean 110
Jiamthinacommilnisys---s- eee: 104
fragilis ...... raph Ga eoetiertet bra) 104
PlObOS aye ee 104
Jamtla ina ay eee We eee ee 104
pallidayee ae sea duane a wa 104
L
Labiosa lineata 25.....189, 205, 436
Leevibuccinum harrisi 2........... 16
Leevicardium serratum ............ 200
Sublimeatumlbieens pecascseeseeseee 200
Weevil denitalliurraee secon se eee 37
WamlelWarialram oajeeeeenseeeeeereya ee I51
Warm PUSTa ie ur ve Ces aus 116
DET OED- 6 aie n Wen OUNAL No MAb iA aia na ine IOI
Latirus infundibulum 77...190,195
200, 279
471 BULLETIN 39 299
IL ANG WIS) Sheet) Wei gana Nan RN i 279
(GANELECIENE) «(UC pean cmpeRE ARG e CRAG SM EG 2&0
Webaibalbore! 275.0005... 0. 6c.c cdc 345
Geulbvaimang zc s sca et) 347
Wedardavalee 27. oss secede, 345
SHMSMMONCISS Drasessgneae, ayaa oe 346
HMM ARMETSIS Wee cscs e see ecee 25
DAMIR UM oat elses ce'stina dae nationcde ses 25
GIETROOINNS -Sassdoanesnostensunseoae: 25
LAME TASS) 27. tes locas seas 346
SaNMO Garam Nese NN Aha 25
Wepellay cwbitcolae se sels i. sdsesses 152
Lepidopleurus pergranatus....... 168
WED LO CHALOM OCR MLE cole 168
Weptothiyraiidutas ce. sete 153
WSU COMMAY ees neceo ho deascseseaaneed: 56
Wencosyadimr CMlOris)) (Sse. 66e: 231
SUSIE CIS ee te sncere essen sess, 63
Sully ooreuama cleanse ee sees 63
EM OCERAS Waseaiaseteeeecanccacece: 63
RVC terol Nene aecee lh us kae gah 62
Leucozonia cingulifera... ..... ..... 85
OCC ate aR Mee UN alii 85
Wevitusush@alei u...s5.. .cc60ccoccese 26
DAOC a ey ease UNG 26
SRST ces eS Henne SaN eB ea ncses ena 7
iLpoongeel, GOlliG eRsececausesenedcodespaceos 194
HimMoOpsismoniis) 29. ...../........ 349
Ne nceacolidiay yo eee see vees. 199, 381
ILO SGT douosenpeeoudene uaaaeiistse sober OL)
IG rtoysla): neni rll ease deapacodepecscacsess 154
PASTTOVANNE)) SoecotcodonpleeHoReaueuees 154
Gri Canin atari escapee decsosuaes 154
WiSSospita proxima.) 6.22. ses... 6. 160
IC sheave) \sxermmoyb:e AeA sda ace ieooasane Ley,
HVelantOStomMal cheeses ss eeee Miay7,
Littorina augulifera.. .............. 136
TENOTALAM Been wetteceeoteseascuer es 136
Tao SSy Osi ebaN paduacsdoooreceeeanodae 136
(CECI S WIE ean aaa Ho Been aBAar Le 137
ZACZAC Hoctenainst nee ceo oe tae de 136
ome chige Su eeten tas ees eee eA 107
Me OVO TATLTTA AN eh ok cy wR 117
Pucapinaadspersavoses:eessen cess 165
Cancellatayy aes p tes 165
Fascia tan Wen eure Ena ee 165
omGurasensisyes.coselccescoeee 165
SURAT Ae eM AUS Eek aN NM 165
Mucinavenistatan wes se. eeee eee 426
ivianiCataliis wos wean dee res 396
PEIN OL ealhs wave oes ee eee ees 19
Guadnisullcatal iss: ee ace 200
MeV AtTaroecavenlovceeatee sees are sacs eee 148
M
Mocoma costaricana 29.....-..-.- 428
elongata en sue weniec Avene aes 19]
SAtWMSUSISHNZO Mes nee eee 429
Dail ik CTSISH Ae ee cens lone esau nes 1G)
WAT Canalis Zon see doce 429
Macrocallista maculata 32...... 189,
406
Mactra alata 37........ 189, 194, 435
Ce Kk OCTANE Bia aan ae NE EH 19!
estrella nay 7nesseseac ss IQI, 435
CxOletaa yee eos 189, 191, 434
SSUBRG EAN DD ie Ay am IT 437
SUMCO]A MOA G hs Saitou eee 436
Mialeatcanauratsosos.s. 186, 191, 200
ello bicaGeeeeaceen eee cteen eae 311
TUG STIS MeN ee yee a Nia 191
Mireur teases ae ee ee eae 149
Manto iliaantomiat ie enst ssc ccnes 71
ASERIC ELAS Masi Mee eae oe 69
atmos ty las snolitacsans uasenanacs 70
Dal bea tae eR MNOR eae Ce MAIC ep 69
DICOMICA ee UO Baal 69
Cos gifs re BR See SS 70
Cerin] ae eer 71
isitm@ rit ell aeneee sae aea eaten 70
Mm elaniti cae eee tes 71
Pemiplas Creriees secncseuscoeccecsas 71
plicosa,...... SbHGHaBObacoRaHabenesess 69
300 BULLETIN 39 472
quadratas../.- 2. iaddagk oescod cof I
DD OL al Y Ke Mr mes ADGA ENS 70
Stell ata ee eee ee eae 70
Ia O) Clary een seas lence celsceliei sa matak 193
Margarita tllosalepeeccnesusccnicecnte 154
Marginella apicina................0. Sel,
AULCOCIMT Cayenne Men an a iis 79
AVEWA Hie ONeers 80, 188, 190, 269
Carn Cai Ne ena UN U ae 77
CESS eae a eS 77
Collina pore en et nen 269
COMEORIMIS se escort eee ene eee 268
conoidalise: (oe 78
denticullatay eee ee eee nians 78
FESS Do TMi ie ain as A MPI, 79
SULATICTISISH eae eee eee see ae 186
Had ria aeRO: 80
laerimelar ed oe nein eee ECOL
VENGISSS Bank B canedenadanadorsuedadsads 268
Lean Ger OMA acecece anaes: 270
macdonaldiyvouse sues 187, 267
WMINATERSISHMON ese Nee eee 268
BIDITIATTV A ese cena enees BBE NAn ese 79
EPRI ROYEAY Noe eno Besdode abe OR ae 79
MAUUSACIMA aM eee 270
Ob Ong ae RU NaN eR 80
Opalanay ie, ve. Bo esdoSceHAnod 78
Ovalliformis meses ee eee eee 80
PellUCtda LOU 78
Var pilsbryal i yige eens. 268
TOStTA TA eau ewecs nsec sees anaes 80
SO retry gee A Ae 80
StOLER a ee eae A ROIs 77
SUCCIMG Al PRD e n Nun aeien a Ne 79
VANE ia tas eset hee een wa eae 78
WATSON ea UE UL gaa ae 78
AOE SEAN Bon cooboncuosuoasucaoace 80
Marseninavamplanne the 151
Martesiarecnrvaii2 assist ee 19
Mathilda yucatecana............... 135
Meioceras deshayesi............. ... 131
TACT CL VET re on IS a ae ns 132
Melampus bidentatus. ............... 55
biplicatus ee eee 56
bulloides eae 56
eCiromalUs hese eo ON nem 55
Cofkenset ees Lie Ee ae 54
Var. oundila cliivesse eee 55
COMMOnMTIS eee
COLREN ALIS Meee secsbisece ee neee seer 55
CTOSSCAMAREEasece seisaceeceene eee ree 56
Havas eee oh ea 55
Bley gle aNbIS) sso cusbeecHcoodce’ dobdocos 55
In\ey ocho) bb Ub ly MoMA NAR Ndia uacobdoodoc 56
f EUDIOO SS Ea AMBER MPMES corns doeeecn: 56
lnmeatus aii...) sae eae 55
TOT OSA erica st ah aie Manne cle 55
Melanellaiibrlineatas ieee nes 106
COMOHC ERE ccocbaddesoosdoboonsesse 105
BLISUS Pca Ns Seat NH ae ale Neg 106
Bibba es Cas eens 105
STaciligy ye eee 105
hemphalliy cece eessoveeeeceeeas 106
Aten dias eee ee eee 105
JAMLAICEMSIS) .n-eseeeeeee eee eee 105
Subcanina tase eee uaa 105
Melongena consors 72...195, 200,
284
COTO ae ee See Aare ane 86
melongenay sees 87, 200, 284
Mere trixadanienaeerssecen tne 404
SRO) aka Fel HasUanoBaseecamaddbo seebeo 25
Mesalia alabamiensis.................26
PU a eee Ue ee ee 26
WAESONETISIS eee eee ee eee 26
Mesodesma costaricensis 3/.....437
MES Or iy 1 Supe eean eee eee eae ere eee 84
IMetiSibar baliig 2 nae eea nen 427
Metula cancellata 7}...... 195, 200,
287
Sabi Mega unesaeeasw anes 287
Joke nee chy Was eee aboneodatiacHandandac: 288
var. limonensis 7}......288
Mexican Mio Cene tusceweescsteneeen 200
Microdrillia rostratula................. 7
473
Mitra alamagrensis............ 188, 275
[DIB ASRIOD Acs. Sscadsaadoda) cocabpdoobe 82
VaTeCOLralloplailal Ose seca 273
GANG ETISIS) Oe) Saciesacellseeesc ce 273,
GASTaUVAVIENIE) ask oho sdbdcbenpjoHoodone 82
ALTE OTS TASTS ecules Welsioe 191
MOTUAMI ON Meenas ulsstenenisians 195, 273
OAS Ober menses sesierimei 188, 274
MUGS: > Macodoo esos eos cccees lays 77/7
GUNTANTAL OEM seindne teondanaas neorocace 82
GS tay Ig AMP Pe ee Ns larisiss as iel asics 82
SUVA SOMMer eis talsac scease cn se-icas 191
swainsoni, var. limonensis
GMa cererisunn ke nivale relents 272
VATICOEIMSIS cessccesdecesceiseeatece 82
Mitromorpha pygmea........ ...... 82
(EAI OMleN Gosnondessonobobens ncdooe 51
Modulus floridanus.................. 129
MITOGMINITIG eae aise sissies eniisslls soni 129
SBTORGM ee einen uve te cuetee slant 25
Montacuta bicuspidata 2.......... 20
I@DSE! A SodasnsoobadsonsbodessnsAH!
Mulinia lateralis, var. sauryen-
SUS iene sea U ean 438
Mitwreiencles), Teverrol onlUbl Aaee sbscosasdosse: 98
Municideahexagonal....0 i102... 98
ATALANTA TL A) tetnus.sssieieclins velee nee 98
CSHIEBIIFEDIOL Gogaodbubasonnone 4oacudd6 98
jOlawlijoyomehaley Seanseeooonadanana: souee 98
Wine xtalS pe terATMU Shee cc ecto ces 96
IDS AU ML i acdateGatiennats 95
[oven aboRoyaGy Aasasaadeecs a aEner ees odtion 96
Cab citaee wes cee ssescwiea eclespelsaleea sey 95
@QUNDB Is acdosd oscadonhosoebauonaE 96
COGMUTKEC EIS eases sno-ee 195, 303
HU WESCSUS opsocoucscssoeauebdecdese 96
MeahebhalSsaSiS.s05 Indeesadacaso, eaoodd 186
PIKOOOSE A poaadsea qoduac on uaERRoneSaae 96
messorius.. .... 95, 190, 195, 303
OCI AES ee eee cetaaln cium sicietre 96
POlyMemAatiCusiassececesdcees-: 186
[DGPS TOs consviendces cabddanuedodasogec 96
Costa RicA M1ockENE— OLSSON 301
Myrtzea limoniana 35...... 200, 393
N
Nassa beaumontensis............--+0. gI
INOW rel yoedopaqopeoosn6ed aodooRdod go
OO OIENE es sonscmododdoucoopasodadasoc go
DUC OMA Ese dosent eyielewsealetestier 12
NiaSsaraimallo ts liitenwnescsesscccosia antes 89
Columibellla tae cere c scala 88
PBs A ton bo dbonenenoRD00d0 HOdodaHOC 88
INaticaallit(calll oSals-eereteeenesticses 147
DONS Heese se ece esse esectiss 186
Cala zZOnaliSmesereeneecate vere. 186
caurena 76..147, 190, 196,
199, 327
Spaoutratbll Wal ococacces asesapeoooobN \ocg98e 26
FORCIATIA Mae bene secnaceecmacees I5I
SUP Py awaw7Owevessaees. 188, 328
J AIVIAICCH SIS see. uescnmnaseernt 146
EVOKE so kescespaccoancnopAnecoudcac 149
Miksysi Gl ca eee Sk aero eet 146
IShoe hull eH are aaa anos HOH hn Asa 26
TAT OCCA AME els acenete renee 146
ON TRBISLE GUAM IA MEER O SA SE AM GH MeN ane 26
jouhveene Nal seh cnoussadsasooscanads 146
TOL TID ee sa aed 146
jONUSTUU ele, b Sansa Dod oHoaeeSboHsonouGo 147
IMENTS sa sodnanoadoncosebonqaudoras 26
Sathorn vate pau Meee Saensies Woaeas 26
Sub Clansaang esse seecaeeacereciee 329
GRU KOH Ya conassaaasonaaceosseedadanes 328
SOULE Ieee ncece ee actaseicerleteaist 196
youngi, var.cocleana 76 327
Naticimaplacteate tia ehescecernesce 149
INGOs iitrattai ape ne ee ee eee eee 112
Nextag pelorontasseyecce-seceeerree 162
tESSell arta ee ee eS acee canara 162
VETSICOL ORAS seen ee ee ae tenet 163
INIGIALGWAE), FOYOL A cacdaaocoasooosaoosceonad 164
HECHivatassissnceuee see uetae 163
Showa Cerise ceecneaeeceee 164
Sparsilimeata yeseycesseeeeee seer 164
WAL Pal ea Sue oueeeetesnen seas 165
302 Costa RicA MIOCENE—OLSSON 474
viridemarisy 73s). 196, 331
VALLE See ese UTR At ga ae 164
ING Veritauibriinmea eye ease anene 150
CamipechienSisniesimsseesnaes 149
Ghuaplicatale se Meaney. 148
foal hbWey WARES SAAN RAB AAUP UNE IQ
moat llariseaeececoseeceeeceeeeas 150
nmereidis eu. 187, 191, 196, 330
TULL eee eee eee TA
fexasiania ya Winns auiy 149
TAD CT eA St bleu Eat ea dee Rian 149
INTIS) Bevel SeS nos gavasdasdooss sasdowobs 106
Splendidiuilay Wayne seecen. 106
Nitidellatparnviuil ayaa eer ues 94
INO ROTA VAs OU id aaah Nea) 74
Northia morthige eer. 190, I9I
northiz, var. miocenica
TZ SSS sbpowesbocobesopaGEbosaaoD 296
Nucula cahuitensis 27............ 343
ASEH Calsheecebu ace entean ty arene 186
ml ediawalaye sen sence eee nee 25
Orbicellay eee eee eens 343
OVA eee UES sal era a 25
tenuisculptay) 27.0 193, 344
Nyctilochus femorale............... 116
0
Obeliscus tesellatusy eee eeeee 107
Meine rae PEA aaa a Go 117
Odostomia acutidens ............... 110
ats Chay ize ea ee 8
CLONKONKG KYM IBERIA ABA a LL A IIo
ATP LESSA A URE eM CaM aEm ed 110
GesaabboneKolah Asscanedes odooboxe hesoS 110
Oligocene: CostapRicas ayes 181
Oliva brevispira /7o......... 195, 260
Carolinensisw saree oe 75
CInCiata elma NU An alae 75
Cylidnicani7o Maes 195, 200, 260
Oliva gatunensis 70............... 261
SATAN see ANTRAL 261
Nay les aR HAS ae A AIA 75
litteratae sue rol ce aaa 75
oo AOEMASWEY WO Aosoanecadsdoness 263
PRELO(OWULN EIS) isbadeoncanacdooane sunoce 260
sayana, var. immortua 70
205, 261
testica, var. costaricensia
7O 190, 262
HESIEIOEE) docoddogseusdpoodson I9I, 192
Olivella, var. bocasensis 70.....266
IDOWUSSAICION 7O NS. ecco 265
(21 bi del(o fs) ar aren aanMmRRerEcn a Sriagocaaae TW
VOT allele nest) ee ian 76
PalvieStOmen SiS) see eee ee eee Te
OMAN WON teste eMeeanee le 264
FaSpidea ys 25 ua eh Meo ua 76
lito menSis) Zoya ee 188, 265
SooKO DEENA Su SABU AAD BR RKO Bua I 26
PTA Ga Cars 0) keer ata a re aa 76
MUticoides ZO 195, 264
TVG A Ne AU e Tenia Oe 76
jOLOKSSUU Fa OR eran ea Lee 34k 76
Semilie nth ca iy wees ese eee 13
SUD temalra ais uy Seen nana ral
LGESTUBIOEE), J ses Hoscooseccoodasqbedcaqon 192
QMS Cia eS ean ae aed eae 118
Oniscidia dennisoni...............068 118
Oocorys abyssorum......... Aeaseiene Il7
Ostrea costaricensis 24...20I1, —
368
PuehdeboleralsilSyarnodiehasdsoanoscboke 186
interme d O1desiizeeeeea se eae 23
MEesod on 27 eNas 189, 191,
194, 199, 367
Scull pturatay ese meee 201
Ovula regularoidea 7......... alee 14
(Oran lotankananveteal mL A eee ie 112
EhotoUblawebbaal aAuranog ;yonsoenaocaney 112
BROWNED HONS BosaanGansedddas Géods 12
@RUROI ETNIES A onodoosdenanee wsboce 112
Subrostratumecees eee 112
475 BULLETIN 39 303
Oxygyrus keraudreni............... It
P
Pachycheilus anagrammatus.....129
Gaynillierarere sca noneancobeseeoncseace 130
SUA WAS een eskascovadenaasenes 130
Raludestrina aldrichi)......00..... I4I
AND CTTATI AH eee ee LeU a 142
CNM OMA bay wenense see vecevncces 141
TAU IR eysls ea seielaetesemeloies 142
SD eNta Ae eee ees ecsnieclcis are cee sibesis sive elss I4I
LUIGI CU Deeps aiaala'se's'sna/ seelesidagiai 142
RanopeambelllSensis) | 3..0..Jes6..-) 22
JERR NEO) os cage cod oeeoeBeaeedn ne sbeebs? at (0)
IPEISHUE AKER, ABNER elo pocanpoaneosenceoneore 7
Reartellanasstcnijeseledesoaccadeegeg ee 58, 151
SOMES ETS oases at eecustsccusomnisesi 143
ROM Cabal eeesen ce cecccsues ie ssteise 143
Pay TAU eAatlameecneasee cess ce seseise 149
RectentallabarienSisicscsscescceccce 25
AlminattensiSs 27s .0) ose. 375
OLMIS MOTE Meco catce acts ceeculees 376
Coraliliplailan) Zones. 188, 371
COStATICEMSIS! (27 yeeneeene ees 373
ChOSAN ee Sasi uils weedeouusaebens 186
SALMMSM SIS Oscars eecses 369
levicostatus 9...187, 200,
371
ISITE) AO eA AM an PRED acon Tel
MACH OMA WOM nsees ssece oe: 370
Olli@OllejontS.njsabanynodooons poseseo be 375
operculariformis ............... 186
IRSA OES) AOsssescadodsocsssas 374
TRSILIG|ENBIS, bao sceesbepdedoedeanee 186
Santarosanus.............- 200, 371
WEMLTANCOS EIS souoscaquducoseasceons 192
Pectunculus castaneus.............. 352
GISCDISEENENIS) co sonadadedsseoos 350, 352
[OSI TOSS jeeaoes dosegone 350, 352
inedicullariavallbiday we eeeneesceses. 113
decussataee se teess RBA E ATED 113
Redipeselonsatusecrsscese-ce ees 54
TaTRbRAN ONDE Senasciou MocoBocoECESeeEeae 54
SOME COHNGKES,,.. Sonaénoodsobusqobseccbes 54
OVA SHeeeeeriencee ee sneiies cies 54
GidadinidenS ees scsatecrsomeees ee 54
CHIGENS Baars ccewthos tocecerecaees 54
Reracleneticuilataycsssscencese esse. 44
Peristernia insula 77.............. 282
WON UbIEXEN) | Wiay lencooasop se stoans 282
Reristichiaraortee masa cssseancscers> Il
WOKE) LOAN be BBS Gacodo dobocd cncoadeon III
Personmay stra lilitadase ees cccs came sce 305
Petaloconchus domingensis 204,318
ELECCEUS ee ereruricieles|sleniselsleniserlseis.9 133
eae Cu atl Sueeeetarsloletsetetsals oat 133
sculpturatus 77...196, 199,
203, 204, 318
VAMTTATIS Hy Wei uennani cic ain cratstare 133
Petricola millestriata.. ......,..... 187
RetrOtTnO Chi S eee scc eens 164
Phacoides actinus 35....... 200, 396
DOCASETISISHMpoestersceseeeseas ce 395
COSEALICENSISMGSieesseteie sees 304
var. medioamericanus........189
RACHA SoM eee RTA 189, 205
radians, var. medioameri-
(CANONS | 267%55) wdoogudtioooobdec 393
(ero IGANEWIS) ncausocose 189, 205, 394
moiuiliferum 75......... 200, 307
Phasian ela ieee neck asc seceme 136
DRE VAS se seU ni eats ue ee 152
pOMOVUHIS, 2) oda anu sudonaecBogoceo Secbe: 232
ORIN ey, Gsosaqcooson506 adoooabAb 152
PONE C ENG BoGocogsssddconnseecadde 333
Dhabi beavers >>:UORE\ cased dqsascuns aéana600 53
Phorus agglutinans.................. BGO
Phos) aritillarcammy ese. eenes | cesteslas 88
IbeteivjenSiSimi/2aaee ese aasnstes 294
(SUAKG Erloncosasaceod! eodusuan oaasdoae 88
elegans eases ancor ... 188
elegans, var. limonensis
D RAR EEE EAE DER BSG Nae 291
EStrellensisimZeesceecoreee see 292
PalvestomeMmsis seeeeces ss eeeee 88
RYOTE | MEeasss5050 000980805 289
grateloupenSiSwerssssacesssess ee 88
304 BULLETIN 29 476
ATCT CALUIS Sse ee Neen Magee ene nnauas 88
00K Mb KO KEKE) VAZERS een asoscoBee 294
mexicanus /72.....200, 201, 289
Moorei, var. costaricensis
EDREAMS Me ASAE AGREE 290
PAT VS Wie ean MeN et Aerials 88
subsemicostatus 72........... 293
NSEC OE een a dogabe 293
Bh yWonOrs eaastencecromeene acre 96
Pitaria boucaryensis }34%........... 4It
CIF CIMA tay eZ. ses IgI, 194, 408
COT AE AR SGOT AG ee rater 187
SUP Pavlos ae eee 409
hilli, var. musacina 34.....409
SDV VOUN AK EI Oe son Ousnh oda conedesoude 409
Va mwatelese ry Gemma 410
Placunanomia lithobleta 27...194,
199, 382
Rlanorbismopiisun vue cunnca nse) 56
Palartay;s clad Gest ie ya nana Murase ease 4o
PLCUROLO MTA NA ROE AAS LON eresaue nat, 63
ACCOM a RUE ON a SUNN Si 26
MNO Ke Leben ine ie ANE L NARA TANS 230
BLESTMO Lata yi ana 209, 234
DTT ae INT A Aa a tie 66
haley Manabbonl) oooh dun esnogadoouededc 220
Pater tetera eaten Ne NG Wain 230
blalkeamalsci ee ram waa alien. 68
nyebey loyeobal ole sce ous ddWoonoe 70
COR TTI AI Ue MIG MUNA RIS Nae Ne 70
COMSO Ras Cea OA Et aR ds 234
Far Mery Wie wee er ukeserene 8
FoR eeeT BLO Eee ACS ee ASE REA ea 69
AAT NSIS nhieu ne Boerne 230
Sve! a porn ds MAA RIAN en ea 8
DOKSH ea HOKORY Ae oA nKe a ogsonmadsadedaae yf
Morekeah oYSrasHIS. Won ldadsnodpdsoaaddaqae 26
WENT, TOBKE OO Ascoasbdoakoe 201
MMSE AVA Le aula naan 26
PETS ale OE TU MOMMA ali ee catia 26
PLC ALA ARSC HNMR eae a a 69
GM SECONLISH Eye Naan teEEnu ts 26
SIMPSON Ai Nee ne nicelcatat 66
Streptoploraa a. haa eley aan 68
VERUS TAMU eae 233
VITO en a ee 230
Pleurotomaria quoyana............ 164
Plaurotomella agassizi............... 68
blakeanay i Cae 68
IDREVES ANNs U3 aa 68
diomedece siege eee 72
EXTECH SAN Mes sevais «ose ceneeeeeeeen 68
IGUBUH EY 21 os MONEE MER EES oad 00.c 68
eal eee lie 5.) a) a 68
ATT CTT SURE No 1 RN Ve 69
TeWCO nda tasers see Se 67
Wales s00\e> OTE s osaopodosce 68
packardaye ces... iss eeaee eres 67
Piicatulaydensatas-.\ 5-5 380
Manoiwatay) Gece 203, 380
Polinices eminuloides..... asta 148
HAbaKeAU ey PN ocosadoaaso dos BARES 148
JOKES TANGO] DHIhg dogtdocosbocessooes$ 148
Veptale ayes si lan a aE USN ea eae 148
stanislas-meunieri 76...196, 329
Subclausaly e7Olsnaeeeee 196, 329
PET TTS eh SU ee AE 147
POULT Aaa as ASR UMS Ray ROD Aaa ae SR 87
JEL YSCLODIGIES). | osoocatasaccsdaadeaeono6ces 4o
Potamides, var gracilior ......... 127
ATL ATS O TMA re se en a 127
Satillensisueemen parece ecens 130
Protocardia costaricensis 30...401
gurabica 30........ 194, 196, 401
harrisisccuaaeceeescosseeaveseeeaen 25
Psammobia harrisi ?............... 24
GyasbUeoll i eos LA penacdoodacsbuouudabodse 23
Psammosolen gatunensis, var.
costaricensis 32.......:. 432
Pseudolivalscalinaeess--eees eee 26
Evab (CeO DONEAL Sngnoqhacagsadaedecoos 26
Rteriainormatayy i eeeeree 194, 367
Ptychosalpinx dentalis 7S...... 187,
281
Puncturella circulars. eerscse 166
triancullata we pos eeeee re reeaes 167
477 Costa R1icA MiocENE—OLSSON 305
Paneturellay trifolimimis yess 166
WIVES OPIMUE i nna Mclain large 166
JPARN FONTAN Dies RAS era ate ae 100
TO CUI OS are eves) pei alenin ene an ys Iol
TAO COU Bb Ibe esayronesenadeuncnase 305
Ryramuidellal candida... 0h.) 20 107
Crermiullanca ye Sea Aan Manure Sn 107
Golalorataye ee i ue le 107
aA CLO SUNOS de aeaHalannoussenbien: 10
ODEWSOIGES ie 7s eee eee a(@)
SVTTHHTS co aa ee 107
isa ita areas oe sk lobe es Soka oe 108
Ryiulagabbreviata... 04. 2..6s0.se.: 101
PAUP AIAN e 6 .scciaaneeccuensnce 119
R
TRavaSI Des oN UAE Bane aii LOS eAM TEA!
Clavelmrartaaen ey ede sire ba 99
CRASS a ee ris tte nicislerecia aaa Slearels 306
IRGCHNSAL IBMIIS ues cso sdedandadeduasedse 49
AZO CIMT SE eee ey cen och eeene ene IO
Rucinmlalnodulosa...)62)5....0- 305
IRGBEMTTD IE seo yeaa SER EEE MGR HEM Sena E Man: 167
AS HUTS ATA geese eecste aaa! 167
Ringicula dominicana.......194, 196,
206d
IOS OPA TARO aioh hodnmbecocakbasene 186
Ke ptochielansy eye eae 47
FTW lh, ososoeaeaaACMeUoreeebe aeolas 47
SEMIS tslaicaa meee eee nie ces 206d
Rissoa acutiocostata................. 139
CMSA A IN Pee Ee PENAL 139
PSRCIPICAT AL. le aes keels 138
Rissoina alabamensis................. 26
LOONATENOE, ccosnagds soso suosdeneE 139
Chesmeliprara tts ssce socks este 139
GECHSSA tA MUU ea: 139
elegamitissimialeneeeo-ce ose 140
Ben raeren czy OUP Sa RAISE ee a 139
SAG AIAM Aes MN nee en seal lne, 140
Rochefortia minuta 2............... 20
Rotella cryptospira.....-:.......... 162
S)
Sabatia bathymophila............... 50
Sanguinolaria alouata 32........ 432
Santo Domingo Miocene........... 92
Saxicavella alabamensis 3........21
aly Sa OUe ie nha e yk etn Kant asian 56
Scallania raat Woe iy webs es OM 102
Scaphander librarius.................49
161(0) Oy LIS ape AnABdrlespacbeonaanocbeen 49
PMU CLOSER EMS Heese enee nee Re esoe 49
WALES OTM! ML eee Lia ad ana 49
Scaphella costaricana............... 187
COStamICenSis 7Speeeeseeecer eee 272
[MELON aN e eee ee ae es 81
Scobinellaymorieretim7scee ee 251
Sconsia bocasensis 75.........-.. 309
Cocleamayynseeeesedsshe 187, 310
ligewlcatalateeecaeees 195, 200, 201
Wate Sallis ieeeenceace 308
7ZAOL OK US UNEASE ENE 193
GUIDE SN ERMA conusanaddstoesaeeaso 200
SYSTUADVSTOWAIE. ys Srisissoguasonocabeqcbabesec: 158
fORMOSA Nasa nsenseacaceeeeeeeens 120
ACOMICAL Maa vensirescustacsedsn nue 121
MONO CINTA tale tenes neee: 120
(UPIGFODEDOS 27, saons adnooadoosbeasdcas 120
Senlavadian sien eesec hase eres 124
Semele claytoni 32....194, 196, 431
DEE VAS HESS UNE rae maa I9l
var. costaricensis 32....430
iewoverckoratlalialey 1 Beso ccuseba ionsseos 18
ONOVOVTOLESUIEHS) | Zocsacuecoccosasocooss 19
SEN Ol S72) Osea ncosdacdecosidasgacee 430
S Era CASSIS sees een ue tiaeed Mane een Neen 118
Semiunellalstearisiy wesw ete seen elt 94
Serpulorbis papulosa 75...196, 200,
317
SCulpPLUravUS ssseeeasese see heee 200
SHEEN TEESE wesc bogne! ob: Hops opoddsmonon. 150
SALUMEMSISM ses aeseee lee sacra eee 186
TEMAS vAptEwOVbbOOl p audod codonoossnde 150
Zab bie ee eoe auan ees naeee ys 186
Siliquaria squamata..............-... 133
IveoKOWOVETHIGTS) cosconaccpoceos TOS
MIO esta ena wi we 134, 188
306 BULLETIN 39 478
Siliquaria modesta, var. limo-
NEMSIS |) 7G i eee: Eta 319
Sinppiapiaterme diay ee eeu sees 112
UBIplicatar i. wees. bb SbodeHOdO 112
Sinum pmacula tums essen eee 150
Spoahbhaol saoNWOKOS Me ndoas4Auas /abrcoaduade 150
PEKSPECEIVU MIME eae sees 150
Siphonalia quadrilineata 2........15
Siphonaria alternata...........0......58
lanate@ arta eae UST ORs Me neN 58
Siphonium nebulosum.. ..... ..... 133
Sistrum nodulosum 7}............ Lol
190, 305
TOS CUTIE MIURA ile aes IoI
Skeneaplanonbis: eeeseselasceaces 140
Solaricllatzeclecsi) uaa eaE5 7
alaibamenSisneees yuna a aeene tate 26
AIMADILIS eee e ee Meee Mie sal 156
Wars ClavaitaaleMuteniM sect. 157
Varo cideal sna nen ie 157
lissoconchiayy swear ie 157
Dub ri Sa se eM ee UNS Ae) 157
SCalbrclus culliaaene en nae ee 156
Solarium gatuniense ie 326
Scat UTM hue ease sstanets 326
STAGES NG hbWaOL oneghdoogonedadodon 26
svatll areal Loni e eu nee meme 200, 326
Solecurtus gatunensis.............. 432
Strieabla tus yea ee aaa 186
Solenosteira chiriquiensis 77...286
dallas: UTD AUS ERR Lee ers 285
var. medioamericano...200
Vvaul shanty ea tenon aua ny 200
var. medioamericana
UA hc cchbaCoo Han todaaugcHECondsSod 285
SPintalisinrostrallisn yay eee eM eE 44
Spondylus americanus.............. 379
SlaheapenlesmeyG’) ey ee 378
gumanomocon 24...... 194, 379
Strepsidura heilprini........... DRL 25
Strigilila) musamicalwi2o4 eae 427
Pisilonmiicieey eye Heey 194, 200, 426
PIStoriis eee eee wanes 189
Strombina ambigua }........... 298
Chiniqmiensis i7opr esses eee 302
costaricensis 7}...... Bae 300
CaCO) | 7 careanocor=cassanoe 299
gibberula, var. galvestonen-
SSRN ae dc cA A a aN gI
NESSEPONEITE, | Ro asrbacdeoussconedc 2098
UL AGLIMV AN OTge eta a. 8h 299
var. musanica 7}........ 301
Sim COL aig 0.55) sete een 301
Strombus accipitrinus.............. 120
(COSWERGBIS) Gadsobote qooscddes sovocuaKe 120
SACUMESISIS| Ose eae 188, 313
pugiloides /6..... 187, 196, 314
UA S ee ee eee 119, 200, 314
Stylio lag wea a ae UVM NaN nae 43
WAL OUD A a aces ea eat e 43
SHsMoOe), Ghashablodoueosooeseacueac sooder: 107
T
Macelisice bushi eee eee 433
Ma Pes Syl Saks Nae Mile Beara 420
Maranisnpulchieliiay assays woneenee 72
Tectarius municatus.../2222.0.04.5;. URz
Teinostoma cryptospira....... eee O2
ROSES). Wicaasecaodsao) dopoocessos 12
subangulata, var. smithi 7...11
Tellidora cristata 29........ 189, 426
HeliinaibelistanaG 25s seeeeeee eee 17
cercadicaniaqa eae 194, 196, 425
Costanicanal Zone cceeees 423
ery stalllimay2oeeeneec 189, I9I,
194, 421
CyMOSLOSSANN eer aceeeneeattee 17
dariena) Zo eerste: 424
equitermiima tae eee peerearee 186
esteliltenmsisiy 2 eeneeeseeee 17, 25
seAN A DNOVOSH 3 54s2ndhoedoue yuocpedoe 429
KEW ake MONA LUEBH ann MAM TU Mm 7
leprcota yi Zones aaeaee 200, 423
VAST APL U. catalase edb geeata 422
479 Costa RicA MI0CRENE—COLSSON 307
THONVEUIEN 21a) AON ee saat 421
TRON AYRE OKG bg ayy Marae R Hn ta aD a 424
SQM GOVAG case nantesnececue ee ees 424
GTA ORDA, VAAN Sadao sot sodocd 18
SemmlinOpit Gama veh ses sae seeeeeeet 18
MOAN OME tanh eet es 2OGM AG
Tenuiactzeon pertenuis 2......... 16
herebrasbembmMalis.) ssc.) 202s 189
var. boscasensis {4...... 213
loerliimeces ) Veaneuoocacuounn 194, 211
ipancitayyegs) sc... 194, 199, 207
Wali OCASCHSIS He sneccees. sees 189
CATO MIMMETISIS Hacer essen ueaee 60
(CINGVETEE. ona oS sen BABAR eH bee car 58
COMEA VAR aE MMe a uel ae NPE LL 59
COSHATICEMSIS/ WI Zin nese tessa: 212
iSO cata weno une Ch a 59
Smell vaey | ess codmoneouosscon 212
SalVeEStOMenSiShess.. sho eeeee nce 60
SALUT SMSISI ua esecces ince scee 208
PAUSAP AVANTE seesieictslerleeiceles leis 210
HMAItEMSIS NZ sce ss scees 194, 207
HATMATCETMSISH (eh. Ub seskiless ess: 58
Te sayexGVovratly od sdeeae ciated an uRal 60
sinner total eee LA ee ce ae 60
INBHtESOSTINS A oodans, eocsduoaseseoancHe 60
War. tmiddenSis) |) ae. 211
ASSUME Ne aseaeatisraiens dete Saige ess 60
TOSENGT a cb eunt gem asnconALaResPE sa onect 59
]OIROUED XU eacnouce encdonod noses uoEsnod 59
PULA DILSTSIS| ieee a EEE Bua
TRIBKG IS ae AN A Se a Men esi 59
SANSA ecu e Gace Wee Lies 58
Gyovberbiese 72 Goosaasoouaoss 194, 210
subsulcifera.......... PN Oi
USS EEE (SOSH ER SER aM EARS 60
\/GUT COSTED un nai nO MSE ee 59
Tr Olbieteavest |" Has Aaabebocoboe 208, 209
Teredo ringens 2............... 1725
ANS WAS FOROS Gav daoee asusodusodsonue 53
WWBWUICob-ald. Beau doncoal sbedemaeaeeHeen 53
MblTanSiG elitoid Casneeewseecseausccees oi 100
Juleehaavalsyovanlalys phan oqunooaeasandane 100
Wags anon shale soceoodcocgoseoc 100
JOENABI EL ouh ca wenoobood aggesonodudodne 99
MINEO COX Ses tee ecenceestelecees sais 163
Thracia gastunensis.............-..- 187
Hite SEA TINY 2G ese elesiselelsleletertsl/=le'or 383
Aone) AVI Ws oaodosagosbesopassede: 119
PETC UKEP sese.speescenelnnnns sons 119
AbloNenl Fal aAsuansodovsdosaosaeadspescesoene 134
orinianCanalikeraseneesscsetasscet 138
CylimGriCa. so... ses eecns ener oxen 138
Tornatella quercollis................. 26
Tornatina) Ovi a cee canoe AT
coix-lacryma.............---+-- 206b
Trailrancdencuilatarcessweseecericelscicts 57
THATS CU) Aeekecmesecescsecelelece 57
MY OSOLIS. ....06.....cceceeereeeee ees 57
PUSIlla..........e ee eee eee seen seen 57
triplicata..........+..-seeeseeeeeee es Ti
Trigonostma ........2.-sesseeee sree eee 74
Triphora abrupta...........--..++0+- 122
bigemma........--....-eseeeeer ee: 122
ENG iecaodehoundaaseoasadoccH ones 121
cylindrella.............2+2++---- 23
GIACOHFEI| sadognosdoscuouedpsoaeusee 121
ISURCLBIS) gouooancdossobs pdgssbaasasonbc 122
sabi NVEbies KES icacHanaundossuoune 2
Hole Ntlsnadcavossbandoass boobeasednde 122
perversa, var. nigrocincta...121
(HONMUKCDU 2), 5 odod6e suoeeeoonDoboSERs 122
RUIeIGYeVe ULE A Gante Hadonanee aaaceogse 121
GBray quence mays dseehosedasaasoadeszeoosaso5500 44
P Rrerkuova i Gdav DMNA AON AR eSB ate oMsGN eee 115
CEhelBEDTSL .Hoguongaas Gs5d “323495 117
lacunellus...........-- -.--.- ----- 99
PAL VUSHesanseor sere ce scent censecian 88
pulchellus...............-.2:+2++- 116
Showaltetivnescwssee beseca scr 26
GyoambUbbadtbISy, onsso5e5 posoHooDoCORRbE 305
A WaltoyaoUl@el. do ouuscoaas4secosonSuaocHcenac 87
Trivia approximans......... ..-.+-- 114
@arverabiolblley candadsosbsupssan Goose 114
308 BULLETIN 39 480
MirivaayclODOSai sens) dear eea asa sacres II4
OLOT a eee OA Matyi alae IIl4
Peddculus yy eee deweNewesidas 113
UL aye UO ee ere ea rale ee are II4
Sphere ee ence eau ieee 114
Suih Sa eee Tue alas 114
Trochus agglutinans. .............-. 332
Ebon iel) on bi aao4 hon Hse ceaete HaEBRaGOu 156
Venticulanise estate: 129
Truncatella bilabiata............... 142
CALIMAETISISH ee see eee eee 142
Pulchellanwrscaseeesstnece seers 142
subcylindricay ey ies.ceteseeese X43
ar boalllpicdateeee sie eeesene castes 153
CAStATT CUS Mp e ssa asceecne estes clieres 152
Crenulatus ee ee eeaeease 153
aA |bbA(CRRUDISYS denadanan | Caseoucaouce 137
oul chre ll ae ee eee es seneleeeter 152
Salts ie Tor wea eaasi nee 333
Spengleniamuse see) canes lene 333
AUR D OMI ae Mey anaaie Nan gna Hl ema ats 107
belothecane ry... Te HeOSHE LT 108
CE GbKO Sa ducdudodedseeeasue acueneac IIo
Chinensisi ii ee ea Gia 9
COTTA CAR eee eee 109
CUT Beem eeu clacie aaecivasactetcleiscts/af= 109
Ge-cU ba riscaelad donachtiag sescadauader 108
INCE LHW play Mey eeee sascue eee eet 108
ObeliScusH see eee. BUEN SA 10g
[DLEDTTEEOEEY | dines Acanopaosecosaauddeda 109
Leb Culatarc ees seesecectse sacl 109
G) nS) bralg SAadsdan ecaadeadacedadaedade sexe)
StylifOrmis ............2- seeeeeeee sate)
tardisculusiy ese cesses eer 9
ilep:< GU DIS RENO BeM EAB Ba SS SRO HeRE 108
tuseahomenSis zee eed eecce= 9
PUREE sen saotueaseooabecasosoocdasrood 109
Vinbaks FERGIE Goobocoa Jacoepeouodneseeec 108
Aurcricuilavlav Amare eeeeen es ee eee IgI
lawanO1deSun7aenssases see IQl, 227
var. limonensis 7....... 228
(gapbariate), |) bicgdqsacos, waceoouos6ae06 229
TIP eyAL V igbyabarckoodaceacedan seed 229
Turris albida 7...62, 190, 194, 199,
230
Webes WAU 6 Gocuseda Keodoodce 62
binronmtatus ene eee ee seraccet eer Ti
denticwla tay eee eee sea serene 6
TmaKOVIHVVENEBI ssacengag dacuenocoosodso 8
TVOYOIONGIENG) Zo ccouddcabe: sadnovdoe- 5
SPOXCHIS | Ua socandonagssnnocoo Buse Ge 6
Miarpitellayacroporayeysseeeeeeeeeee 134
alalpatmienSiSs ve-cs-ceeneeeserece 25
evlonbbieh, 17775545 akaoe [SS B2l 322
var. chiriquiensis 77...322
var. costaricensis 77...323
Chipolama aes Pas eeeeee cree 135
COUTAGT AIRS 2 5) eee met 320
ExOletarn alia a aeeEnees 134, 188
var. limonensis 77.....224
Pal DU OS eek cen ena 221
SALT SISIS 7 pears eee eee 320
lohbhasVSCost le ccdabou=ob sorodoondooos0d 25
var. limonensis........... 188
UONHATSWES | IH cae ooaasooapegadehes 321
TNOLLO MI yee eane ener ecerase 25
OMAXOOOD 17? docchongonascecyeons 324
Sata ea Cha se ues eal eye 135
Subexumdiheras se. tane eee 135
tenebmatoriadsh eee eeeanere 135
CeTmMeESSeEMSISH 4. coe neceee aoe 25
WOMEN HEI de GugencconadUsesconeooeoud 323
NW/BNEISTINE) | Lobo sgnaseonsed opoccca0n 134
VWillCalecaiasee nasser cere 134
diy plhisvalbatiism eee erste 195, 304
eFN ODP UAE Aap banabuoodassangoouspoe 186
linguliferus, var, costarien-
se aedannnsndenonascHooGood 304
U
(OMS TEES SAcgusdedenweGasesess cosoccoco000 112
Uimbonium jbaimdieeens.. eee 158
Wrosalpitsx| cinereus: pencnaeen 97, 282
OVATE Coos setnnoudgcssoodoosdeos 98
perrugatus......-....0............-. 97
{iehan OB DOSS So go nndoodcosecedocosoade 97
48X Costa RicA MIocENE—OLSSON 309
TONES SAE a AINE 48
V
Vaginella chipolana .................. 43
Wateoatnttll aise ansciecsiaisccevetulelceusscsseee 58
Venericardia conradiana, var.
ItmonmenSiS: 9/29... ..e.sce- 389
scabricostata 35...194, 200, 388
CaTaysIMMME OM ss eaaldceciesece se as 388
WHULGOREIIGIS peauetosesueBedosouesoae 23
Wemusmblanidiana. 62 .5.... stec eens ne
CIOSRBEMIVAT soaacase wsdeoneeodeseay 200
AITACIUAL ARE Ne ccs) scciioecies wetioes 4oo
AINA SHAM CARMA setae cients eee 412
TTMUNELCOS teas asc eee ees ee 4it
PU SOS ARM Ae siete) eslerceieisw eis susiee( 412
Vermetus papulosus................. 317
UChr reo soem scate sacar 200
SCHIP EMA LTS Ie ese eee eee aes 318
Vermicularia nigricans............ 133
Sioitertaemaudey unas sa! ALUN 132
Vermiculus spirata............-..-.- 132
Veronicella floridana.................58
Vitrinella hemphilli........... Naa 159
TIDNESHCTT Bl ORE Seounoneaneeacencosoaaon 159
TNOOREAM As scone ne wen scene meee 159
AMAUtCaTAMatas.- 2). 4h.ss.e os 161
IGEN WOME SAL aA eae RRS ream alotay
Chill ONAN sc pApcuocoseodéocuceoudsonDdceN. 81
VATESCENS patina us rovesesecesnnenans 81
Volutilithes limopsis ................. 25
TOBA GNSS Gagos sdadogsosdonodoooudAnAge 25
WOE) oobaGdasssobanbooDsHoccdoodaded 80
Wel Aichi ke Wee aaioRtrricoc ha caiSceSHensbanoaea 48
WO EWaITelS ooscodbaopoade acodonquaccacogace So
Volvula cylindrica.....190, 194, 199,
206¢
Volvulella micratracta...... ..... 186
WwW
Wolliliatratulenr eb Sipeeereean ease iessesce: 58
X
Xan CUSMS COIS Hees seseielssiece eet 284
SOO) DUNKS 7.22.55 Socoseo \aobensn0n 283
Xenophora agglutinans............ 146
Cais DaSalyeniasaee woes nace aenioceercisee 136
conchyliophora........... 145, 332
Y
Yoldia bocasensis 37/........ ...-. 348
eborea........ GON I AN ea ca Naan 25
Z
TINA AO WBMP 4, Seocbaneanpooosode Se 36000° 154
"
ADDENDA ET CORRIGENDA
NotEe.—-All page references are to the Volume and not the separate
Page
47
58
WS
A
81
IOI
104
164
188
206a
275
306
318
405
Bulletins.
For RNIGICULA read RINGICULA.
For Siphonaria lineclata d’Orb. (reme frecectziec)
substitute S. xaufraga Stearns.
For Olftva circinata Marrat substitute O. sayana Ra-
venal (Cat. p. 19, 1834) and place civcinata in
synonymy.
For Marginella aptcinca read apicina.
For Scaphella junonia substitute Maculopeplum junon-
zum, this species being the type of Maculopeplum
Dall, 1906.
For Epztomium read Epitonium.
Janthina communts is regarded by Johnson (Nautilus,
1922) asidentical with /. janthina. Bartsch, how-
ever, considered communis and fragilis (equals
7anthina) to be distinct (Bull U. S. N. M. 1915).
The latter opinion is followed here.
For Petrotrochus read Perotrochus.
For Chione Rowelli read ChioneRowleet.
After Acteon costaricensis, n..sp._ insert Pl. 15, fig. 15.
For fuses mtocosmins read fusus miocosmius.
After Bursa crassa, tor Pl. 15, fig. 15 read Pl. 15 fig. 19.
Home Belwea readily GaWeas
For Gafrarium limonensts read Gafrarium limonense.
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